PBDB Collection

Swain Quarry (Paleocene of the United States)

Also known as CM loc 217

Where: Carbon County, Wyoming (41.3° N, 107.7° W: paleocoordinates 47.7° N, 86.2° W)

• coordinate stated in text

• small collection-level geographic resolution

When: Fort Union Formation, Torrejonian (63.8 - 60.9 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: terrestrial; lithology not reported

Size classes: macrofossils, mesofossils

Collection methods: surface (in situ),

Primary reference: R. M. West. 1970. Tetraclaenodon puercensis (Mammalia: Phenacodontidae), Goler Formation, Paleocene of California, and distribution of the genus. Journal of Paleontology 44(5):851-857 [J. Alroy/J. Alroy/M. Carrano]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 14886: authorized by John Alroy, entered by John Alroy on 18.02.1993, edited by Roger Benson, Terri Cleary and Patricia Holroyd

Creative Commons license: CC BY (attribution)

Taxonomic list

• there are seven other localities in the same section, but none of the material is described

•Mesodma pygmaea, i.e. sp. P, not listed, but mentioned by Krause 1977

Show authors, comments, and common names

Amphibia
Proteida - Batrachosauroididae	Opisthotriton kayi⁵ Opisthotriton kayi⁵ Auffenberg 1961 salamander
Caudata - Urodela	Habrosaurus sp.⁵, Scapherpeton tectum⁵ Habrosaurus sp.⁵ Gilmore 1928 siren Scapherpeton tectum⁵ Cope 1876 salamander
Mammalia
Multituberculata - Ptilodontidae	Ptilodus mediaevus⁹, Baiotomeus douglassi⁸ Ptilodus mediaevus⁹ Cope 1881 multituberculate Baiotomeus douglassi⁸ Simpson 1935 multituberculate one specimen of "Ptilodus mediaevus"
Multituberculata - Neoplagiaulacidae	Xanclomys mcgrewi n. sp.⁹, "Ectypodus sylviae n. sp." = Parectypodus sylviae⁹, "Parectypodus clemensi" = Krauseia clemensi¹², Xyronomys swainae n. sp.⁹, Ectypodus szalayi¹², Neoplagiaulax nelsoni¹³ Xanclomys mcgrewi n. sp.⁹ Rigby 1980 multituberculate "Ectypodus sylviae n. sp." = Parectypodus sylviae⁹ Rigby 1980 multituberculate "Parectypodus clemensi" = Krauseia clemensi¹² Sloan 1981 multituberculate may include Rigby's new "Ectypodus sp.," said to be the same as the Bangtail species by Gingerich et al. 1983, who do not cite Sloan Xyronomys swainae n. sp.⁹ Rigby 1980 multituberculate Ectypodus szalayi¹² Sloan 1981 multituberculate Neoplagiaulax nelsoni¹³ Sloan 1987 multituberculate N. "cf. hunteri"
Proteutheria - Palaeoryctidae	Palaeoryctes puercensis⁹ Palaeoryctes puercensis⁹ Matthew 1913 eutherian
Leptictida - Leptictidae	Myrmecoboides montanensis⁹, "Acmeodon hyoni n. sp." = Acmeodon secans⁹, "Diacodon concordiarcensis" = Prodiacodon concordiarcensis⁹ Myrmecoboides montanensis⁹ Gidley 1915 eutherian "Acmeodon hyoni n. sp." = Acmeodon secans⁹ Matthew and Granger 1921 eutherian "Diacodon concordiarcensis" = Prodiacodon concordiarcensis⁹ Simpson 1935 eutherian
Macroscelidea - Apheliscidae	Haplaletes disceptatrix⁹, Litomylus dissentaneus⁹ Haplaletes disceptatrix⁹ Simpson 1935 elephant shrew Litomylus dissentaneus⁹ Simpson 1935 elephant shrew
