Bore Lullingen 1, depth 39m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61896: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Foraminifera
 Planorbulinacea - Cibicididae
Cibicides lobatulus, Cibicides pseudoungerianus
 Rotaliacea - Elphidiidae
Elphidium inflatum
  - Spiroplectamminidae
Spiroplectammina carinata
 Textulariacea - Valvulinidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
 Textulariacea - Textulariidae
Textularia gramen
Textularia gramen dOrbigny 1846
  - Nodosariidae
Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) gibba, Lenticulina (Robulus) cultrata, Lenticulina (Vaginulinopsis) hauerina
 Nodosariacea - Polymorphinidae
Sigmomorphina sp., Globulina gibba, Guttulina austriaca, Guttulina problema, Pseudopolymorphina subnodosa
Sigmomorphina sp. Cushman and Ozawa 1928
Guttulina problema d'Orbigny 1826
Pseudopolymorphina subnodosa
incl. 1 specimen P. subn. for. rugosa
 Nodosariacea - Lagenidae
Dentalina "sp. 1", Dentalina konincki
Dentalina "sp. 1" d'Orbigny 1826
Dentalina konincki Reuss 1861
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
Polychaeta
  -
Polychaeta indet.1
Polychaeta indet.1 Grube 1850
Errantida indet. A
 Sabellida - Serpulidae
Serpulidae indet.1, Ditrupa cornea1
Serpulidae indet.1 Rafinesque 1835
Serpulidae indet. B
Gymnolaemata
 Cheilostomata - Cellariidae
Cellaria marginata1
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
 Cheilostomata - Membraniporidae
Biflustra savartii1, Discoporella umbellata1, Reussirella haidingeri1
Biflustra savartii1 Audouin 1826
Discoporella umbellata1 Defrance 1823
Rhynchonellata
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
Scaphopoda
 Dentaliida - Dentaliidae
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1, Fissidentalium twistringense1, Lentigodentalium pseudomutabile1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1
Bivalvia
 Nuculida - Nuculidae
Nuculoma haesendoncki1, Nucula (Lamellinucula) jeffreysi1, Nucula (Nucula) nucleus1
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
Nucula (Lamellinucula) jeffreysi1 Bellardi 1875 nut clam
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
 Protobranchia - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Protobranchia - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
 Arcida - Limopsidae
Limopsis (Limopsis) aurita1, "Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1, Bathyarca pectunculoides1
Bathyarca pectunculoides1 Scacchi 1834 clam
 Ostreida - Ostreidae
Ostreidae indet.1
Ostreidae indet.1 Rafinesque 1815 oyster
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Pectinida - Pectinoidae
Pseudamussium lilli1, Aequipecten angelonii1, Korobkovia woodi1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Limidae
Limea (Limea) aff. strigilata1
Limea (Limea) aff. strigilata1 Brocchi 1814 file clam
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Mytilida - Crenellidae
Musculus (Musculus) cf. sternbergensis1
 Mytilida - Mytilidae
Amygdalum aff. longum1
 Carditida - Condylocardiidae
Erycinella chavani1
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1, Goodallia (Goodallia) angulata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 27 specimens, Astarte radiata fa. convexior 5 specimens
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium straeleni1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
 Cardiida - Tellinidae
"Macoma elliptica" = Laciolina magna1, Angulus (Laceolina) benedeni1
"Macoma elliptica" = Laciolina magna1 Spengler 1798 tellin clam
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Psammobiidae
Gari (Psammotaena) aff. angusta1
 Cardiida - Solecurtidae
Solecurtus basteroti1
 Cardiida - Semelidae
sensu lato Abra lehmanni1, Abra (Abra) antwerpiensis1, Abra (? Abra) sorgenfreii1
Abra (Abra) antwerpiensis1 Glibert 1945 abra
 Cardiida - Mactridae
Lutraria frangula1
 Cardiida - Veneridae
"Gouldia minima" = Gouldia (Gouldia) minima1, Callista chione1, Venus (Ventricoloidea) multilamella1, Pelecyora (Cordiopsis) polytropa1
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
Callista chione1 Linnaeus 1758 venus clam
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Pelecyora (Cordiopsis) polytropa1 venus clam
P. (C.) polytropa nysti
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Leptonidae
Properycina (Properycina) edlaueri1, Lepton (? Lepton) transversarium1
 Cardiida - Lasaeidae
"Mysella (Mysella) bidentata" = Diarmaidia bidentata1
"Mysella (Mysella) bidentata" = Diarmaidia bidentata1 Montague 1803 clam
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Myidae
Sphenia pusilla1
Sphenia pusilla1 Sorgenfrel 1958 softshell clam
 Solenida - Pharidae
Cultellus (Cultellus) aff. cultellatus1, Ensis hausmanni1
 Poromyida - Cuspidariidae
Cuspidaria sp.1
Cuspidaria sp.1 Dall 1886 clam
 Thraciida - Periplomatidae
Periplomatidae indet.1
Periplomatidae indet.1 Dall 1895 clam
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
 Lucinida - Lucinidae
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1, "Lucinoma borealis" = Lucinoma (Lucinoma) borealis1
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
Gastropoda
 Heterostropha - Architectonicidae
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3, Architectonica (Architectonica) briarti3
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3 snail
