Madygen, PIN collection 2555, Dzhailoucho area (Triassic of Kyrgyzstan)

Where: Osh, Kyrgyzstan (40.1° N, 70.2° E: paleocoordinates 36.5° N, 60.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Madygen Formation, Ladinian (242.0 - 237.0 Ma)

• Aristov (2020): The age of the Madygen locality is usually reported as the Ladinian or Carnian of the Middle or Upper Triassic (Shcherbakov, 2008). This concept emerged, because I.A. Dobruskina (1995) did not discriminate between Ladinian and Carnian floras. Our data show that only three of the six families found at Madygen are known from younger sediments (Fig. 1), and therefore we can confirm the assumption of the Ladinian age of the Madygen locality (A.G. Ponomarenko, personal communication, 2018).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - small; lithified siliciclastic sediments

• Oxbow lake; Madygen basin; tectonically-induced intermontane basin. Complex facies pattern consistent with active tectonics in upland region.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106088: authorized by Matthew Clapham, entered by Matthew Clapham on 02.03.2011, edited by Lindy Edwards, Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Diptera - Psychotipidae
Psychotipa depicta n. sp.31 Shcherbakov 1995 fly
PIN 2555/661
 Diptera - Nadipteridae
Nadiptera pulchella n. sp.31 Lukashevich 1995 fly
PIN 2555/660
 Eoblattida - Megakhosaridae
Mesoblattogryllus intermedius n. gen. n. sp.33 Storozhenko 1990 insect
PIN 2555/984
Megakhosarodes paulivenosus n. sp.36 Storozhenko 1993 insect
PIN 2555/699
Metakhosara sharovi36 Storozhenko 1993 insect
PIN 2555/757, 779
Megablattogryllus austerus33 Storozhenko 1990 insect
PIN 2555/727
Megablattogryllus magister33 Storozhenko 1990 insect
PIN 2555/691
"Microblattogryllus variabilis n. gen. n. sp." = Protoblattogryllus variabilis33 Storozhenko 1990 insect
PIN 2555/744, 674, 770, 777, 830, 833
Protoblattogryllus asiaticus n. sp.33 Storozhenko 1990 insect
PIN 2555/786, 848
 Eoblattida - Mesorthopteridae
Parastenaropodites fluxa n. gen. n. sp.38 Storozhenko 1996 insect
PIN 2555/985,789,842,906,911
Parastenaropodites longiuscula38 Storozhenko 1996 insect
Sharovites alexanderi3 Aristov and Storozhenko 2013 insect
PIN 2555/729, 2781
Belmophenopterum rasnitsyni n. sp.40 Storozhenko and Aristov 2021 insect
PIN 2555/820
Mesoidelia ignorata n. gen. n. sp.38 Storozhenko 1996 insect
PIN 2555/676,991
Mesoidelia faceta38 Storozhenko 1996 insect
PIN 2555/686
Locustoblattina segmentata n. sp.2 Aristov 2017 insect
PIN 2555/2001
"Parastenaropodites nervosa" = Austroidelia nervosa38 Storozhenko 1996 insect
Austroidelia asiatica38 Storozhenko 1996 insect
 Eoblattida - Blattogryllidae
Baharellinus pectinatus n. sp.35 Storozhenko 1992 insect
PIN 2555/967
Baharellinus dimidiatus n. gen. n. sp.35 Storozhenko 1992 insect
PIN 2555/759, 807, 935
Dorniella primitiva n. sp.35 Storozhenko 1992 insect
PIN 2555/709(931)
Baharellus madygensis n. sp.35 Storozhenko 1992 insect
PIN 2555/722
Costatoviblatta aenigmatosa35 Storozhenko 1992 insect
PIN 2555/672, 680, 735, 739, 758, 762, 767, 771, 809, 844, 856, 858, 875, 881, 890, 901, 910, 915, 922, 933, 998, 999
Costatoviblatta similis n. sp.1 Aristov 2011 insect
 Reculida - Geinitziidae
Shurabia ferganensis26 Rasnitsyn 1982 winged insect
PIN 2555/787
Shurabia minuta26 Rasnitsyn 1982 winged insect