Macroscelidea -	"Mckennatherium martinezi n. sp." = Adunator martinezi⁹, "Mckennatherium fredericki n. sp." = Adunator fredericki⁹ "Mckennatherium martinezi n. sp." = Adunator martinezi⁹ Rigby 1980 elephant shrew "Mckennatherium fredericki n. sp." = Adunator fredericki⁹ Rigby 1980 elephant shrew
Insectivora -	Insectivora indet.¹ Insectivora indet.¹ Bowdich 1821 insectivoran
Tribosphenida - Mixodectidae	Mixodectes sp.⁹, Mixodectes malaris⁹ Mixodectes sp.⁹ Cope 1883 placental Mixodectes malaris⁹ Cope 1884 placental confirmed by Gunnell 1989
Primates - Picrodontidae	Picrodus silberlingi⁹ Picrodus silberlingi⁹ Douglass 1908 primate
Primates - Palaechthonidae	"Palaechthon problematicus" = Plesiolestes problematicus⁹, Palenochtha weissae n. sp.⁹, Palenochtha minor⁶ "Palaechthon problematicus" = Plesiolestes problematicus⁹ Jepsen 1930 primate Palenochtha weissae n. sp.⁹ Rigby 1980 primate Palenochtha minor⁶ Gidley 1923 primate
Primates - Paromomyidae	Paromomys depressidens⁹, Paromomys maturus⁹ Paromomys depressidens⁹ Gidley 1923 primate confirmed by Gunnell 1989 Paromomys maturus⁹ Gidley 1923 primate confirmed by Gunnell 1989
Tribosphenida - Pantolestoidae	Bessoecetor septentrionalis¹¹, Pantolestinae indet.⁹ Bessoecetor septentrionalis¹¹ Russell 1929 placental Pantolestinae indet.⁹ Simpson 1937 placental
Tribosphenida -	"Prodiacodon cf. puercensis" = Diacodon puercensis⁹ "Prodiacodon cf. puercensis" = Diacodon puercensis⁹ Matthew 1918 placental
Soricomorpha - Nyctitheriidae	Leptacodon cf. tener⁹ Leptacodon cf. tener⁹ Matthew and Granger 1921 placental
Acreodi - Triisodontidae	Goniacodon levisanus⁹ Goniacodon levisanus⁹ Cope 1883 placental
Acreodi - Mesonychidae	Dissacus navajovius⁹ Dissacus navajovius⁹ Cope 1881 condylarth
Acreodi - Oxyclaenidae	Prothryptacodon cf. furens⁹, Prothryptacodon hilli n. sp.⁹ Prothryptacodon cf. furens⁹ Simpson 1935 placental Prothryptacodon hilli n. sp.⁹ Rigby 1980 placental
Tribosphenida - Periptychidae	Periptychus carinidens⁹, Haploconus sp.⁹, Anisonchus sectorius⁹, Anisonchus willeyi n. sp.⁹ Periptychus carinidens⁹ Cope 1881 condylarth Haploconus sp.⁹ Cope 1882 condylarth Anisonchus sectorius⁹ Cope 1881 condylarth Anisonchus willeyi n. sp.⁹ Rigby 1980 condylarth
Procreodi - Arctocyonidae	Chriacus pelvidens⁹, "Mimotricentes subtrigonus" = Tricentes subtrigonus⁹, Arctocyon ferox⁹, "Protogonodon criswelli n. sp." = Loxolophus criswelli⁹ Chriacus pelvidens⁹ Cope 1881 condylarth "Mimotricentes subtrigonus" = Tricentes subtrigonus⁹ Cope 1881 condylarth Arctocyon ferox⁹ Cope 1883 condylarth "Protogonodon criswelli n. sp." = Loxolophus criswelli⁹ Rigby 1980 condylarth
Tribosphenida - Phenacodontidae	Tetraclaenodon puercensis⁹ Tetraclaenodon puercensis⁹ Cope 1881 condylarth
Ferae - Viverravidae	"Bryanictis vanvaleni" = Intyrictis vanvaleni⁹, "Protictis haydenianus" = Protictis haydenianus⁹, "Prolimnocyon macfaddeni n. sp." = Protictis haydenianus⁹, Simpsonictis jaynanneae n. sp.⁹ "Bryanictis vanvaleni" = Intyrictis vanvaleni⁹ MacIntyre 1966 placental "Protictis haydenianus" = Protictis haydenianus⁹ Cope 1882 placental "Prolimnocyon macfaddeni n. sp." = Protictis haydenianus⁹ Cope 1882 placental macfaddeni Simpsonictis jaynanneae n. sp.⁹ Rigby 1980 placental
