Bore Kevelaer, depth 85m (Miocene to of Germany)

Where: North Rhein-Westfalia, Germany (51.6° N, 6.3° E: paleocoordinates 51.8° N, 4.6° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Early/Lower Miocene to Early/Lower Miocene (23.0 - 11.6 Ma)

• boundary Reinbek-Tichelh./Hemmoor stage

• bed-level stratigraphic resolution

Environment/lithology: marine; unlithified, fine-grained, brown sandstone

Size classes: macrofossils, mesofossils, microfossils

Preservation: original aragonite

Collection methods: core, mechanical, peroxide, sieve,

• well drilling, 1994

Primary reference: G. Wienrich. 1997. Die Fauna des marinen Miozäns von Kevelaer (Niederrhein). Band 1 Foraminiferen, Anthozoen. Backhuys Publishers, Leiden 1:1-187 [M. Aberhan/S. Nurnberg/S. Nurnberg]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 62884: authorized by Martin Aberhan, entered by Sabine Nürnberg on 01.08.2006

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Globothalamea
 Rotaliida - Rosalinidae
Rosalina globularis
Rosalina globularis d'Orbigny 1826
Tubothalamea
 Miliolida - Hauerinidae
Triloculina inflata, Triloculina scapha
 Miliolida - Miliolidae
Miliolidae indet., Sigmoilina tenuis, Pyrgo bulloides
Miliolidae indet. Ehrenberg 1839
Foraminifera
 Discorbacea - Bagginidae
Cancris auriculus
Cancris auriculus Fichtel and Moll 1798
 Discorbacea - Eponididae
Eponides nanus
 Planorbulinacea - Planulinidae
Planulina austriaca
 Planorbulinacea - Cibicididae
Cibicides lobatulus, Cibicides aknerianus, Cibicides dutemplei, Cibicides pseudoungerianus
Cibicides lobatulus
incl. 4 specimens C. lobatulus var. ornatus
Cibicides dutemplei dOrbigny 1846
 Rotaliacea - Elphidiidae
Elphidium inflatum, Elphidium ungeri
 Buliminacea - Uvigerinidae
Uvigerina semiornata, Uvigerina tenuipustulata
Uvigerina semiornata
U. semiornata for. semiornata
 Nonionacea - Nonionidae
Nonion boueanum, Nonion affine, Nonion (Porosononion) soldanii, Nonion sp.
Nonion sp. Monfort 1808
 Asterigerinacea - Asterigerinidae
Asterigerina guerichi
Asterigerina guerichi
A. gürichi staeschi
  - Buliminidae
Virgulina (Virgulinella) pertusa
  - Spiroplectamminidae
Spiroplectammina carinata
 Textulariacea - Valvulinidae
Karreriella siphonella, Martinottiella communis
Martinottiella communis Huang et al. 2019
 Textulariacea - Textulariidae
Textularia gramen, Bigenerina nodosaria
Textularia gramen dOrbigny 1846
  - Nodosariidae
Lenticulina (Robulus) grandis, Lenticulina (Lenticulina) inornata, Lenticulina (Robulus) ornata, Lenticulina (Vaginulinopsis) gerlachi, Lenticulina (Vaginulinopsis) hauerina, Lenticulina (Planularia) grundensis, Lenticulina (Planularia) kubinyi, Lenticulina (Robulus) vortex
Lenticulina (Robulus) vortex Fichtel and Moll 1798
 Nodosariacea - Polymorphinidae
Sigmomorphina sp., Globulina gibba, Guttulina austriaca, Guttulina problema, Pseudopolymorphina incerta, Pseudopolymorphina ovalis
Sigmomorphina sp. Cushman and Ozawa 1928
