Bore Lullingen 1, depth 45m (Miocene of Germany)

Where: North Rhein-Westfalia, Germany (51.5° N, 6.2° E: paleocoordinates 51.8° N, 4.8° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Middle Miocene (16.0 - 11.6 Ma)

• Reinbek-Bislicher stage

• bed-level stratigraphic resolution

Environment/lithology: transition zone or lower shoreface; unlithified, fine-grained, silty sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1992

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 61905: authorized by Martin Aberhan, entered by Sabine Nürnberg on 07.07.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Globothalamea
 Rotaliida - Cibicididae
Cibicides lobatulus, Cibicides dutemplei
Cibicides dutemplei dOrbigny 1846
Nodosariata
 Nodosariida - Nodosariidae
Vaginulina obtusicosta, Dentalina grossicosta, Dentalina "sp. 1", Dentalina konincki, Nodosaria bacillum
Dentalina "sp. 1" d'Orbigny 1826
Dentalina konincki Reuss 1861
 Polymorphinida - Polymorphinidae
Globulina gibba, Guttulina austriaca, Guttulina problema
Globulina gibba d'Orbigny 1832
Guttulina problema d'Orbigny 1826
 Vaginulinida - Vaginulinidae
Lenticulina (Lenticulina) inornata, Lenticulina (Planularia) grundensis, Lenticulina (Robulus) grandis, Lenticulina (Robulus) cultrata, Lenticulina (Lenticulina) rotulata, Lenticulina (Lenticulina) calcar
Foraminifera
 Lituolacea - Ataxophragmiidae
Martinottiella communis
Martinottiella communis Huang et al. 2019
Tubothalamea
 Miliolida - Miliolidae
Spiroloculina badensis
Foraminifera
 Lituolida - Lituolidae
Textularia gramen
Textularia gramen dOrbigny 1846
 Rotalioidea - Elphidiidae
Elphidium inflatum
  - Spiroplectamminidae
Spiroplectammina carinata
Sedentaria
 Sabellida - Serpulidae
Serpulidae indet.1, Spirorbis sp.1
Serpulidae indet.1 Rafinesque 1835
Serpulidae indet. A
Spirorbis sp.1 Daudin 1800
Rhynchonellata
 Rhynchonellida - Cryptoporidae
Cryptopora sp.1
Cryptopora sp.1 Jeffreys 1869
 Terebratulida - Terebratulidae
Terebratula sp.1
Terebratula sp.1 Müller 1776
Gymnolaemata
 Cheilostomata - Membraniporidae
Discoporella umbellata1, Cupuladria canariensis1, Reussirella haidingeri1
Discoporella umbellata1 Defrance 1823
Cupuladria canariensis1
C. canariensis cavernosa
 Cheilostomata - Lunulitidae
Lunulites voigti1
Bivalvia
 Nuculanida - Yoldiidae
"Portlandia (Yoldiella) pygmaea" = Yoldiella1, Yoldia (Yoldia) glaberrima1
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
 Nuculanida - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Nuculida - Nuculidae
Nucula (Nucula) nucleus1, Nucula (Lamellinucula) jeffreysi1, Nuculoma haesendoncki1
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
Nucula (Lamellinucula) jeffreysi1 Bellardi 1875 nut clam
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
 Pectinida - Pectinoidae
Pseudamussium lilli1, Korobkovia woodi1, Aequipecten angelonii1, Aequipecten seniensis1
Pseudamussium lilli1 Pusch 1837 scallop
 Pectinida - Anomiidae
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1, Pododesmus (Monia) anitae1
"Pododesmus (Heteranomia) squamula" = Heteranomia squamula1 Linnaeus 1758 scallop
Pododesmus (Monia) anitae1 Janse and Janssen 1983 scallop
 Ostreida - Pinnidae
Atrina pectinata1
Atrina pectinata1 Linnaeus 1758 oyster
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1, Bathyarca pectunculoides1
Bathyarca pectunculoides1 Scacchi 1834 clam
 Arcida - Limopsidae
Limopsis (Limopsis) aurita1, "Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
 Mytilida - Mytilidae
Modiolula phaseolina1, Arcoperna sericea1
Modiolula phaseolina1 Philippi 1844 mussel
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Goodallia (Goodallia) angulata1, Astarte (Astarte) radiata1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 86 specimens, Astarte radiata fa. convexior 10 specimens
 Lucinida - Lucinidae
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1, "Lucinoma borealis" = Lucinoma (Lucinoma) borealis1
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
 Hiatellida - Hiatellidae
Panopea (Panopea) meynardi1, "Hiatella (Hiatella) arctica" = Hiatella arctica1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Mactridae
Lutraria frangula1, Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Veneridae
Venus (Ventricoloidea) multilamella1, "Gouldia minima" = Gouldia (Gouldia) minima1
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium straeleni1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
 Cardiida - Tellinidae