PIN 2555/792
Shurabia anomala n. sp.26 Rasnitsyn 1982 winged insect
PIN 2555/788
 Blattaria - Caloblattinidae
Thuringoblatta sogdianensis43 Vishnyakova 1998 cockroach
PIN 2555/135, 140, 142, 147-152, 154, 170
Sogdoblatta nana n. sp.43 Vishnyakova 1998 cockroach
PIN 2555/191, 213, 506
 Dicondylia - Permosialidae
"Perloblatta ordinata n. gen. n. sp." = Permonka triassica35, Permonka unica n. sp.25, Permosialis triassica n. sp.20
"Perloblatta ordinata n. gen. n. sp." = Permonka triassica35 Rasnitsyn 1977 winged insect
PIN 2555/723a
Permonka unica n. sp.25 Rasnitsyn 1977 winged insect
PIN 2555/1060
Permosialis triassica n. sp.20 Novokshonov and Zhuzhgova 2004 winged insect
PIN 2555/2066, 2067, 2068, 3582
 Neuroptera - Archeosmylidae
Osmylotriasia superba n. gen. n. sp.13
Osmylotriasia superba n. gen. n. sp.13 Khramov 2024 lacewing
PIN 2555/2096
 Coleoptera - Ademosynidae
Dolichosyne rostrata n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1735
Ademosyne kirghizica22 Ponomarenko 1969 beetle
PIN 2555/1696
Cephalosyne capitata22 Ponomarenko 1969 beetle
PIN 2555/1732
 Coleoptera - Triaplidae
Triaplus laticoxa4 Ponomarenko 1977 beetle
PIN 2555/1720
 Coleoptera - Phoroschizidae
Salebroferus asper n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1707
Salebroferus confragosus n. gen. n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1692
Catabrycus hoplites n. gen. n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1702
Triassocoleus tortulosus22 Ponomarenko 1969 beetle
PIN 2555/1708
Thnesidius xyphophorus22 Ponomarenko 1969 beetle
PIN 2555/1709
Schizophoroides rugosus22 Ponomarenko 1969 beetle
PIN 2555/1710
Hadeocoleus pelopius22 Ponomarenko 1969 beetle
PIN 2555/1701, 1719, 1699
Hadeocoleus catachtonius22 Ponomarenko 1969 beetle
PIN 2555/1704
Hadeocoleus gigas22 Ponomarenko 1969 beetle
PIN 2555/1698, 1700
 Coleoptera - Ommatidae
Lithocupes gigas n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1668
 Coleoptera -
"Notocupes rostratus n. sp." = Rhabdocupes rostratus22, "Notocupes tenuis" = Rhabdocupes tenuis22, "Notocupes laticella n. sp." = Rhabdocupes laticella22, Rhabdocupes baculatus n. sp.22
"Notocupes rostratus n. sp." = Rhabdocupes rostratus22 Ponomarenko 1969 beetle
PIN 2555/1664
"Notocupes tenuis" = Rhabdocupes tenuis22 Ponomarenko 1969 beetle
PIN 2555/1683,1685
"Notocupes laticella n. sp." = Rhabdocupes laticella22 Ponomarenko 1969 beetle
PIN 2555/1663, 1662, 1686
Rhabdocupes baculatus n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1666
 Coleoptera - Triadocupedidae
Cupesia sepulta n. sp.22
Cupesia sepulta n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1682, 1658
 Mecoptera - Mesopsychidae
"Mesopsyche justa" = Turbidapsyche justa19, "Mesopsyche gentica n. sp." = Turbidapsyche gentica19, "Mesopsyche shcherbakovi n. sp." = Turbidapsyche shcherbakovi17, Prochoristella longa16
"Mesopsyche justa" = Turbidapsyche justa19 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2555/1950
"Mesopsyche gentica n. sp." = Turbidapsyche gentica19 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2555/2013
"Mesopsyche shcherbakovi n. sp." = Turbidapsyche shcherbakovi17 Novokshonov 1997 scorpionfly
PIN 2555/2037
Prochoristella longa16 Novokshonov 1997 scorpionfly
PIN 2555/1956
 Mecoptera - Permochoristidae
