Bore Lullingen 1, depth 57m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E)

• Paleocoordinates: 49.0° N, 8.5° E (Wright 2013)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Tichelh. stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61900: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names
Sedentaria
 Sabellida - Serpulidae
Ditrupa cornea1
Gymnolaemata
 Cheilostomata - Membraniporidae
Biflustra savartii1
Biflustra savartii1 Audouin 1826
 Cheilostomata - Celleporidae
Celleporidae indet.1, Palmiscenea aff. alata1, Turbicellepora compressa1
 Cheilostomata - Cupuladriidae
Reussirella haidingeri1
 Cheilostomata - Lunulitidae
Lunulites voigti1
Gastropoda
 Heterostropha - Pyramidellidae
Syrnola hoernesi3, Syrnola neumayri3, Syrnola "sp. 3"3, Turbonilla (Mormula) cf. miomutiniensis3, Turbonilla (Turbonilla) koeneniana3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) tuexeni3, Chrysallida (Pyrgulina) pygmaea3, Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3
Syrnola "sp. 3"3 Adams 1860 snail
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
Ch. pygmaea f. belgica
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
 Heterostropha - Architectonicidae
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3, Architectonica (Pseudotorinia) sensu lato obtusa3, Architectonica (Architectonica) briarti3
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3 snail
 Heterostropha - Acteonidae
Acteon (Acteon) semistriatus3, Acteon (Acteon) "sp. 2"3, Crenilabium terebelloides3
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 2"3 Montfort 1810 snail
spelled as 'Actaeon'
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3, Ringicula (Ringiculina) ventricosa3
 Epitonioidea - Epitoniidae
Epitonium (? Hyaloscala) "sp. 3"2, "Amaea (Scalina) subreticula" = Scalina2, Opalia (Nodiscala) rugatina2, ? Acirsa sp.2, Acirsa (Hemiacirsa) lanceolata2, "Epitonium (Clathrus) clathrus" = Clathrus2
Epitonium (? Hyaloscala) "sp. 3"2 Bolten 1798 wentletrap
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
? Acirsa sp.2 Mörch 1857 wentletrap
? Acirsa (s. lat.) sp.
"Epitonium (Clathrus) clathrus" = Clathrus2 wentletrap
 Vanikoroidea - Eulimidae
Balcis (Balcis) alba2, Niso (? Niso) sp.2, Niso (Niso) terebellum2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
Balcis (Balcis) alba2 snail
incl. 17 specimens B. alba for. lactea
Niso (? Niso) sp.2 Risso 1826 snail
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3
 Neogastropoda - Nassariidae
Amyclina facki2, Nassarius bocholtensis2, Nassarius cavatus2, Nassarius karinae2
Nassarius karinae2 Wienrich 2001 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) hosiusi2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Fasciolariidae
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2, Streptodictyon sexcostatus2
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2 snail
 Neogastropoda - Cancellariidae
Babylonella fusiformis2, Sveltia varicosa2, "Trigonostoma (Ventrilia) geslini" = Ventrilia2, "Trigonostoma (Ventrilia) pouwi" = Ventrilia2, Cancellaria (Merica) contorta2
Sveltia varicosa2 snail
S. varicosa paucicostata
"Trigonostoma (Ventrilia) geslini" = Ventrilia2 snail
T. geslini josephinae
"Trigonostoma (Ventrilia) pouwi" = Ventrilia2 snail
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3
 Neogastropoda - Raphitomidae
Teretia anceps3, Favriella sinuosula3, Metuonella grippi3
 Neogastropoda - Terebridae
Strioterebrum fabritzi3, Strioterebrum basteroti3, Terebra (Myurellina) acuminata3
Strioterebrum basteroti3 Nyst 1843 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Mangeliidae
Sorgenfreispira sorgenfreii3, Benthomangelia aff. obtusangula3, Benthomangelia grippi3, sensu lato Mangelia miorugulosa3
Benthomangelia grippi3 Anderson 1964 snail
sensu lato Mangelia miorugulosa3 Kautsky 1925 snail
 Neogastropoda - Borsoniidae
Asthenotoma pannus3, Bathytoma jugleri3
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Turridae
Nitidiclavus maitrejus3, Boreodrillia hosiusi3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) vermicularis3, Gemmula (Gemmula) zimmermanni3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Fusiturris aquensis3, Fusiturris duchastelii3
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
sensu lato Unedogemmula boreoturricula3 turrid
incl. 19 specimens U. (s. lat.) boreoturricula x hanseata ?