"Macoma elliptica" = Laciolina magna1, Angulus (Moerella) aff. donacillus1
"Macoma elliptica" = Laciolina magna1 Spengler 1798 tellin clam
 Cardiida - Semelidae
sensu lato Abra lehmanni1, Abra (? Abra) sorgenfreii1
 Cardiida - Solecurtidae
Azorinus (Azorinus) chamasolen1
Azorinus (Azorinus) chamasolen1 clam
A. chamasolen miocaenicus
 Cardiida - Lasaeidae
Scacchia (Scacchia) degrangei1
Scacchia (Scacchia) degrangei1 Cossmann and Peyrot 1911 kelly clam
 Thraciida - Thraciidae
Thracia (Thracia) aff. inflata1
Scaphopoda
 Dentaliida - Dentaliidae
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1, Lentigodentalium pseudomutabile1, Fissidentalium twistringense1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1
Gastropoda
 Pseudomelanoidea - Pseudomelaniidae
sensu lato Inquisitor borealis3
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Epitonioidea - Epitoniidae
Epitonium (Spiniscala) frondiculum2, Acirsa (Hemiacirsa) aff. duvergieri2, Acirsa (Plesioacirsa) mioplicatula2, "Amaea (Scalina) subreticula" = Scalina2, sensu lato Cirsotrema ertborni2, Cirsotrema (Cirsotrema) crassicostatum2, Opalia (Pliciscala) pertusa2, Opalia (Nodiscala) rugatina2
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Cirsotrema (Cirsotrema) crassicostatum2 Deshayes 1839 wentletrap
 Vanikoroidea - Eulimidae
Balcis (Balcis) alba2, Balcis (? Balcis) sp.2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
Balcis (Balcis) alba2 snail
incl. 1 specimen B. alba for. eichwaldi and 3 specimens B. alba for. lactea
Balcis (? Balcis) sp.2 Leach 1847 snail
B. (? Balcis) sp. nov.
 Cerithioidea - Cerithiidae
"Bittium (Bittium) spina" = Bittium spina2, Bittium (Bittium) tenuispina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Cerithioidea - Turritellidae
Turritella subangulata2, Turritella "sp. 2"2
Turritella "sp. 2"2 Lamarck 1799 turret shell
 Sorbeoconcha - Newtoniellidae
"Cerithiopsidella (Vatopsis) dautzenbergi" = Vatopsis2
"Cerithiopsidella (Vatopsis) dautzenbergi" = Vatopsis2 snail
 Sorbeoconcha - Triforidae
Cerithiella (Cerithiella) genei2
 Sorbeoconcha - Triphoridae
Norephora (Norephora) fritschi2
 Sorbeoconcha - Naticidae
Naticidae indet.2, Neverita josephinia2, Euspira staringi2, Euspira sp.2, Euspira helicina2, Polinices (Polinices) miocolligens2, Tectonatica miopusilla2, "Natica neglecta" = Cochlis neglecta2, "Natica sensu lato tigrina" = Cochlis tigrina2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
Euspira staringi2 moon snail
Euspira sp.2 Agassiz 1837 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
 Sorbeoconcha - Rissoidae
Ovirissoa gottscheana2, Ovirissoa westfalica2, sensu lato Alvania holsatica2, Alvania (Alvania) partschi2, Alvania (Alvania) dingdensis2
Alvania (Alvania) dingdensis2 Janssen 1967 snail
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Neogastropoda - Fasciolariidae
? Exilifusus attenuatus2, Fredenia ronaldi n. sp.2, Streptodictyon sexcostatus2, "Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2
Fredenia ronaldi n. sp.2 Wienrich 2001 snail
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2 snail
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius tenuistriatus2, Nassarius woodwardi2, Nassarius karinae2, Nassarius prysmaticus2, Amyclina facki2
Nassarius karinae2 Wienrich 2001 snail
Nassarius prysmaticus2 Brocchi 1814 snail
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Columbellidae
Mitrella (Macrurella) nassoides2, Anachis (Costoanachis) hosiusi2, Anachis (Costoanachis) terebralis2, Anachis (Costoanachis) "sp. 1"2, Columbellopsis cf. astensis2
Columbellopsis cf. astensis2 snail
C. ?. sp. n. cf. astensis
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Cancellariidae
Cancellaria (Merica) contorta2, Cancellaria (Bivetiella) cancellata2, "Trigonostoma (Ventrilia) pouwi" = Ventrilia2, Babylonella fusiformis2, Trigonostoma (Trigonostoma) banardi2, Trigonostoma (Trigonostoma) cf. spinifera2, Sveltia varicosa2
Cancellaria (Merica) contorta2 snail
C. contorta gelriana
Cancellaria (Bivetiella) cancellata2 Linnaeus 1767 snail
C. cancellata praecedens
"Trigonostoma (Ventrilia) pouwi" = Ventrilia2 snail
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Raphitomidae
Metuonella grippi3, Favriella "sp. 2"3, Teretia anceps3
Favriella "sp. 2"3 Hornung 1920 snail
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Mangeliidae
Sorgenfreispira "sp. 2"3, Sorgenfreispira tenella3, Benthomangelia aff. obtusangula3
