PBDB

Bore Lullingen 1, depth 21m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Dingdener stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61895: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Foraminifera
Planorbulinacea - Cibicididae	Cibicides lobatulus, Cibicides dutemplei, Cibicides pseudoungerianus Cibicides lobatulus incl. 1 specimen C. lob. var. ornatus Cibicides dutemplei dOrbigny 1846 Cibicides pseudoungerianus Cushman 1922
Rotaliacea - Elphidiidae	Elphidium inflatum Elphidium inflatum
Nonionacea - Nonionidae	Nonion boueanum Nonion boueanum
- Spiroplectamminidae	Spiroplectammina carinata Spiroplectammina carinata d'Orbigny 1846
Textulariacea - Textulariidae	Textularia gramen, Bigenerina nodosaria Textularia gramen dOrbigny 1846 Bigenerina nodosaria
- Nodosariidae	Lenticulina (Lenticulina) calcar, Lenticulina (Robulus) grandis, Lenticulina (Robulus) cultrata, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) depauperata, Lenticulina (Planularia) grundensis, Lenticulina (Planularia) kubinyi, Lenticulina (Robulus) clypeiformis Lenticulina (Lenticulina) calcar Lenticulina (Robulus) grandis Lenticulina (Robulus) cultrata Lenticulina (Lenticulina) inornata Lenticulina (Robulus) depauperata L. (L.) depauperata for. costata Lenticulina (Planularia) grundensis Lenticulina (Planularia) kubinyi Lenticulina (Robulus) clypeiformis d'Orbigny 1846
Nodosariacea - Polymorphinidae	Globulina gibba, Guttulina problema, Pseudopolymorphina subnodosa, Pyrulina fusiformis Globulina gibba Guttulina problema d'Orbigny 1826 Pseudopolymorphina subnodosa P. subn. for. rugosa Pyrulina fusiformis
Nodosariacea - Lagenidae	Vaginulina obtusicosta, Nodosaria bacillum, Nodosaria raphanus, "Glandulina ovula" = Nodosaria (Glandulina), Dentalina intermittens, Dentalina bifurcata, Dentalina grossicosta, Dentalina pauperata, Dentalina konincki, Frondicularia annularis Vaginulina obtusicosta Nodosaria bacillum Nodosaria raphanus Brady 1884 "Glandulina ovula" = Nodosaria (Glandulina) Dentalina intermittens Dentalina bifurcata Dentalina grossicosta Dentalina pauperata d'Orbigny 1846 Dentalina konincki Reuss 1861 Frondicularia annularis
Anthozoa
Scleractinia - Micrabaciidae	Stephanophyllia imperialis Stephanophyllia imperialis Michelin 1841 stony coral
Scleractinia - Flabellidae	Flabellum sp., Flabellum pompeckji, Flabellum tuberculatum Flabellum sp. Lesson 1831 stony coral Flabellum pompeckji Krejci 1926 stony coral Flabellum tuberculatum Keferstein 1859 stony coral
Polychaeta
-	Polychaeta indet.¹ Polychaeta indet.¹ Grube 1850 Sedentarida indet.
Sabellida - Serpulidae	Josephella sp.¹, Mercierella sp.¹, Protula cf. canavarii¹ Josephella sp.¹ Caullery and Mesnil 1896 Mercierella sp.¹ Fauvel 1923 Protula cf. canavarii¹
Gymnolaemata
Cheilostomata - Lunulitidae	Lunulites voigti¹, Lunulites burdigalensis¹ Lunulites voigti¹ Lunulites burdigalensis¹
Cheilostomata - Membraniporidae	Reussirella haidingeri¹ Reussirella haidingeri¹
Cephalopoda
Spirulida - Spirulirostridae	Spirulirostra hoernesi¹ Spirulirostra hoernesi¹
Scaphopoda
Dentaliida - Dentaliidae	Lentigodentalium pseudomutabile¹, Antalis dumasi¹ Lentigodentalium pseudomutabile¹ tusk shell Antalis dumasi¹ tusk shell Antalis dumasi subsp.
