Shapaja (TAR-20) (Oligocene to of Peru)

Also known as TAR 20

Where: Peru (6.6° S, 76.3° W)

• Paleocoordinates: 6.6° S, 65.7° W (Wright 2013)

• coordinate stated in text

• small collection-level geographic resolution

When: Deseadan mammal zone, Pozo Shales Member (Pozo Formation), Rupelian to Rupelian (33.9 - 23.0 Ma)

• Campbell et al. (2021). Our BTD results are also inconsistent with the latest Eocene and earliest Oligocene ages that Antoine et al. (2021) have proposed for the Shapaja localities. Though our analyses do not correctly predict the sequence of the Shapaja localities, the mean estimates for individual species nevertheless range from 31.6 to 20.7 Ma, with composite means for the section ranging from 29.1 to 28.3 Ma. The composite mean estimates for the Shapaja section are all very close to those for Santa Rosa, differing by only 0.42 to 1.0 My across analyses with both broad (effectively “unknown”) and more restricted age priors. These results are consistent with the close correlation of the Shapaja and Santa Rosa sections originally proposed by Boivin et al. (2018). Antoine et al. (2021) note that their placement of the Shapaja localities near the Eocene–Oligocene boundary implies that these faunas include 14 new first appearances in South America (either at the genus, family, or superfamily level), with most of the taxa in question otherwise first known from Deseadan (Upper Oligocene) localities. With the geochronological evidence now available from Santa Rosa, and the biochronological evidence for a close correlation of the Shapaja and Santa Rosa sections, we propose that a more parsimonious explanation for the occurrence of otherwise Deseadan taxa in the Shapaja localities is that those localities are, in fact, Deseadan in age. Given such an age estimate, we suggest that the two positive δ13C excursions identified in the Shapaja chemostratigraphies would be better correlated with those that occur at ∼26.5 and ∼25.5 Ma in the generalized global record (ref. 28, their figure 28.11).

• bed-level stratigraphic resolution

Environment/lithology: fluvial-deltaic; conglomerate

• from Antoine et al. 2021: a marine/brackish influence can be hypothesized in the middle part of the section (between TAR-20 and TAR-22) thanks to the conspicuous presence of myliobatid and pristid selachians (bat rays and sawfish, respectively). These taxa co-occur with a freshwater ichthyo- fauna dominated by characiform actinopterygians and dipnoans, thus pointing to an estuarine environment. The presence of neonate/young batoid specimens at TAR-20, TAR-13, and TAR-72 might further support brackish conditions for the concerned interval, as many Myliobatis females enter estuaries and coastal plains to give birth today (Mianzan et al., 2001). Accordingly, we can hypothesize that the concerned set- tings were close to such protected environments playing a nursery role for bat rays (even if the occurrence of a minute euryhaline/freshwater myliobatid species of unknown affinities cannot be fully discarded).
• Yellow-greenish microconglomerate with gypsum and plant remains.

Size classes: macrofossils, mesofossils

Collected in 2014

Primary reference: M. Boivin, L. Marivaux, F. Pujos, R. Salas-Gismondi, J. V. Tejada-Lara, R. M. Varas-Malca, and P.-O. Antoine. 2018. Early Oligocene caviomorph rodents from Shapaja, Peruvian Amazonia. Palaeontographica Abteilung A 311(1-6):87-156 [P. Mannion/P. Mannion]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 199560: authorized by Philip Mannion, entered by Philip Mannion on 31.01.2019, edited by Grace Varnham

Creative Commons license: CC0 (CC0)

Taxonomic list

unclassified
  -
Malacostraca
 Decapoda - Trichodactylidae
Trichodactylidae indet. Milne-Edwards 1853 crab
claw and shell fragments
Gastropoda
  -
Gastropoda indet. Cuvier 1795 snail
turriform snail inner cast
 Heterostropha - Planorbidae
Helisoma sp. Swainson 1940 snail
inner cast
Actinopteri
 Teleostei -
 Characiformes - Cynodontidae
cf. Hydrolycus sp. Lundberg 1997
teeth
 Characiformes - Anostomidae
Osteichthyes
 Ceratodontiformes - Lepidosirenidae
Lepidosirenidae indet. Stirton 1953 Australian lungfish
tooth plates
Amphibia
 Salientia -
Anura indet. frog
humerus and radio-ulna
Reptilia
 Crocodylia - Alligatoridae
Caimaninae "indet. new genus new species" Brochu 1999 crocodilian
new genus and new species, skull 45cm long
Caimaninae indet. Brochu 1999 crocodilian
 Crocodylia -
Gavialoidea indet. Brochu 1997 crocodilian
Mammalia
 Pyrotheria -
Pyrotheria indet. Ameghino 1895 placental
 Notoungulata -
Toxodontia indet. Ameghino 1887 notoungulate
 Rodentia -
 Rodentia - Agoutidae
Eoincamys valverdei Boivin et al. 2018 caviomorph
MUSM 3334-3336
Eoincamys parvus Boivin et al. 2018 caviomorph
MUSM 3337
 Rodentia -
cf. Tarapotomys sp. Boivin et al. 2018 caviomorph
MUSM 3327-3333
Tarapotomys subandinus Boivin et al. 2018 caviomorph
 Metatheria -
 Polydolopimorphia -
Argyrolagoidea indet. Ameghino 1904 metatherian
 Polydolopimorphia - Argyrolagidae
cf. Proargyrolagus sp. Wolff 1984 metatherian
Chondrichthyes
 Myliobatiformes - Myliobatidae
Myliobatis sp. Cuvier 1816 eagle ray
one neonate tooth
unclassified
  -
Plantae indet. Haeckel 1866
Silicified wood
Angiospermae
 Vitales - Vitaceae
Vitaceae indet. Jussieu 1789
seeds
 Poales - Poaceae
Poaceae indet. Barnhart 1895 grass
seeds
Magnoliopsida
  -
Magnoliopsida indet. Cronquist et al. 1996
PZ5 leaves
Charophyceae
  -
Charophyceae indet. Smith 1938
oogonia