Agetopanorpa consueta n. sp.16 Novokshonov 1997 scorpionfly
PIN 2555/1812
Liassochorista ? utilis16 Novokshonov 1997 scorpionfly
PIN 2555/1964
Mesageta pertrita n. sp.16 Novokshonov 1997 scorpionfly
PIN 2555/1927
Mesageta ignava n. sp.16 Novokshonov 1997 scorpionfly
PIN 2555/1890, 1867
 Mecoptera - Parachoristidae
Parachorista asiatica n. sp.15 Novokshonov 1997 scorpionfly
PIN 2555/2017
Parachorista sana n. sp.18 Novokshonov 2001 scorpionfly
PIN 2555/836
Parachorista arguta18 Novokshonov 2001 scorpionfly
PIN 2555/1989, 2009
Parachorista multivena n. sp.15 Novokshonov 1997 scorpionfly
PIN 2555/1941
Triassochorista kirgizica n. sp.15 Novokshonov 1997 scorpionfly
PIN 2555/1835, 2045, 2052
 Trichoptera - Cladochoristidae
Cladochorista multivenosa n. sp.42 Sukatsheva 1973 caddisfly
PIN 2555/174
 Trichoptera - Philopotamidae
Prophilopotamus asiaticus n. gen. n. sp.42
Prophilopotamus asiaticus n. gen. n. sp.42 Sukatsheva 1973 finger-net caddisfly
PIN 2555/1780
 Hymenoptera - Xyelidae
Madygenius primitivus n. sp.24, Triassoxyela orycta n. sp.24, Ferganoxyela sogdiana n. gen. n. sp.24, Ferganoxyela destructa n. sp.24, Oryctoxyela anomala n. gen. n. sp.24
Madygenius primitivus n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1755
Triassoxyela orycta n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1757
Ferganoxyela sogdiana n. gen. n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1752
Ferganoxyela destructa n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1753
Oryctoxyela anomala n. gen. n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1754
 Paoliida - Paoliidae
Ideliopsina nana39 Storozhenko 1996 winged insect
PIN 2555/687, 738, 846, 992
Ideliopsina ornata n. sp.39 Storozhenko 1996 winged insect
PIN 2555/895, 766
 Dicondylia - Subioblattidae
Subioblatta madygenica21 Papier and Nel 2001 winged insect
PIN 2555/76; 38, 39, 43, 46, 48, 51, 53, 59, 62, 66, 80, 81, 82, 86, 87, 89, 98, 100, 104 in Hinkelman 2022
 Embioptera - Alexarasniidae
Nestorembia sp.12 Shcherbakov 2015 webspinners
PIN 2555/1873
 Dicondylia - Xiphopteridae
Xiphopterum curvatum n. gen. n. sp.
Xiphopterum curvatum n. gen. n. sp. Sharov 1968 winged insect
PIN 2555/1554
 Dicondylia - Aeroplanidae
"Paraplana affinis" = Sharovoplana affinis
"Paraplana affinis" = Sharovoplana affinis Sharov 1968 winged insect
PIN 2555/1563
 Dicondylia - Prochresmodidae
Triassophasma minutissimum n. sp.8 Gorochov 1994 winged insect
PIN 2555/1614
"Prochresmoda brevipoda n. sp." = Triassophasma brevipoda Sharov 1968 winged insect
PIN 2555/1594
Triassophasma intermedium8 Gorochov 1994 winged insect
PIN 2555/1557, 1616, 1596
Prochresmoda minuta n. sp.8 Gorochov 1994 winged insect
PIN 2555/1611
 Plecoptera - Perlariopseidae
Tritaniella synneura32 Sinitshenkova 1987 stonefly
PIN 2555/1024, 1029
Tritaniella mera n. gen. n. sp.32 Sinitshenkova 1987 stonefly
PIN 2555/1036
Dicronemoura acaulis32 Sinitshenkova 1987 stonefly
PIN 2555/1014, 1016, 1020, 1022, 1028, 1031
Fritaniopsis brevicaulis n. gen. n. sp.32 Sinitshenkova 1987 stonefly
PIN 2555/1013, 1017
Fritaniopsis remota n. sp.32 Sinitshenkova 1987 stonefly
PIN 2555/1027, 1019
 Orthoptera - Xenopteridae
Axenopterum venosum n. gen. n. sp.10, Proxenopterum primitivum n. gen. n. sp.10, Pteroferganodes rieki n. gen. n. sp.7, Ferganopterus clarus, Ferganopterodes reductus n. gen. n. sp.