Gadilida - Gadilidae	Gadila dingdenensis¹ Gadila dingdenensis¹ tusk shell
Bivalvia
Nuculida - Nuculidae	Nuculoma haesendoncki¹ Nuculoma haesendoncki¹ nut clam N. haesendoncki hanseata
Protobranchia - Nuculanidae	"Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ "Nuculana (Saccella) westendorpi" = Saccella westendorpi¹ Nyst and Westendorp 1839 pointed nut clam
Protobranchia - Yoldiidae	Yoldia (Yoldia) glaberrima¹, "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ Yoldia (Yoldia) glaberrima¹ von Münster 1837 clam "Portlandia (Yoldiella) pygmaea" = Yoldiella¹ clam
Arcida - Limopsidae	Limopsis (Limopsis) aurita¹, "Limopsis (Pectunculina) anomala" = Limopsis¹, Limopsis (Pectunculina) lamellata¹ Limopsis (Limopsis) aurita¹ Brocchi 1814 clam "Limopsis (Pectunculina) anomala" = Limopsis¹ clam Limopsis (Pectunculina) lamellata¹ Lehmann 1885 clam
Arcida - Arcidae	Scapharca (Scapharca) diluvii¹ Scapharca (Scapharca) diluvii¹ Lamarck 1805 clam
Ostreida - Pinnidae	Atrina pectinata¹ Atrina pectinata¹ Linnaeus 1758 oyster
Pectinida - Pectinoidae	Pseudamussium lilli¹, Aequipecten angelonii¹, Korobkovia woodi¹ Pseudamussium lilli¹ Pusch 1837 scallop Aequipecten angelonii¹ scallop Korobkovia woodi¹ scallop
Carditida - Carditidae	Cyclocardia sp.¹ Cyclocardia sp.¹ Conrad 1867 clam
Carditida - Astartidae	Astarte (Astarte) radiata¹ Astarte (Astarte) radiata¹ Nyst and Westendorp 1839 clam Astarte radiata fa. radiata 13 specimens, Astarte radiata fa. convexior 52 specimens
Cardiida - Cardiidae	"Habecardium subturgidum" = Laevicardium (Habecardium)¹, Parvicardium straeleni¹ "Habecardium subturgidum" = Laevicardium (Habecardium)¹ cockle H. subturgidum subturgidum Parvicardium straeleni¹ cockle
Cardiida - Tellinidae	Angulus (Moerella) aff. donacillus¹ Angulus (Moerella) aff. donacillus¹ tellin clam
Cardiida - Solecurtidae	Solecurtus basteroti¹ Solecurtus basteroti¹ clam
Cardiida - Semelidae	sensu lato Abra lehmanni¹, Abra (? Abra) sorgenfreii¹ sensu lato Abra lehmanni¹ abra Abra (? Abra) sorgenfreii¹ abra
Cardiida - Mactridae	Lutraria frangula¹ Lutraria frangula¹ clam
Cardiida - Veneridae	"Gouldia minima" = Gouldia (Gouldia) minima¹, Venus (Ventricoloidea) multilamella¹ "Gouldia minima" = Gouldia (Gouldia) minima¹ Montagu 1803 venus clam Venus (Ventricoloidea) multilamella¹ Lamarck 1818 venus clam V. (V.) multilamella multilamella
Cardiida - Glossidae	Glossus (Glossus) lunulatus¹ Glossus (Glossus) lunulatus¹ clam
Cardiida - Kelliellidae	Spaniodontella nitida¹ Spaniodontella nitida¹ clam
Cardiida - Erycinidae	"Hemilepton mionitidum" = Erycina (Hemilepton)¹ "Hemilepton mionitidum" = Erycina (Hemilepton)¹ clam
Pholadida - Teredinidae	Teredinidae indet.¹ Teredinidae indet.¹ Rafinesque 1815 clam
Pholadida - Corbulidae	Corbula (Varicorbula) gibba¹ Corbula (Varicorbula) gibba¹ Oliva 1792 clam C. (V.) gibba gibba
Solenida - Pharidae	Ensis hausmanni¹ Ensis hausmanni¹ clam
Hiatellida - Hiatellidae	Panopea (Panopea) meynardi¹ Panopea (Panopea) meynardi¹ clam
Poromyida - Verticordiidae	Verticordia (Verticordia) punctata¹ Verticordia (Verticordia) punctata¹ clam