Condylarthra - Hyopsodontidae	Litaletes sp.¹⁰, Promioclaenus acolytus⁹ Litaletes sp.¹⁰ Simpson 1935 condylarth said to be new and not the same as "L. mantiensis", a jun. syn. of L. disjunctus Promioclaenus acolytus⁹ Cope 1882 condylarth
Placentalia -	Deltatheridia indet.⁹ Deltatheridia indet.⁹ placental
Cimolesta - Apatemyidae	Jepsenella praepropera⁹ Jepsenella praepropera⁹ Simpson 1940 eutherian
Cimolesta - Cimolestidae	Gelastops joni n. sp.⁹, Paleotomus milleri n. sp.⁹ Gelastops joni n. sp.⁹ Rigby 1980 eutherian Paleotomus milleri n. sp.⁹ Rigby 1980 eutherian
Taeniodonta - Stylinodontidae	Psittacotherium multifragum⁹ Psittacotherium multifragum⁹ Cope 1882 taeniodont
Tribosphenida - Herpetotheriidae	Swaindelphys cifellii n. sp.⁷ Swaindelphys cifellii n. sp.⁷ Johanson 1996 marsupial
Reptilia
Squamata -	Squamata indet.¹⁴ Squamata indet.¹⁴ squamates "indeterminate Sauria" AMNH 12041, ventral part of basicranium; AMNH 12046, 14301, 14322, 14323, 14324, 14325, 14326, 15895, 15896, 15897, 15903, 15904, 15906, 15921, 15922, 15924, 15927, 15930, 15941, 15942, 15944, 15948, 15968, 15997, 15998, 16005, 16009, 16013, 16016, 16018, 16020, 16154, 16158, 16160, 16163, 16166, 16167, 16168, 16171, 16172, 16173, 16174, 16177, 16178, 16179, 16180, 16181, 16183, 16188, 16189, 16190, 16192, 16193, (?) 16194, 16196, 16197, 16198, 16216, 16217, 16218, partial dentaries; AMNH 15932, 15934, 15939, (?) 15946, 15947, 15950, 15964, 16025, partial maxillae; AMNH 15943, 15954, 16195, tooth-bearing fragments; AMNH 15958, proximal end of femur; AMNH 15962, trunk vertebra; AMNH 16056, distal end of humerus; AMNH 16137, caudal vertebra.
Squamata - Iguanidae	Swainiguanoides milleri n. gen. n. sp.¹⁴ Swainiguanoides milleri n. gen. n. sp.¹⁴ Sullivan 1982 squamates AMNH 12082 (holotype), medial part of right dentary; 12081, medial part of right dentary; 15902, medial part of left dentary; 15933, medial part of left dentary; 12048, medial part of left maxilla
Squamata - Xantusiidae	Palaeoxantusia fera¹⁴, Palaeoxantusia fera⁴ Palaeoxantusia fera¹⁴ Hecht 1956 night lizard AMNH 12028, posterior part of right maxilla; AMNH 12036, posterior part of left spleniodentary; AMNH 12038, posterior half of left spleniodentary; AMNH 12039, anterior part of right maxilla; AMNH 12040, left spleniodentary; AMNH 12041, posterior part of right spleniodentary; AMNH 12049, medial part of left spleniodentary; AMNH 12064, nearly complete left spleniodentary; AMNH 12071, anterior part of right spleniodentary; AMNH 12072, anterior part of right spleniodentary; AMNH 12073, posterior part of right spleniodentary; AMNH 12074, posterior part of right spleniodentary; AMNH 12075, left spleniodentary; AMNH 12076, nearly complete right spleniodentary; AMNH 12077, posterior part of right