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Heterostropha - Acteonidae
Acteon (Acteon) laevigatus3, Acteon (Acteon) "sp. 1"3, Acteon (Acteon) "sp. 2"3
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 1"3 Montfort 1810 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 2"3 Montfort 1810 snail
 Heterostropha - Pyramidellidae
Chrysallida (Pyrgulina) "sp. 2"3, Chrysallida (Pyrgulina) pygmaea3, Chrysallida (Parthenina) cimbrica3, Chrysallida (Parthenina) modesta3, Chrysallida (Parthenina) "sp. 1"3, Chrysallida (Parthenina) "sp. 3"3, Eulimella (Eulimella) acicula3, Eulimella (Eulimella) scillae3, Syrnola hoernesi3, Syrnola "sp. 2"3, Turbonilla (Mormula) "sp. 4"3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Odostomia) sp.3, Odostomia (Megastomia) tuexeni3, Odostomia (Megastomia) cf. conoidea3, Evalea sp.3
Chrysallida (Pyrgulina) "sp. 2"3 Carpenter 1856 snail
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
incl. 2 specimens Chrysallida pygmaea f. belgica
Chrysallida (Parthenina) "sp. 1"3 Bucquoy et al. 1883 snail
Chrysallida (Parthenina) "sp. 3"3 Bucquoy et al. 1883 snail
Syrnola "sp. 2"3 Adams 1860 snail
Turbonilla (Mormula) "sp. 4"3 Adams 1864 snail
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
Odostomia (Odostomia) sp.3 Fleming 1817 snail
Evalea sp.3 Adams 1860 snail
 Cocculinoidea - Cocculinidae
Cocculina miocaenica2
 Thecosomata - Limacinidae
Limacina miorostralis3
 Opisthobranchia -
Roxania utriculus3
 Opisthobranchia - Retusidae
Retusa (Cylichnina) elongata3, Volvulella (Volvulella) acuminata3
 Opisthobranchia - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3, Scaphander grateloupi3
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
 Trochoidea - Skeneidae
Skenea minuta2
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2
 Sorbeoconcha - Epitoniidae
Opalia (Pliciscala) pertusa2, Opalia (Nodiscala) rugatina2, Acirsa (Plesioacirsa) mioplicatula2, "Amaea (Scalina) subreticula" = Scalina2, Epitonium (Spiniscala) frondiculum2, sensu lato Cirsotrema ertborni2, Cirsotrema (Cirsotrema) crassicostatum2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Epitonium (Spiniscala) frondiculum2 Wood 1842 wentletrap
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Sorbeoconcha - Rissoidae
Ovirissoa gottscheana2, Ovirissoa westfalica2, sensu lato Alvania holsatica2
 Sorbeoconcha - Eulimidae
Balcis (Balcis) alba2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
Balcis (Balcis) alba2 snail
incl. 2 specimens B. alba for. lactea
 Sorbeoconcha - Naticidae
Naticidae indet.2, Sinum (Sinum) aquense2, "Natica neglecta" = Cochlis neglecta2, "Natica sensu lato tigrina" = Cochlis tigrina2, Euspira staringi2, Euspira helicina2, Tectonatica miopusilla2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus partimstriatus2, Circulus hennei2
Circulus praecedens2 snail
C. praecedens for. praecedens
Circulus hennei2 Glibert 1952 snail
 Sorbeoconcha - Tornidae
Tornus (Tornus) trigonostoma2
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Neogastropoda - Terebridae
Strioterebrum hoernesi3, Strioterebrum ronaldi3, Terebra (Myurellina) acuminata3
Strioterebrum ronaldi3 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Mangeliidae
Sorgenfreispira sorgenfreii3, Sorgenfreispira tenella3, sensu lato Mangelia andersoni3, Benthomangelia aff. obtusangula3
Sorgenfreispira tenella3 Conrad 1848 snail
sensu lato Mangelia andersoni3 Wienrich and Janssen 2007 snail
 Neogastropoda - Turridae
Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Fusiturris aquensis3, Fusiturris duchastelii3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) zimmermanni3, sensu lato Inquisitor borealis3, Perrona obliquiplicatula3
sensu lato Unedogemmula boreoturricula3 turrid
incl. 8 specimens U. (s. lat.) boreoturricula x hanseata ?
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
 Neogastropoda - Borsoniidae
"Genota ramosa" = Genota valeriae3, Bathytoma jugleri3, Asthenotoma pannoides3, Asthenotoma pannus3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Raphitomidae
Favriella sinuosula3, Metuonella grippi3, Raphitoma aff. praehispida3, Teretia anceps3, sensu lato Teretia "sp. 2"3
sensu lato Teretia "sp. 2"3 Norman 1888 snail
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Cancellariidae
Trigonostoma (Trigonostoma) extractrix2, Babylonella fusiformis2, Cancellaria (Merica) bellardi2, Cancellaria (Merica) contorta2, Sveltia varicosa2
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Muricidae
Coralliophila (Aldrichia) cf. granifera2, Lyrotyphis (Eotyphis) fistulosus2, Typhis (Typhis) ? pungens2, "Murex (Haustellum) inornatus" = Haustellum2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius voorthuyseni2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, ? Exilifusus attenuatus2
 Neogastropoda - Columbellidae
Anachis (Costoanachis) "sp. 1"2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Costellariidae
"Vexillum (Uromitra) gliberti" = Uromitra3, Orthosurcula steinvorthi3
"Vexillum (Uromitra) gliberti" = Uromitra3 ribbed mitre
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2, "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Cerithiopsidae
Cerithiopsis (Cerithiopsis) vogeli2
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Turritellidae
Turritella subangulata2
 Sorbeoconcha - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007