Sorgenfreispira "sp. 2"3 Moroni 1979 snail
Sorgenfreispira tenella3 Conrad 1848 snail
 Neogastropoda - Turridae
sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula stoffelsii3, Boreodrillia hosiusi3, Nitidiclavus maitrejus3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) vermicularis3, Gemmula (Gemmula) zimmermanni3, Fusiturris aquensis3, Fusiturris duchastelii3, Acamptogenotia escheri3
sensu lato Unedogemmula boreoturricula3 turrid
incl. 20 specimens U. (s. lat.) boreoturricula x hanseata ?
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
 Neogastropoda - Terebridae
Strioterebrum hoernesi3, Strioterebrum ronaldi3, Terebra (Myurellina) acuminata3
Strioterebrum ronaldi3 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Clathurellidae
Pleurotomoides luisae3, Pleurotomoides simplex3, ? Ceritoturris hannoverana3
 Neogastropoda - Borsoniidae
Bathytoma jugleri3, Asthenotoma pannoides3, Asthenotoma pannus3, "Genota ramosa" = Genota valeriae3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
 Neogastropoda - Mitridae
Mitra (Tiara) substriatula3
Mitra (Tiara) substriatula3 d'Orbigny 1852 mitre shell
 Neogastropoda - Costellariidae
Orthosurcula steinvorthi3, "Vexillum (Uromitra) boreocinctum" = Uromitra3, "Vexillum (Uromitra) pliactulum" = Uromitra3
"Vexillum (Uromitra) boreocinctum" = Uromitra3 ribbed mitre
"Vexillum (Uromitra) pliactulum" = Uromitra3 ribbed mitre
 Neogastropoda - Muricidae
Coralliophila (Pseudomurex) bracteata2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, "Murex (Haustellum) inornatus" = Haustellum2, Lyrotyphis (Eotyphis) fistulosus2, Typhis (Typhis) ? pungens2
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Vitrinellidae
Circulus praecedens2, Circulus hennei2
Circulus praecedens2 snail
C. praecedens for. praecedens
Circulus hennei2 Glibert 1952 snail
 Trochoidea - Trochidae
Calliostoma (Ampullotrochus) ? laureatum2, Calliostoma (Ampullotrochus) serratocostatum2
Calliostoma (Ampullotrochus) serratocostatum2 top snail
C. serratocostatum mülleri
 Lepetellida - Lepetellidae
Lepetella compressiuscala2
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3
 Heterostropha - Acteonidae
Crenilabium terebelloides3, Acteon (Acteon) laevigatus3, Acteon (Acteon) semistriatus3
Acteon (Acteon) laevigatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
 Heterostropha - Architectonicidae
Architectonica (Architectonica) simplex3
 Heterostropha - Pyramidellidae
Syrnola dubia3, Syrnola hoernesi3, Syrnola neumayri3, Syrnola "sp. 1"3, Pyramidella (Voluspa) plicosa3, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, "Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3, Eulimella (Eulimella) acicula3, Eulimella (Eulimella) scillae3, Odostomia (Brachystomia) cf. sallespissiensis3, Odostomia (Megastomia) cf. conoidea3, Odostomia (Megastomia) tuexeni3, Odostomia (Megastomia) "sp. 1"3, Odostomia (Megastomia) "sp. 3"3, Turbonilla (Mormula) "sp. 4"3, Turbonilla (Turbonilla) gastaldi3, Turbonilla (Turbonilla) koeneniana3, Chrysallida (Parthenina) cimbrica3, Chrysallida (Parthenina) "sp. 1"3, Chrysallida (Parthenina) "sp. 3"3, Chrysallida (Pyrgulina) pygmaea3
Syrnola "sp. 1"3 Adams 1860 snail
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) undulata" = Pyrgolampros3 snail
sensu JANSSEN
Turbonilla (Mormula) "sp. 4"3 Adams 1864 snail
Turbonilla (Turbonilla) gastaldi3 snail
sensu JANSSEN
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
Chrysallida (Parthenina) "sp. 1"3 Bucquoy et al. 1883 snail
Chrysallida (Parthenina) "sp. 3"3 Bucquoy et al. 1883 snail
Chrysallida (Pyrgulina) pygmaea3 Grateloup 1838 snail
Ch. pygmaea f. belgica
 Heterostropha - Murchisonellidae
Ebala (Ebala) vandervlerki3
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
 Opisthobranchia -
Roxania utriculus3
 Opisthobranchia - Retusidae
Volvulella (Volvulella) acuminata3, Retusa (Cylichnina) elongata3
 Opisthobranchia - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3, Bullichna paucistriata3, Scaphander grateloupi3
Anthozoa
 Scleractinia - Caryophylliidae
Deltocyathus reussi
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
 Scleractinia - Turbinoliidae
"Cylindrophyllia duncani" = Peponocyathus duncani
"Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
 Scleractinia - Dendrophylliidae
"Eupsammia praelonga" = Balanophyllia praelonga
"Eupsammia praelonga" = Balanophyllia praelonga Michelotti 1838 stony coral
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007