Globulina gibba
incl. 1 specimen G. gibba for. tubulosa
Guttulina problema d'Orbigny 1826
 Nodosariacea - Lagenidae
Vaginulina obtusicosta, Nodosaria bacillum, Nodosaria venusta, "Glandulina ovula" = Nodosaria (Glandulina), Lagena apiculata, Lagena striata, Dentalina intermittens, Dentalina inornata, Dentalina grossicosta, Dentalina acuticauda, Dentalina konincki
"Glandulina ovula" = Nodosaria (Glandulina)
Lagena striata Williamson 1848
Dentalina konincki Reuss 1861
Anthozoa
 Scleractinia - Dendrophylliidae
"Eupsammia calyculus" = Balanophyllia
"Eupsammia calyculus" = Balanophyllia stony coral
 Scleractinia - Micrabaciidae
Stephanophyllia imperialis
Stephanophyllia imperialis Michelin 1841 stony coral
 Scleractinia - Flabellidae
Flabellum sp., Flabellum cristatum
Flabellum sp. Lesson 1831 stony coral
Flabellum cristatum Milne-Edwards and Haime 1851 stony coral
F. cristatum for. dietrichi
 Scleractinia - Caryophylliidae
Ceratocyathus granulatus
Ceratocyathus granulatus Goldfuss 1826 stony coral
 Scleractinia - Turbinoliidae
"Cylindrophyllia duncani" = Peponocyathus duncani
"Cylindrophyllia duncani" = Peponocyathus duncani Reuss 1871 stony coral
Polychaeta
  -
Polychaeta indet.1
Polychaeta indet.1 Grube 1850
Sedentarida indet.
 Sabellida - Serpulidae
Filograna sp.1
Filograna sp.1 Oken 1815
Gymnolaemata
 Cheilostomata - Lunulitidae
Lunulites voigti1, Lunulites burdigalensis1
 Cheilostomata - Membraniporidae
Discoporella umbellata1, Reussirella haidingeri1
Discoporella umbellata1 Defrance 1823
Scaphopoda
 Dentaliida - Dentaliidae
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1, Lentigodentalium pseudomutabile1, Antalis dumasi1, Antalis aff. exlamarcki1, "Dentalium bouei" = Antalis bouei1
"Pseudantalis aturensis" = Dentalium (Pseudantilis)1 tusk shell
Antalis dumasi1 tusk shell
Antalis dumasi subsp.
"Dentalium bouei" = Antalis bouei1 Deshayes 1825 tusk shell
 Gadilida - Gadilidae
Gadila dingdenensis1, Cadulus aff. subfusiformis1
Bivalvia
 Nuculida - Nuculidae
Nuculoma haesendoncki1, Nucula (Nucula) aff. nitidosa1, Nucula (Nucula) nucleus1
Nuculoma haesendoncki1 nut clam
N. haesendoncki hanseata
Nucula (Nucula) aff. nitidosa1 Winckworth 1930 nut clam
Nucula (Nucula) nucleus1 Linnaeus 1758 nut clam
 Protobranchia - Nuculanidae
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1
"Nuculana (Saccella) westendorpi" = Saccella westendorpi1 Nyst and Westendorp 1839 pointed nut clam
 Protobranchia - Yoldiidae
Yoldia (Yoldia) glaberrima1, "Portlandia (Yoldiella) pygmaea" = Yoldiella1
Yoldia (Yoldia) glaberrima1 von Münster 1837 clam
"Portlandia (Yoldiella) pygmaea" = Yoldiella1 clam
 Arcida - Limopsidae
Limopsis (Limopsis) aurita1, "Limopsis (Pectunculina) anomala" = Limopsis1, Limopsis (Pectunculina) lamellata1
Limopsis (Limopsis) aurita1 Brocchi 1814 clam
"Limopsis (Pectunculina) anomala" = Limopsis1 clam
 Arcida - Glycymerididae
Glycymeris (Glycymeris) obovata1