Axenopterum venosum n. gen. n. sp.10 Gorochov 2005 grasshopper
PIN 2555/1316
Proxenopterum primitivum n. gen. n. sp.10 Gorochov 2005 grasshopper
PIN 2555/1313, 1307, 1312, 1315
Pteroferganodes rieki n. gen. n. sp.7 Gorochov 1989 grasshopper
PIN 2555/2240
Ferganopterus clarus Sharov 1968 grasshopper
text says 121 imprints but doesn't specify collection
Ferganopterodes reductus n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1280
 Orthoptera - Mesoedischiidae
Mesoedischia madygenica Sharov 1968 grasshopper
PIN 2555/1250,1256,1249,1251,1252,1253,1254,1255,1257,1326
 Orthoptera - Proparagryllacrididae
Madygenia ovalis6 Gorochov 1987 grasshopper
PIN 2555/1342
Madygenia orientalis Sharov 1968 grasshopper
PIN 2555/1340
"Fergania reducta n. gen. n. sp." = Kashgarlimahmutia reducta Sharov 1968 grasshopper
PIN 2555/1445
Gryllacrimima perfecta Sharov 1968 grasshopper
 Orthoptera - Bintoniellidae
Provitimia pectinata n. gen. n. sp., "Provitimia ramivenosa n. sp." = Proshiella ramivenosa
Provitimia pectinata n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1420
"Provitimia ramivenosa n. sp." = Proshiella ramivenosa Sharov 1968 grasshopper
PIN 2555/1436
 Orthoptera - Tuphellidae
Tuphella rohdendorfi n. sp.5 Gorochov 1986 grasshopper
PIN 2555/1081
Tuphella sharovi n. sp.5 Gorochov 1986 grasshopper
PIN 2555/1159
 Orthoptera - Haglidae
Sonohagla chopardi n. sp.5, Sonohagla saussurei n. gen. n. sp.5, Archihagla tenuis n. sp.5, Voliopinae indet.5, Turkestania deviata n. gen. n. sp., Zavoliopus densus n. gen. n. sp.5, Stenovoliopus elongatus n. gen. n. sp.5, Platyvoliopus maximus n. gen. n. sp.5, Lyrohagla pravdini n. sp.5, Lyrohagla decipiens n. sp.5, Protshorkuphlebia similis n. sp.5, Protshorkuphlebia triassica n. gen. n. sp., Proisfaroptera martynovi, Hagloptera intermedia n. gen. n. sp.5, Eumaraga madygenica n. gen. n. sp.5, "Maraga tshorkuphlebioides n. sp." = Triassaga tshorkuphlebioides5, Microhagla minuta n. gen. n. sp.5, Dinohagla corrugata n. gen. n. sp.5, Dulcihagla beybienkoi n. gen. n. sp.5
Sonohagla chopardi n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1118
Sonohagla saussurei n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1102
Archihagla tenuis n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1128
Voliopinae indet.5 Gorochov 1986 hump-winged cricket
PIN 2555/1079
Turkestania deviata n. gen. n. sp. Sharov 1968 hump-winged cricket
PIN 2555/1142
Zavoliopus densus n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1148
Stenovoliopus elongatus n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1119
Platyvoliopus maximus n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1168
Lyrohagla pravdini n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1165
Lyrohagla decipiens n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1125
Protshorkuphlebia similis n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1092
Protshorkuphlebia triassica n. gen. n. sp. Sharov 1968 hump-winged cricket
PIN 2555/1088,1097
Proisfaroptera martynovi Sharov 1968 hump-winged cricket
PIN 2555/1114
Hagloptera intermedia n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1141
Eumaraga madygenica n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1085
"Maraga tshorkuphlebioides n. sp." = Triassaga tshorkuphlebioides5 Gorochov 1986 hump-winged cricket
PIN 2555/1099
Microhagla minuta n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1134
Dinohagla corrugata n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1137
Dulcihagla beybienkoi n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1150
 Orthoptera - Mesotitanidae
Mesotitanodes tillyardi n. gen. n. sp., "Prototitan similis n. sp." = Mesotitanodes similis, "Prototitan primitivus" = Mesotitan primitivus, Gigatitan similis n. sp., "Nanotitan extentus" = Gigatitan extensus, "Nanotitan magnificus n. gen. n. sp." = Gigatitan magnificus, Gigatitan ovatus n. sp.
