Tuanwang, Laiyang City (Shandong Prov. Museum coll.) (Cretaceous of China)

Where: Shandong, China (36.8° N, 120.6° E)

• Paleocoordinates: 35.9° N, 133.2° E (Wright 2013)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Laiyang Formation, Aptian (121.4 - 113.2 Ma)

Environment/lithology: lacustrine - large; lithified shale

• Large (several hundred km2) lake in the Laiyang basin. In addition to the abundant entomofauna, plant fossil fragments, fish and a single bird feather also have been found. Water plants were apparently rare, and at the centre of the lake I have found no plant or animal fossils. Extensional regime in response to the progressive attenuation of the thickened lithosphere and orogenic collapse.
• The Third Member of the Laiyang Formation consists of 250-705 m of calcareous sandstone, siltstone and fine-granted sandstone intercalated with thin, paper-like, insect-rich shales.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collection methods: Repository: Shandong Provincial Museum

Primary reference: J. F. Zhang. 1992. Fossil Coleoptera from Laiyang, Shandong Province. Acta Entomologica Sinica 35:331-338 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 125196: authorized by Matthew Clapham, entered by Matthew Clapham on 06.03.2012

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names
Insecta
 Hemiptera - Palaeoaphididae
Caudaphis leptoneura n. sp.6, Caudaphis spinalis n. gen. n. sp.6
Caudaphis leptoneura n. sp.6 Zhang et al. 1989 aphid
L84117
Caudaphis spinalis n. gen. n. sp.6 Zhang et al. 1989 aphid
L84116
 Hemiptera - Oviparosiphidae
"Mesoviparosiphum tuanwangense n. gen. n. sp." = Archeoviparosiphum tuanwangense6, "Paroviparosiphum camptotropum n. sp." = Archeoviparosiphum camptotropum6, "Mesoviparosiphum malacum n. sp." = Archeoviparosiphum malacum6, "Paroviparosiphum opimum n. gen. n. sp." = Archeoviparosiphum opimum6
"Mesoviparosiphum tuanwangense n. gen. n. sp." = Archeoviparosiphum tuanwangense6 Zhang et al. 1989 aphid
L84121
"Paroviparosiphum camptotropum n. sp." = Archeoviparosiphum camptotropum6 Zhang et al. 1989 aphid
L84129
"Mesoviparosiphum malacum n. sp." = Archeoviparosiphum malacum6 Zhang et al. 1989 aphid
L84122
"Paroviparosiphum opimum n. gen. n. sp." = Archeoviparosiphum opimum6 Zhang et al. 1989 aphid
L84126
 Hemiptera - Sinaphididae
Tartaraphis peregrina n. gen. n. sp.6
Tartaraphis peregrina n. gen. n. sp.6 Zhang et al. 1989 aphid
L84124(5)
 Hemiptera - Saldidae
Mesolygaeus laiyangensis4
Mesolygaeus laiyangensis4 Ping 1928 shore bug
L88232, 88240, 88242
 Hemiptera - Archegocimicidae
Schizopteryx lacustris4, Schizopteryx shandongensis4
Schizopteryx lacustris4 Zhang 1991 shore bug
L88257
Schizopteryx shandongensis4 Hong 1984 shore bug
L88245, 88246, 88247, 88248, 88249, 88253
 Hemiptera - Archijassidae
Mesoccus advenus n. sp.1, Mesoccus lutarius n. gen. n. sp.1
Mesoccus advenus n. sp.1 Zhang 1985 true bug
L85103
Mesoccus lutarius n. gen. n. sp.1 Zhang 1985 true bug
L85101, 85104
 Hymenoptera - Megalyridae
Stemmogaster celata n. gen. n. sp.1
Stemmogaster celata n. gen. n. sp.1 Zhang 1985 wasp
L85113
 Hymenoptera -
"Oligoneuroides huadongensis n. gen. n. sp." = Rasnitsynia huadongensis1
"Oligoneuroides huadongensis n. gen. n. sp." = Rasnitsynia huadongensis1 Zhang 1985 wasp
L85115
 Hymenoptera - Angarosphecidae
"Shandongodes lithodes n. gen. n. sp." = Angarosphex lithodes1, "Mataeosphex venulosus n. gen. n. sp." = Angarosphex venulosus1
"Shandongodes lithodes n. gen. n. sp." = Angarosphex lithodes1 Zhang 1985 apoid wasp
L85110
"Mataeosphex venulosus n. gen. n. sp." = Angarosphex venulosus1 Zhang 1985 apoid wasp
L85112-1
 Hymenoptera -
Shandongodes necrodes n. sp.1
Shandongodes necrodes n. sp.1 Zhang 1985 winged insect
L85111
 Hymenoptera - Tenthredinidae
Palaeathalia laiyangensis n. gen. n. sp.1
Palaeathalia laiyangensis n. gen. n. sp.1 Zhang 1985 wood wasp
L85121
 Diptera - Rhagionidae
Pseudopetia exilis n. sp.2
Pseudopetia exilis n. sp.2 Zhang 1987 snipe fly
L84135
 Diptera - Platypezidae
Mesopetia tuanwangensis n. gen. n. sp.2
Mesopetia tuanwangensis n. gen. n. sp.2 Zhang 1987 flat-footed fly
L84131
 Diptera - Pediciidae
"Palaeolimnobia laiyangensis n. gen. n. sp." = Zhangobia laiyangensis5
"Palaeolimnobia laiyangensis n. gen. n. sp." = Zhangobia laiyangensis5 Zhang et al. 1986 hairy-eyed cranefly
L84101(2)
 Diptera - Chironomidae
Oryctochlus contiguus n. sp.3
Oryctochlus contiguus n. sp.3 Zhang 1991 non biting midge
L86812, 86813
 Diptera - Chaoboridae
Mesochaoborus zhangshanyingensis5, Mesochaoborus pallens n. sp.5, "Petiolatendipes shouchangensis" = Chironomaptera vesca5
Mesochaoborus zhangshanyingensis5 Hong 1984 phantom midge
L84112
Mesochaoborus pallens n. sp.5 Zhang et al. 1986 phantom midge
L84113
"Petiolatendipes shouchangensis" = Chironomaptera vesca5 Kalugina 1980 phantom midge
L84104, L84114
 Trichoptera -
Tuanwangica aethoneura n. gen. n. sp.1
Tuanwangica aethoneura n. gen. n. sp.1 Zhang 1985 caddisfly
L85106
 Coleoptera - Coptoclavidae
Coptoclavisca grandioculus n. sp.
Coptoclavisca grandioculus n. sp. Zhang 1992 beetle
L85433
 Orthoptera - Locustopsidae
Pseudoacrida cf. costata1
Pseudoacrida cf. costata1 Lin 1982 grasshopper
L84985(84988)
 Orthoptera -
Falsirameus ravus n. gen. n. sp.1
Falsirameus ravus n. gen. n. sp.1 Zhang 1985 grasshopper
L84991(2)
1Zhang 1985; 2Zhang 1987; 3Zhang 1991; 4Zhang 1991; 5Zhang et al. 1986; 6Zhang et al. 1989