Glycymeris (Glycymeris) obovata1 Lamarck 1819 clam
G. obovata baldii
 Arcida - Arcidae
Scapharca (Scapharca) diluvii1
 Pectinida - Pectinoidae
Pseudamussium lilli1, sensu lato Palliolum guersi n. sp.1, Aequipecten angelonii1, Aequipecten seniensis1, Korobkovia woodi1
Pseudamussium lilli1 Pusch 1837 scallop
sensu lato Palliolum guersi n. sp.1 Wienrich 1999 scallop
 Pectinida - Anomiidae
Pododesmus (Monia) anitae1
Pododesmus (Monia) anitae1 Janse and Janssen 1983 scallop
 Carditida - Carditidae
Cyclocardia sp.1
Cyclocardia sp.1 Conrad 1867 clam
 Carditida - Astartidae
Astarte (Astarte) radiata1, Goodallia (Goodallia) angulata1, Digitaria cf. beyschlageri1
Astarte (Astarte) radiata1 Nyst and Westendorp 1839 clam
Astarte radiata fa. radiata 274 specimens, Astarte radiata fa. convexior 56 specimens, Astarte radiata fa. goldfussi 32 specimens
Goodallia (Goodallia) angulata1 Lehmann 1885 clam
Digitaria cf. beyschlageri1 clam
[entered as Digitaria beyschlagi]
 Cardiida - Cardiidae
"Habecardium subturgidum" = Laevicardium (Habecardium)1, Habecardium dingdense1, Parvicardium ovale1, Parvicardium straeleni1
"Habecardium subturgidum" = Laevicardium (Habecardium)1 cockle
H. subturgidum subturgidum
Habecardium dingdense1 Lehmann 1885 cockle
 Cardiida - Tellinidae
Angulus (Laceolina) benedeni1
Angulus (Laceolina) benedeni1 Nyst and Westendorp 1839 tellin clam
A. benedeni fallax
 Cardiida - Semelidae
sensu lato Abra lehmanni1, Abra (? Abra) sorgenfreii1
 Cardiida - Mactridae
Spisula elliptica1
Spisula elliptica1 Brown 1827 clam
 Cardiida - Veneridae
"Gouldia minima" = Gouldia (Gouldia) minima1, Venus (Ventricoloidea) multilamella1, Pelecyora (Cordiopsis) polytropa1
"Gouldia minima" = Gouldia (Gouldia) minima1 Montagu 1803 venus clam
Venus (Ventricoloidea) multilamella1 Lamarck 1818 venus clam
V. (V.) multilamella multilamella
Pelecyora (Cordiopsis) polytropa1 venus clam
P. (C.) polytropa nysti
 Cardiida - Glossidae
Glossus (Glossus) lunulatus1
 Cardiida - Kelliellidae
Spaniodontella nitida1
 Cardiida - Montacutidae
Mioerycina coarctata1
Mioerycina coarctata1 Wood 1840 clam
 Cardiida - Lasaeidae
Aligena sp.1
Aligena sp.1 Lea 1843 aligena
 Pholadida - Corbulidae
Corbula (Varicorbula) gibba1
Corbula (Varicorbula) gibba1 Oliva 1792 clam
C. (V.) gibba gibba
 Pholadida - Myidae
Sphenia pusilla1
Sphenia pusilla1 Sorgenfrel 1958 softshell clam
 Hiatellida - Hiatellidae
"Hiatella (Hiatella) arctica" = Hiatella arctica1
"Hiatella (Hiatella) arctica" = Hiatella arctica1 Linnaeus 1767 clam
H. (H.) arctica arctica
 Lucinida - Lucinidae
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1, "Lucinoma borealis" = Lucinoma (Lucinoma) borealis1
"Cavilucina (Goniomyrtea) droueti" = Gonimyrtea droueti1 Nyst 1861 clam
C. droueti droueti
"Lucinoma borealis" = Lucinoma (Lucinoma) borealis1 Linnaeus 1767 clam
 Lucinida - Thyasiridae
Axinulus germanicus1
Axinulus germanicus1 Janssen 1972 clam
Gastropoda