spleniodentary; AMNH 12079, anterior part of right spleniodentary; AMNH 12080, posterior part of right spleniodentary; AMNH 12083 (Text-fig. 6), posterior part of left maxilla; AMNH 14302, medial part of right spleniodentary; AMNH 14312, 4 teeth; AMNH 14313, medial part of right spleniodentary; AMNH 14314, posterior part of right maxilla; AMNH 15907, medial part of left spleniodentary; AMNH 15917, medial part of left spleniodentary; AMNH 15920, greater anterior part of left spleniodentary; AMNH 15926, posterior part of left maxilla; AMNH 15928, medial part of left spleniodentary; AMNH 15 929, greater anterior part of right spleniodentary; AMNH 15938, posterior part of left maxilla; AMNH 15949, posterior part of right spleniodentary; AMNH 15952, posterior part of right maxilla; AMNH 15953, medial part of left spleniodentary; AMNH 16007, nearly complete left spleniodentary; AMNH 16156, medial part of left spleniodentary; AMNH 16157, anterior part of left spleniodentary; AMNH 16170, anterior part of left spleniodentary. Also referred tentatively: AMNH 12043, medial part of right spleniodentary; AMNH 12059, medial part of right spleniodentary; AMNH 15925, nearly complete left maxilla; AMNH 15955, incomplete medial part of right spleniodentary; AMNH 16049, spleniodentary fragment; AMNH 16159, medial part of left spleniodentary; AMNH 16182, incomplete posterior part of right maxilla. Palaeoxantusia fera⁴ Hecht 1956 night lizard scincid
Squamata - Necrosauridae	cf. Provaranosaurus sp.¹⁴ cf. Provaranosaurus sp.¹⁴ Gilmore 1942 squamates AMNH 14306, medial part of left dentary; AMNH 14307, medial part of left dentary; AMNH 14309, medial part of right dentary
Squamata -	cf. Saniwa sp.¹⁴, cf. Saniwa sp.⁴ cf. Saniwa sp.¹⁴ Leidy 1870 squamates AMNH 15960, trunk vertebra; AMNH 16003, incomplete trunk vertebra. cf. Saniwa sp.⁴ Leidy 1870 squamates AMNH 15957, medial part of left dentary with 3 tooth bases.
Squamata - Anguidae	Anguidae indet.¹⁴, Odaxosaurus sp.², Odaxosaurus piger¹⁴, Odaxosaurinae indet.¹⁴, Glyptosaurinae indet.¹⁴, Machaerosaurus torrejonensis¹⁴ Anguidae indet.¹⁴ Gray 1825 squamates AMNH 12052, 12053, 12056, 12057, 12058, 12093, partial dentaries. Odaxosaurus sp.² Gilmore 1928 squamates CM 87168 Odaxosaurus piger¹⁴ Gilmore 1928 squamates AMNH 12030, posterior part of right dentary; AMNH 12032, right dentary; AMNH 12033, medial part of right dentary; AMNH 12035, posterior part of right dentary; AMNH 12060, medial part of left dentary; AMNH 12084, medial part of left dentary; AMNH 12085, posterior part of left dentary; AMNH 12086, posterior part of left maxilla; AMNH 14310, medial part of left dentary; AMNH 15900, posterior part of left maxilla; AMNH 15901, posterior part of right maxilla; AMNH 15940, posterior part of right dentary. Also tentatively referred: AMNH 15905, posterior part of right dentary; AMNH 15918, medial part of right dentary. Odaxosaurinae indet.