Mesotitanodes tillyardi n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1524,1455
"Prototitan similis n. sp." = Mesotitanodes similis Sharov 1968 grasshopper
PIN 2555/1481,1521
"Prototitan primitivus" = Mesotitan primitivus Sharov 1968 grasshopper
PIN 2555/1473
Gigatitan similis n. sp. Sharov 1968 grasshopper
PIN 2555/1462
"Nanotitan extentus" = Gigatitan extensus Sharov 1968 grasshopper
PIN 2555/1513
"Nanotitan magnificus n. gen. n. sp." = Gigatitan magnificus Sharov 1968 grasshopper
PIN 2555/1509
Gigatitan ovatus n. sp. Sharov 1968 grasshopper
PIN 2555/1504,1461
 Orthoptera - Paratitanidae
Paratitan venosus n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1456
Paratitan modestus n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1479
Paratitan libelluloides n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1530
Paratitan intermedius n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1520
Paratitan latispeculum n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1478
 Orthoptera - Dzhajloutshellidae
Adzhajloutshella plana n. sp.11 Gorochov 2005 grasshopper
PIN 2555/1440
 Orthoptera - Locustavidae
Locustavus deformatus n. sp.11, Locustavus intermedius11, "Ferganopsis lanceolatus n. gen. n. sp." = Locustavus lanceolatus, Locustavus minutus11, Locustavus problematicus n. sp.11
Locustavus deformatus n. sp.11 Gorochov 2005 grasshopper
PIN 2555/1396
Locustavus intermedius11 Gorochov 2005 grasshopper
PIN 2555/1393
"Ferganopsis lanceolatus n. gen. n. sp." = Locustavus lanceolatus Sharov 1968 grasshopper
PIN 2555/1392
Locustavus minutus11 Gorochov 2005 grasshopper
PIN 2555/1399, 1406, 1417
Locustavus problematicus n. sp.11 Gorochov 2005 grasshopper
PIN 2555/1403
 Cnemidolestodea - Necrophasmidae
Ferganamadygenia plicata n. gen. n. sp.41
Ferganamadygenia plicata n. gen. n. sp.41 Storozhenko and Vršanský 1995 winged insect
PIN 2555/717
 Grylloblattodea -
Pseudoshurabia pallidula n. gen. n. sp.39
Pseudoshurabia pallidula n. gen. n. sp.39 Storozhenko 1996 ice crawler
PIN 2555/716
 Grylloblattodea - Gorochoviidae
Gorochovia individua n. gen. n. sp.37
Gorochovia individua n. gen. n. sp.37 Storozhenko 1994 ice crawler
PIN 2555/731,763,873,857
 Grylloblattodea - Madygenophlebiidae
Micromadygenophlebia obscura n. gen. n. sp.34 Storozhenko 1992 ice crawler
PIN 2555/826
Madygenophlebia primitiva n. sp.34 Storozhenko 1992 ice crawler
PIN 2555/718
Madygenophlebia bella n. gen. n. sp.34 Storozhenko 1992 ice crawler
PIN 2555/819,748,824,828
 Grylloblattodea -
Madygenidelia conjuncta n. gen. n. sp.39
Madygenidelia conjuncta n. gen. n. sp.39 Storozhenko 1996 ice crawler
PIN 2555/791
 Hemiptera - Saaloscytinidae
Tingiopsis reticulata28 Becker-Migdisova 1953 true bug
PIN 2555/2213
 Hemiptera - Naibiidae
Coccavus supercubitus n. gen. n. sp.27
Coccavus supercubitus n. gen. n. sp.27 Shcherbakov 2007 true bug
PIN 2555/2312, 2392, 2936
 Hemiptera - Palaeontinidae
Papiliontina spectans n. sp.30 Shcherbakov 2022 true bug
PIN 2555/2137
Papiliontina machaon n. sp.30 Shcherbakov 2022 true bug
PIN 2555/2314