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Muricidae
Typhis (Typhis) ? pungens2, Lyrotyphis (Eotyphis) fistulosus2, "Murex (Haustellum) inornatus" = Haustellum2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
 Neogastropoda - Volutidae
Scaphella (Scaphella) bolli2
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Vitrinellidae
Circulus hennei2
Circulus hennei2 Glibert 1952 snail
 Sorbeoconcha - Iravadiidae
Ceratia regiorivi2
 Sorbeoconcha - Naticidae
Naticidae indet.2, Tectonatica miopusilla2, "Natica sensu lato tigrina" = Cochlis tigrina2, Neverita josephinia2, Euspira nysti2, Euspira staringi2, Euspira helicina2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
Euspira nysti2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
 Sorbeoconcha - Rissoidae
Onoba semicostata2, sensu lato Alvania holsatica2, Ovirissoa gottscheana2
Onoba semicostata2 Montagu 1803 snail
 Cerithioidea - Turritellidae
Turritella subangulata2
 Cerithioidea - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
 Trochoidea - Skeneidae
Tubiola sp.2
Tubiola sp.2 Adams 1863 snail
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
 Opisthobranchiata - Retusidae
Volvulella (Volvulella) acuminata3, Retusa (Cylichnina) elongata3
 Opisthobranchiata - Cylichnidae
Bullichna paucistriata3, Cylichna (Cylichna) pseudoconvoluta3, Cylichna (Cylichna) subcylindrica3
 Opisthobranchiata -
Roxania utriculus3
Bivalvia
 Carditida - Astartidae
Astarte (Astarte) radiata1, Goodallia (Goodallia) angulata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Poromyida - Cuspidariidae
Cuspidaria sp.1, Cardiomya (Cardiomya) costellata1
Cuspidaria sp.1 Dall 1886 clam
 Hiatellida - Hiatellidae
Panopea (Panopea) meynardi1, "Hiatella (Hiatella) arctica" = Hiatella arctica1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Cardiida - Lasaeidae
"Mysella (Mysella) bidentata" = Diarmaidia bidentata1
"Mysella (Mysella) bidentata" = Diarmaidia bidentata1 Montague 1803 kelly clam
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Glossidae
Meiocardia harpa1
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Semelidae
Abra (Abra) antwerpiensis1
Abra (Abra) antwerpiensis1 Glibert 1945 abra
 Cardiida - Cardiidae
Parvicardium ovale1, Parvicardium straeleni1, "Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
 Pholadida - Teredinidae
Teredinidae indet.1
Teredinidae indet.1 Rafinesque 1815 clam
 Pholadida - Corbulidae
"Lentidium (Lentidium) donaciformis" = Lentidium (Ajanssenium) donaciforme1, Corbula (Varicorbula) gibba1
"Lentidium (Lentidium) donaciformis" = Lentidium (Ajanssenium) donaciforme1 Nyst 1836 clam
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Lucinida - Lucinidae
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1, "Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
 Mytilida - Mytilidae
Modiolula phaseolina1
Modiolula phaseolina1 Philippi 1844 mussel
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1
 Arcida - Glycymerididae
Glycymeris (Glycymeris) obovata1
Glycymeris (Glycymeris) obovata1 Lamarck 1819 clam
G. obovata baldii
 Arcida - Limopsidae
"Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1, Limopsis (Limopsis) aurita1
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
 Pectinida - Limidae
Limea (Limea) aff. strigilata1
Limea (Limea) aff. strigilata1 Brocchi 1814 file clam
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
 Pectinida - Pectinoidae
Korobkovia woodi1, Pseudamussium lilli1, Aequipecten seniensis1
Pseudamussium lilli1 Pusch 1837 scallop
 Nuculanida - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculida - Nuculidae
Nucula (Nucula) nucleus1, Nucula (Lamellinucula) jeffreysi1, Nuculoma haesendoncki1, Nuculoma laevigata1
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
Nucula (Lamellinucula) jeffreysi1 Bellardi 1875 nut clam
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
Scaphopoda
 Dentaliida - Dentaliidae
Lentigodentalium pseudomutabile1, "Pseudantalis aturensis" = Dentalium (Pseudantilis)1, Antalis dumasi1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
Antalis dumasi1 tusk shell
Antalis dumasi subsp.
 Dentaliida - Gadilinidae
Gadilina sp.1
Gadilina sp.1 Foresti 1895 tusk shell
Anthozoa
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Turbinoliidae
"Cylindrophyllia duncani" = Peponocyathus duncani, "Cylindrophyllia cf. ottnangensis" = Peponocyathus
"Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
"Cylindrophyllia cf. ottnangensis" = Peponocyathus stony coral
 Scleractinia - Flabellidae
Flabellum sp.
Flabellum sp. Lesson 1831 stony coral
Globothalamea
 Rotaliida - Nonionidae
Nonion boueanum
 Rotaliida - Cibicididae
Cibicides dutemplei, Cibicides lobatulus
Cibicides dutemplei dOrbigny 1846
Cibicides lobatulus Walker and Jacob 1798
Foraminifera
 Rotalioidea - Elphidiidae
Elphidium inflatum
 Spiroplectamminacea - Spiroplectamminidae
Spiroplectammina carinata
Tubothalamea
 Miliolida - Hauerinidae
Quinqueloculina haidingeri, Triloculina inflata, Triloculina tricarinata
 Miliolida - Miliolidae
Miliolidae indet., Pyrgo clypeata, Pyrgo sinedente
Miliolidae indet. Ehrenberg 1839
Pyrgo sinedente Wienrich 1997
 Miliolida - Cornuspiridae
Cornuspira involvens
Nodosariata
 Nodosariida - Nodosariidae
Dentalina "sp. 2", Dentalina guttifera, Dentalina semiplicata, Dentalina konincki, Frondicularia cf. nysti
Dentalina "sp. 2" d'Orbigny 1826
Dentalina konincki Reuss 1861
 Vaginulinida - Vaginulinidae
Lenticulina (Astacolus) compressa, Lenticulina (Lenticulina) inornata, Lenticulina (Lenticulina) gibba, Lenticulina (Lenticulina) calcar, Lenticulina (Lenticulina) rotulata
 Polymorphinida - Polymorphinidae
Pseudopolymorphina subnodosa, Guttulina austriaca, Guttulina problema, Globulina gibba, Sigmomorphina schwageri
Guttulina problema d'Orbigny 1826
Globulina gibba d'Orbigny 1832
Foraminifera
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
 Lituolida - Lituolidae
Textularia laevigata, Textularia gramen
Textularia gramen dOrbigny 1846
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007