¹⁴ Sullivan 1982 squamates AMNH 12089, 21 body osteoderms; AMNH 15910, posterior part of right maxilla; AMNH 15916, right maxilla fragment with tooth; AMNH 15937, posterior part of right maxilla; AMNH 15956, posterior part of left maxilla; AMNH 16004, medial part of right dentary; AMNH 16011, medial part of right dentary; AMNH 16012, anterior part of right dentary; AMNH 16109, frontal fragment; AMNH 16124 and 16125, body osteoderms, associated with frontal fragments TEXT-FIG. 11-Glyptosaurinae, genus and species indeterminate (AMNH 12088), right pterygoid (palatal view), x10. and garscales; AMNH 16176, maxilla fragment; AMNH 16209, (?) cheek osteoderm. Glyptosaurinae indet.¹⁴ Marsh 1872 squamates AMNH 12031, posterior part of right maxilla; AMNH 12042, medial part of left dentary; AMNH 12044, premaxilla; AMNH 12045, anterior part of left dentary; AMNH 12065, medial part of right dentary; AMNH 12070, anterior part of right maxilla; AMNH 12088, tooth bearing region of right pterygoid; AMNH 12091, medial part of (?) right maxilla Machaerosaurus torrejonensis¹⁴ Gilmore 1928 squamates AMNH 12047, medial part of left dentary; AMNH 12063, posterior part of right maxilla; AMNH 12066, medial part of right dentary; AMNH 12068, anterior part of right maxilla; AMNH 12087, left frontal; AMNH 12094, medial part of left dentary; AMNH 12095, posterior part of right maxilla; AMNH 12097, anterior part of left maxilla; AMNH 12099, nearly complete right dentary; AMNH 14311, posterior part of right dentary; AMNH 15911, left dentary; AMNH 15914, posterior part of left dentary; AMNH 16100, nearly complete right frontal. Also tentatively referred: AMNH 12034, posterior part of left maxilla; AMNH 12050, greater part of left dentary; AMNH 12051, posterior part of right dentary; AMNH 12055, medial part of left dentary; AMNH 12092, posterior part of right dentary; AMNH 14303, greater part of right dentary; AMNH 14327, posterior part of right maxilla; AMNH 15898, anterior part of left dentary; AMNH 15899, left maxilla; AMNH 15908, posterior part of a left maxilla; AMNH 15909, greater part of right maxilla; AMNH 15913, posterior part of left maxilla; AMNH 15931, posterior part of left dentary; AMNH 15935, anterior part of right maxilla; AMNH 15936, maxilla fragment; AMNH 15951, posterior part of right maxilla; AMNH 16008, posterior part of left dentary; AMNH 16010, posterior part of right maxilla.
Squamata - Xenosauridae	Exostinus cf. lancensis¹⁴ Exostinus cf. lancensis¹⁴ Gilmore 1928 squamates AMNH 14315, medial part of right dentary; AMNH 14316, medial part of left dentary; AMNH 14317, medial part of left dentary; AMNH 14318, anterior part of left dentary.
Testudinata - Baenidae	"Plesiobaena putorius" = Cedrobaena putorius³ "Plesiobaena putorius" = Cedrobaena putorius³ Gaffney 1972 turtle AMNH 4950, anterior portion of skull roof; 4949, posterior portion of skull roof

¹Alroy 2002; ²ElShafie 2024; ³Gaffney 1972; ⁴Gao and Fox 1996; ⁵Gardner and DeMar 2013; ⁶Gunnell 1989; ⁷Johanson 1996; ⁸Krause 1987; ⁹Rigby 1980; ¹⁰Robison 1986; ¹¹Scott et al. 2002; ¹²Sloan 1981; ¹³Sloan 1987; ¹⁴Sullivan 1982