Lucinida - Lucinidae	"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹, "Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti¹ Nyst 1861 clam C. droueti droueti "Lucinoma borealis" = Lucinoma (Lucinoma) borealis¹ Linnaeus 1767 clam
Gastropoda
Heterostropha - Ringiculidae	Ringicula (Ringiculina) buccinea³ Ringicula (Ringiculina) buccinea³ Sowerby 1823 snail
Heterostropha - Acteonidae	Acteon (Acteon) semistriatus³, Acteon (Acteon) "sp. 2"³ Acteon (Acteon) semistriatus³ snail spelled as 'Actaeon' Acteon (Acteon) "sp. 2"³ Montfort 1810 snail spelled as 'Actaeon'
Heterostropha - Pyramidellidae	Pyramidella (Voluspa) elata³, Pyramidella (Voluspa) plicosa³, Kleinella (Kleinella) sp.³, Chrysallida (Pyrgulina) pygmaea³, Chrysallida (Parthenina) "sp. 1"³, Eulimella (Eulimella) scillae³, Syrnola hoernesi³, Syrnola neumayri³, Turbonilla (Turbonilla) gastaldi³, Turbonilla (Turbonilla) koeneniana³, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³, Odostomia (Megastomia) cf. pallidaeformis³, Odostomia (Megastomia) tuexeni³, Odostomia (Megastomia) cf. conoidea³ Pyramidella (Voluspa) elata³ snail Pyramidella (Voluspa) plicosa³ snail Kleinella (Kleinella) sp.³ Adams 1860 snail Chrysallida (Pyrgulina) pygmaea³ Grateloup 1838 snail Ch. pygmaea f. belgica Chrysallida (Parthenina) "sp. 1"³ Bucquoy et al. 1883 snail Eulimella (Eulimella) scillae³ snail Syrnola hoernesi³ snail Syrnola neumayri³ snail Turbonilla (Turbonilla) gastaldi³ snail sensu JANSSEN Turbonilla (Turbonilla) koeneniana³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros³ snail sensu JANSSEN "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros³ snail sensu JANSSEN Odostomia (Megastomia) cf. pallidaeformis³ snail Odostomia (Megastomia) tuexeni³ snail Odostomia (Megastomia) cf. conoidea³ Brocchi 1814 snail
Cocculinoidea - Cocculinidae	Cocculina (Cocculina) miocaenica² Cocculina (Cocculina) miocaenica² snail
Opisthobranchia -	Roxania utriculus³ Roxania utriculus³ snail
Opisthobranchia - Retusidae	Retusa (Cylichnina) elongata³, Retusa (Cylichnina) subangistoma³, Volvulella (Volvulella) acuminata³ Retusa (Cylichnina) elongata³ snail Retusa (Cylichnina) subangistoma³ snail Volvulella (Volvulella) acuminata³ snail
Opisthobranchia - Cylichnidae	Cylichna (Cylichna) pseudoconvoluta³, Bullichna paucistriata³, Scaphander grateloupi³ Cylichna (Cylichna) pseudoconvoluta³ snail Bullichna paucistriata³ snail Scaphander grateloupi³ snail
Lepetellida - Lepetellidae	Lepetella compressiuscula² Lepetella compressiuscula² snail
Trochoidea - Trochidae	Calliostoma (Ampullotrochus) ? laureatum², Calliostoma (Ampullotrochus) serratocostatum² Calliostoma (Ampullotrochus) ? laureatum² top snail Calliostoma (Ampullotrochus) serratocostatum² top snail C. serratocostatum mülleri
Sorbeoconcha - Epitoniidae	sensu lato Turriscala straeleni², Acirsa (Hemiacirsa) aff. duvergieri², Epitonium (Spiniscala) frondiculum², sensu lato Cirsotrema ertborni², Cirsotrema (Cirsotrema) crassicostatum² sensu lato Turriscala straeleni² wentletrap T. straeleni for. germanica Acirsa (Hemiacirsa) aff. duvergieri² wentletrap Epitonium (Spiniscala) frondiculum² Wood 1842 wentletrap sensu lato Cirsotrema ertborni² wentletrap Cirsotrema (Cirsotrema) crassicostatum² Deshayes 1839 wentletrap