 Heterostropha - Architectonicidae
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3, Architectonica (Architectonica) briarti3, Architectonica (Architectonica) simplex3, Architectonica (Architectonica) ariei n. sp.3
"Architectonica (Pseudotorinia) planulata" = Pseudotorinia3 snail
Architectonica (Architectonica) ariei n. sp.3 Wienrich and Janssen 2007 sundial
 Heterostropha - Ringiculidae
Ringicula (Ringiculina) buccinea3, Ringicula (Ringiculina) ventricosa3
 Heterostropha - Acteonidae
Acteon (Acteon) semistriatus3, Acteon (Acteon) "sp. 2"3, Crenilabium terebelloides3
Acteon (Acteon) semistriatus3 snail
spelled as 'Actaeon'
Acteon (Acteon) "sp. 2"3 Montfort 1810 snail
spelled as 'Actaeon'
 Heterostropha - Murchisonellidae
Ebala (Ebala) vandervlerki3
 Heterostropha - Pyramidellidae
Pyramidella (Voluspa) elata3, Pyramidella (Voluspa) plicosa3, Chrysallida (Parthenina) toftlundensis3, Syrnola hoernesi3, Syrnola neumayri3, Turbonilla (Turbonilla) koeneniana3, "Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3, Odostomia (Megastomia) tuexeni3
Turbonilla (Turbonilla) koeneniana3 snail
sensu JANSSEN
"Turbonilla (Pyrgolampros) pseudoterebralis" = Pyrgolampros3 snail
sensu JANSSEN
 Heterostropha - Mathildidae
"Mathilda (Fimbriatella) clathratula" = Fimbriatella3
"Mathilda (Fimbriatella) clathratula" = Fimbriatella3 snail
 Thecosomata - Limacinidae
Limacina miorostralis3, Limacina valvatina3
Limacina valvatina3 Reuss 1867 snail
 Thecosomata - Cavoliniidae
Vaginella austriaca3
 Opisthobranchia -
Roxania utriculus3
 Opisthobranchia - Retusidae
Retusa sp.3, Retusa (Cylichnina) elongata3, Volvulella (Volvulella) acuminata3
Retusa sp.3 Brown 1827 snail
 Opisthobranchia - Cylichnidae
Cylichna (Cylichna) pseudoconvoluta3, Bullichna paucistriata3, Scaphander grateloupi3
 Lepetellida - Lepetellidae
Lepetella compressiuscula2
 Sorbeoconcha - Epitoniidae
Opalia (Nodiscala) rugatina2, Opalia (Nodiscala) sp.2, "Amaea (Scalina) subreticula" = Scalina2, Epitonium (? Hyaloscala) "sp. 2"2
Opalia (Nodiscala) sp.2 De Boury 1890 wentletrap
"Amaea (Scalina) subreticula" = Scalina2 wentletrap
Epitonium (? Hyaloscala) "sp. 2"2 Bolten 1798 wentletrap
 Sorbeoconcha - Capulidae
Capulus ungaricus2
Capulus ungaricus2 Linnaeus 1758 snail
 Sorbeoconcha - Rissoidae
Ovirissoa gottscheana2, sensu lato Alvania antwerpiensis2
 Sorbeoconcha - Eulimidae
Niso (Niso) terebellum2, Balcis (Balcis) alba2, Eulima (Eulima) bilineata2, Eulima (Eulima) glabra2
Balcis (Balcis) alba2 snail
incl. 2specimens B. alba for. eichwaldi
 Sorbeoconcha - Naticidae
Naticidae indet.2, Sinum (Sinum) aquense2, "Natica neglecta" = Cochlis neglecta2, "Natica sensu lato tigrina" = Cochlis tigrina2, Polinices (Polinices) miocolligens2, Euspira gottschei2, Euspira mioaperta2, Euspira nysti2, Euspira staringi2, Euspira helicina2, Neverita josephinia2, Tectonatica miopusilla2
Naticidae indet.2 Guilding 1834 moon snail
Naticidae juv. et indet.
