Bardenas Reales Barranco del Fraile 1 (BF1) (Miocene to of Spain)

Where: Navarre, Spain (42.1° N, 1.5° W: paleocoordinates 42.0° N, 3.2° W)

• coordinate estimated from map

• small collection-level geographic resolution

When: MN2b-3 mammal zone, Tudela Formation, Aquitanian to Aquitanian (23.0 - 16.0 Ma)

• Late Aquitanian-Early Burdigalian

•Larrasoaña et al. 2006: fossil localities of the Tudela Formation assigned to the "Ramblian" continental stage. The lower boundary of the Ramblian is placed at the lower part of chron C6r (ca. 20.4 Ma), and the boundary between the lower and upper Ramblian is located at the middle of chron C6n (ca. 19.6Ma). The upper boundary of the Ramblian, and hence the base of the Aragonian, is placed at the upper middle part of chron C5Cr (16.8–17Ma). A revised calibration of European mammal zone MN3 suggest the lower boundary of zone MN3 is placed at the boundary between chrons C6r and C6n (ca. 20.1Ma), and its upper boundary is located at the upper middle part of chron C5Cr (16.8–17Ma).

• bed-level stratigraphic resolution

Environment/lithology: mire or swamp; gray, blue marl

• The lithological features and sedimentary structures observed in the Bardenas sites suggest that the vertebrate remains were deposited in the distal areas of an alluvial system. The presence of mudcracks and bioturbation traces in limestones suggests a shallow, not permanent water environment. This is indicative of a wetland.
• Blue to gray marls

Size class: macrofossils

Primary reference: X. Murelaga, X. Pereda Xuberbiola, F. Lapparent Broin, J. C. Rage, S. Duffaud, H. Astibia, and A. Badiola. 2002. Amphibians and reptiles from the Early Miocene of the Bardenas Reales of Navarre (Ebro Basin, Iberian Peninsula). Geobios 35:347-365 [R. Butler/J. Benito Moreno/T. Cleary]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 153751: authorized by Philip Mannion, entered by Philip Mannion on 13.01.2014, edited by Juan Benito Moreno

Creative Commons license: CC BY (attribution)

Taxonomic list

 Squamata - Colubridae
Colubridae indet. Oppel 1811 colubrid snake
BF1-156, 1 caudal vertebra
 Squamata - Boidae
? Eryx sp. Daudin 1803 true sand boa
BF-137 to 155, 19 trunk vertebrae
 Squamata - Anguidae
Ophisaurus sp. Daudin 1803 glass lizard
BF1-123, 1 trunk vertebra, BF-133 to 135, 3 caudal vertebrae, BF-136, 1 osteoscute
 Crocodylia -
Diplocynodon sp.2 Pomel 1847 crocodilian
BF1 material
 Testudines - Geoemydidae
Ptychogaster (Temnoclemmys) bardenensis Murelaga et al. 1999 turtle
 Testudines - Chelydridae
Chelydropsis apellanizi n. sp.1 Murelaga et al. 1999 turtle
BF-58, left hyoplastron, BF-1:31, posterior part of left epiplastron; BF-64, right hypoplastron; BF-11.3. right xiphiplastron; BF-20, right peripheral 9: BF-19, left peripheral 10: BF-21, pygal: BF-23. neural 1. BF-74, odd neural.
 Testudines - Trionychidae
Trionychinae indet. softshell turtle
 Caudata - Urodela
Salamandridae indet. Gray 1825 newt
1 trunk vertebra (BF1-159)
 Salientia -
Anura indet. Fischer von Waldheim 1813 frog
1 ilium (BF1-160), 1 urostyle (BF1-161)