Sorbeoconcha - Rissoidae	sensu lato Alvania holsatica², Alvania (Alvania) partschi², Alvania (Alvania) basisulcata² sensu lato Alvania holsatica² snail Alvania (Alvania) partschi² snail Alvania (Alvania) basisulcata² snail
Sorbeoconcha - Eulimidae	Niso (Niso) terebellum², Balcis (Balcis) alba² Niso (Niso) terebellum² snail Balcis (Balcis) alba² snail incl. 5 specimens B. alba for. lactea
Sorbeoconcha - Naticidae	Naticidae indet.², "Natica neglecta" = Cochlis neglecta², "Natica sensu lato tigrina" = Cochlis tigrina², Euspira staringi², Euspira helicina², Natica koeneni², Tectonatica miopusilla² Naticidae indet.² Guilding 1834 moon snail Naticidae juv. et indet. "Natica neglecta" = Cochlis neglecta² Mayer 1858 moon snail "Natica sensu lato tigrina" = Cochlis tigrina² Röding 1798 moon snail Euspira staringi² moon snail Euspira helicina² Brocchi 1814 moon snail E. aggr. helicina Natica koeneni² moon snail Tectonatica miopusilla² moon snail
Sorbeoconcha - Vitrinellidae	Circulus praecedens², Circulus hennei² Circulus praecedens² snail 2 specimens C. praecedens for. praecedens and 1 specimen C. praecedens for. gliberti Circulus hennei² Glibert 1952 snail
Sorbeoconcha - Triviidae	Erato (Erato) germanica² Erato (Erato) germanica² Schilder 1929 snail
Neogastropoda - Terebridae	Strioterebrum hoernesi³, Strioterebrum ronaldi³, Terebra (Myurellina) acuminata³ Strioterebrum hoernesi³ auger snail Strioterebrum ronaldi³ auger snail Terebra (Myurellina) acuminata³ auger snail T. acuminata neglecta
Neogastropoda - Mangeliidae	Sorgenfreispira plicatelloides³, Sorgenfreispira "sp. 1"³, Sorgenfreispira tenella³, sensu lato Mangelia andersoni³ Sorgenfreispira plicatelloides³ snail Sorgenfreispira "sp. 1"³ Moroni 1979 snail Sorgenfreispira tenella³ Conrad 1848 snail sensu lato Mangelia andersoni³ Wienrich and Janssen 2007 snail
Neogastropoda - Turridae	Boreodrillia hosiusi³, Nitidiclavus maitrejus³, sensu lato Unedogemmula boreoturricula³, sensu lato Unedogemmula hanseata³, sensu lato Unedogemmula stoffelsii³, Fusiturris aquensis³, Fusiturris duchastelii³, Gemmula (Gemmula) sp.³, Gemmula (Gemmula) vermicularis³, Gemmula (Gemmula) zimmermanni³, sensu lato Inquisitor borealis³ Boreodrillia hosiusi³ turrid Nitidiclavus maitrejus³ turrid sensu lato Unedogemmula boreoturricula³ turrid sensu lato Unedogemmula hanseata³ turrid sensu lato Unedogemmula stoffelsii³ turrid Fusiturris aquensis³ turrid Fusiturris duchastelii³ turrid F. duchastelii flexiplicata Gemmula (Gemmula) sp.³ Weinkauff 1875 turrid Gemmula (Gemmula) vermicularis³ turrid Gemmula (Gemmula) zimmermanni³ turrid sensu lato Inquisitor borealis³ turrid
Neogastropoda - Borsoniidae	Bathytoma jugleri³, Asthenotoma pannoides³ Bathytoma jugleri³ snail Asthenotoma pannoides³ snail
Neogastropoda - Conidae	"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ "Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)³ cone shell
Neogastropoda - Clathurellidae	Pleurotomoides luisae³ Pleurotomoides luisae³ snail