Papiliontina dracomima n. gen. n. sp.30 Shcherbakov 2022 true bug
PIN 2555/2136
 Hemiptera - Maguviopseidae
Falcarta bella n. gen.28, Phyllotexta latens n. gen.28, Asiocula lima n. gen.28, Nevicia imitans n. gen.28
Falcarta bella n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2508
Phyllotexta latens n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2751
Asiocula lima n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2245
Nevicia imitans n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2382
 Hemiptera - Mesojabloniidae
Scytachile cf. emeljanovi n. gen.28, Fulgobole evansi n. gen. n. sp.28
Scytachile cf. emeljanovi n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2499
Fulgobole evansi n. gen. n. sp.28 Shcherbakov 2011 true bug
PIN 2555/2228,2762
 Hemiptera - Curvicubitidae
Beaconiella pulchra29 Shcherbakov 2021 true bug
PIN 2555/2360, 2368(2386), 2429, 2908, 2320
Beaconiella cincta29 Shcherbakov 2021 true bug
PIN 2555/2192(2866), 3514(3642)
 Dicondylia - Paurophlebiidae
Nonymophlebia venosa n. gen. n. sp.23, Cladophlebia parvula n. gen. n. sp.23, Cladophlebia brevis n. sp.23, Paurophlebia lepida n. gen. n. sp.23, Neritophlebia longa n. sp.23
Nonymophlebia venosa n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/649, 653
Cladophlebia parvula n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/617, 654
Cladophlebia brevis n. sp.23 Pritykina 1981 winged insect
PIN 2555/634
Paurophlebia lepida n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/624
Neritophlebia longa n. sp.23 Pritykina 1981 winged insect
PIN 2555/614
 Dicondylia - Triadophlebiidae
Triadophlebia magna n. sp.23 Pritykina 1981 winged insect
PIN 2555/635, 645
Triadophlebia honesta n. sp.23 Pritykina 1981 winged insect
PIN 2555/646
Triadophlebia madygenica n. sp.23 Pritykina 1981 winged insect
PIN 2555/639
Triadophlebia modica n. sp.23 Pritykina 1981 winged insect
PIN 2555/642
Triadophlebia minuta n. sp.23 Pritykina 1981 winged insect
PIN 2555/635
 Dicondylia - Mitophlebiidae
Mitophlebia enormis n. gen. n. sp.23
Mitophlebia enormis n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/641
 Dicondylia - Xamenophlebiidae
Xamenophlebia ornata14 Pritykina 1981 winged insect
PIN 2555/2074
 Dicondylia - Zygophlebiidae
Zygophlebiella curta n. gen. n. sp.23, Mixophlebia mixta n. gen. n. sp.14, Zygophlebia ramosa n. gen. n. sp.23
Zygophlebiella curta n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/633, 631
Mixophlebia mixta n. gen. n. sp.14 Pritykina 1981 winged insect
PIN 2555/625
Zygophlebia ramosa n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/619, 656
 Dicondylia - Kennedyidae
Kennedya carpenteri n. sp.23 Pritykina 1981 winged insect
PIN 2555/616
 Dicondylia -
Terskeja tenuis n. sp.23, Terskeja pumilio n. sp.23
Terskeja tenuis n. sp.23 Pritykina 1981 winged insect
PIN 2555/632
Terskeja pumilio n. sp.23 Pritykina 1981 winged insect
PIN 2555/630
 Odonata - Triassolestidae
Triassolestodes asiaticus n. gen. n. sp.23
Triassolestodes asiaticus n. gen. n. sp.23 Pritykina 1981 dragonfly
PIN 2555/637