"Natica neglecta" = Cochlis neglecta2 Mayer 1858 moon snail
"Natica sensu lato tigrina" = Cochlis tigrina2 Röding 1798 moon snail
Euspira gottschei2 moon snail
Euspira mioaperta2 moon snail
Euspira nysti2 moon snail
Euspira staringi2 moon snail
Euspira helicina2 Brocchi 1814 moon snail
E. aggr. helicina
Neverita josephinia2 Risso 1826 moon snail
N. josephinia olla
 Sorbeoconcha - Vitrinellidae
Circulus hennei2
Circulus hennei2 Glibert 1952 snail
 Sorbeoconcha - Triviidae
Erato (Erato) germanica2
Erato (Erato) germanica2 Schilder 1929 snail
 Neogastropoda - Terebridae
Strioterebrum fabritzi3, Strioterebrum basteroti3, Terebra (Myurellina) acuminata3
Strioterebrum basteroti3 Nyst 1843 auger snail
Terebra (Myurellina) acuminata3 auger snail
T. acuminata neglecta
 Neogastropoda - Clavatulidae
Clavatula boreointerrupta3
 Neogastropoda - Mangeliidae
Sorgenfreispira sorgenfreii3, Sorgenfreispira tenella3, sensu lato Mangelia miorugulosa3, Benthomangelia aff. obtusangula3, Benthomangelia grippi3
Sorgenfreispira tenella3 Conrad 1848 snail
sensu lato Mangelia miorugulosa3 Kautsky 1925 snail
Benthomangelia grippi3 Anderson 1964 snail
 Neogastropoda - Turridae
Boreodrillia hosiusi3, Nitidiclavus maitrejus3, sensu lato Unedogemmula boreoturricula3, sensu lato Unedogemmula hanseata3, sensu lato Unedogemmula stoffelsii3, Acamptogenotia escheri3, Fusiturris aquensis3, Fusiturris duchastelii3, Gemmula (Gemmula) schafferi3, Gemmula (Gemmula) sp.3, Gemmula (Gemmula) denticula3, Gemmula (Gemmula) vermicularis3, Gemmula (Gemmula) zimmermanni3, sensu lato Inquisitor borealis3
Fusiturris duchastelii3 turrid
F. duchastelii flexiplicata
Gemmula (Gemmula) sp.3 Weinkauff 1875 turrid
Gemmula (Gemmula) denticula3 turrid
G. denticula borealis
 Neogastropoda - Borsoniidae
"Genota ramosa" = Genota valeriae3, Borsonia uniplicata3, Bathytoma jugleri3, Asthenotoma pannus3
"Genota ramosa" = Genota valeriae3 Hoernes and Auinger 1891 snail
Borsonia uniplicata3 Koenen 1872 snail
 Neogastropoda - Conidae
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3
"Conus (Conolithus) sensu lato antediluvianus" = Conus (Conolithes)3 cone shell
 Neogastropoda - Clathurellidae
? Ceritoturris hannoverana3, Pleurotomoides borealis3, Pleurotomoides luisae3, Pleurotomoides simplex3
 Neogastropoda - Drilliidae
Splendrillia selenkae3
 Neogastropoda - Raphitomidae
Favriella sinuosula3, Favriella "sp. 1"3, Metuonella grippi3, Teretia anceps3, Teretia "sp. 1"3
Favriella "sp. 1"3 Hornung 1920 snail
Teretia "sp. 1"3 Norman 1888 snail
 Neogastropoda - Cochlespiridae
Cochlespira corneti3
 Neogastropoda - Ancillariidae
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2
"Ancilla (Baryspira) obsoleta" = Amalda obsoleta2 Brocchi 1814 snail
 Neogastropoda - Olividae
Oliva (Strephona) dufresnei2
Oliva (Strephona) dufresnei2 Basterot 1825 olive snail
 Neogastropoda - Mitridae
Mitra (Tiara) grateloupi3
Mitra (Tiara) grateloupi3 d'Orbigny 1852 mitre shell
 Neogastropoda - Cancellariidae