Neogastropoda - Drilliidae	Splendrillia selenkae³ Splendrillia selenkae³ snail
Neogastropoda - Raphitomidae	Metuonella grippi³, Mioawateria andersoni³, Teretia anceps³ Metuonella grippi³ snail Mioawateria andersoni³ snail Teretia anceps³ snail
Neogastropoda - Ancillariidae	"Ancilla (Baryspira) obsoleta" = Amalda obsoleta² "Ancilla (Baryspira) obsoleta" = Amalda obsoleta² Brocchi 1814 snail
Neogastropoda - Olividae	Oliva (Strephona) dufresnei² Oliva (Strephona) dufresnei² Basterot 1825 olive snail
Neogastropoda - Mitridae	Mitra (Tiara) substriatula³ Mitra (Tiara) substriatula³ d'Orbigny 1852 mitre shell
Neogastropoda - Cancellariidae	Babylonella fusiformis², "Trigonostoma (Ventrilia) pouwi" = Ventrilia², Cancellaria (Merica) contorta² Babylonella fusiformis² snail "Trigonostoma (Ventrilia) pouwi" = Ventrilia² snail Cancellaria (Merica) contorta² snail C. contorta gelriana
Neogastropoda - Muricidae	Lyrotyphis (Eotyphis) fistulosus², Typhis (Typhis) ? pungens², "Murex (Haustellum) inornatus" = Haustellum² Lyrotyphis (Eotyphis) fistulosus² murex snail Typhis (Typhis) ? pungens² Solander in Brander and Brander 1766 murex snail "Murex (Haustellum) inornatus" = Haustellum² murex snail
Neogastropoda - Volutidae	Scaphella (Scaphella) bolli² Scaphella (Scaphella) bolli² volute
Neogastropoda - Nassariidae	Nassarius bocholtensis², Nassarius tenuistriatus² Nassarius bocholtensis² snail Nassarius tenuistriatus² snail
Neogastropoda - Fasciolariidae	Streptodictyon sexcostatus², ? Exilifusus attenuatus² Streptodictyon sexcostatus² snail ? Exilifusus attenuatus² snail
Neogastropoda - Columbellidae	Mitrella (Macrurella) nassoides² Mitrella (Macrurella) nassoides² snail
Neogastropoda - Costellariidae	Orthosurcula steinvorthi³ Orthosurcula steinvorthi³ ribbed mitre
Sorbeoconcha - Aporrhaidae	Aporrhais (Aporrhais) alata² Aporrhais (Aporrhais) alata² snail
Sorbeoconcha - Xenophoridae	Xenophora deshayesi² Xenophora deshayesi² Michelotti 1847 carrier shell
Sorbeoconcha - Cassidae	"Semicassis (Echinophoria) bicoronata" = Echinophoria² "Semicassis (Echinophoria) bicoronata" = Echinophoria² snail
Sorbeoconcha - Ranellidae	"Charonia (Sassia) tarbelliana" = Sassia² "Charonia (Sassia) tarbelliana" = Sassia² triton shell
Sorbeoconcha - Cerithiopsidae	Cerithiopsis (Dizoniopsis) "sp. 3"² Cerithiopsis (Dizoniopsis) "sp. 3"² Hanley 1849 snail
Sorbeoconcha - Triforidae	Cerithiella (Cerithiella) genei² Cerithiella (Cerithiella) genei² snail
Sorbeoconcha - Triphoridae	Norephora (Norephora) fritschi² Norephora (Norephora) fritschi² snail
Sorbeoconcha - Turritellidae	Turritella subangulata² Turritella subangulata² turret shell
Sorbeoconcha - Cerithiidae	Bittium (Bittium) tenuispina², "Bittium (Bittium) spina" = Bittium spina² Bittium (Bittium) tenuispina² cerith snail "Bittium (Bittium) spina" = Bittium spina² Partsch 1856 cerith snail
Neotaenioglossa - Calyptraeidae	Calyptraea (Calyptraea) chinensis² Calyptraea (Calyptraea) chinensis² Linnaeus 1758 slipper shell
Entomotaeniata - Cimidae	Actaeopyramis elatus³ Actaeopyramis elatus³ snail

¹Wienrich 1999; ²Wienrich 2001; ³Wienrich and Janssen 2007