Trigonostoma (Trigonostoma) aperta2, Trigonostoma (Trigonostoma) banardi2, Trigonostoma (Trigonostoma) protrigonostoma2, Babylonella fusiformis2, Aneurystoma canaliculatum2, Sveltia varicosa2
Sveltia varicosa2 snail
S. varicosa paucicostata
 Neogastropoda - Harpidae
Morum cf. dunkeri2
 Neogastropoda - Muricidae
Lyrotyphis (Eotyphis) fistulosus2, Typhis (Typhis) ? pungens2, Hadriania coelata2, "Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2, "Murex (Haustellum) inornatus" = Haustellum2
Typhis (Typhis) ? pungens2 Solander in Brander and Brander 1766 murex snail
Hadriania coelata2 Dujardin 1837 murex snail
"Murex (Tubicauda) spinicosta" = Murex (Promurex) spinicosta2 Bronn 1831 murex snail
"Murex (Haustellum) inornatus" = Haustellum2 murex snail
 Neogastropoda - Volutidae
Athleta (Athleta) ficulina2, Scaphella (Scaphella) bolli2
Athleta (Athleta) ficulina2 Lamarck 1811 volute
 Neogastropoda - Nassariidae
Nassarius bocholtensis2, Nassarius schroederi2, Nassarius sulcatus2, Nassarius cavatus2, Nassarius woodwardi2, Nassarius serraticosta2, Nassarius "sp. 2"2, Nassarius karinae2, "Nassarius andersoni" = Nassarius (Catilon) andersoni2, Amyclina facki2
Nassarius "sp. 2"2 Duméril 1806 snail
Nassarius karinae2 Wienrich 2001 snail
"Nassarius andersoni" = Nassarius (Catilon) andersoni2 Weaver 1912 snail
 Neogastropoda - Buccinidae
Phos (Phos) decussatus2
 Neogastropoda - Fasciolariidae
Streptodictyon sexcostatus2, "Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2, ? Exilifusus attenuatus2
"Streptolathyrus rothi" = Streptochetus (Streptolathyrus)2 snail
 Neogastropoda - Columbellidae
Anachis (Costoanachis) hosiusi2, Anachis (Costoanachis) terebralis2, Mitrella (Macrurella) nassoides2
 Neogastropoda - Costellariidae
"Vexillum (Uromitra) gliberti" = Uromitra3, Orthosurcula steinvorthi3
"Vexillum (Uromitra) gliberti" = Uromitra3 ribbed mitre
 Sorbeoconcha - Aporrhaidae
Aporrhais (Aporrhais) alata2
 Sorbeoconcha - Ficidae
Ficus (Ficus) conditus2
Ficus (Ficus) conditus2 Brongniart 1823 fig shell
 Sorbeoconcha - Xenophoridae
Xenophora deshayesi2
Xenophora deshayesi2 Michelotti 1847 carrier shell
 Sorbeoconcha - Cassidae
"Semicassis (Echinophoria) bicoronata" = Echinophoria2, "Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2
"Semicassis (Echinophoria) bicoronata" = Echinophoria2 snail
"Semicassis (Semicassis) miolaevigata" = Phalium (Semicassis) miolaevigata2 Sacco 1890 snail
 Sorbeoconcha - Ranellidae
"Charonia (Sassia) tarbelliana" = Sassia2
"Charonia (Sassia) tarbelliana" = Sassia2 triton shell
 Sorbeoconcha - Turritellidae
Turritella subangulata2
 Sorbeoconcha - Cerithiidae
Bittium (Bittium) tenuispina2, "Bittium (Bittium) spina" = Bittium spina2
"Bittium (Bittium) spina" = Bittium spina2 Partsch 1856 cerith snail
 Neotaenioglossa - Calyptraeidae
Calyptraea (Calyptraea) chinensis2
Calyptraea (Calyptraea) chinensis2 Linnaeus 1758 slipper shell
 Entomotaeniata - Cimidae
Cima neglecta3, Actaeopyramis elatus3
1Wienrich 1999; 2Wienrich 2001; 3Wienrich and Janssen 2007