Akespe (Oligocene of Kazakhstan)

Also known as Agyspe; Kumbulak Cliffs

Where: Kazakhstan (46.8° N, 60.6° E: paleocoordinates 47.6° N, 58.1° E)

• coordinate estimated from map

• small collection-level geographic resolution

When: Aral Formation, Late/Upper Oligocene (28.4 - 23.0 Ma)

• late Oligocene

• bed-level stratigraphic resolution

Environment/lithology: terrestrial; gray, green, yellow, calcareous shale and claystone

• grayish yellow-green calcareous shale and claystone

Size classes: macrofossils, mesofossils

Collected by N.S. Shevyreva in 1968

Primary reference: A. V. Lopatin. 1996. The Stratigraphy and Small Mammals from the Aral Formation, the Altynshokysu Locality, Northern Aral Area. Stratigraphy and Geological Correlation 4(2):65-79 [P. Mannion/P. Mannion]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 64413: authorized by Mark Uhen, entered by Mark Uhen on 08.09.2006, edited by Philip Mannion and Evangelos Vlachos

Creative Commons license: CC BY (attribution)

Taxonomic list

 Caudata - Urodela
Mioproteus sp.7 Estes and Darevsky 1978 mudpuppy
 Rodentia - Muridae
"Eumysodon spurius n. gen. n. sp." = Eumysodon spurius6, "Eumysodon orlovi n. sp." = Eumysodon spurius6, "Aralomys gigas n. sp." = Tachyoryctoides4, Tachyoryctoides obrutschewi6
"Eumysodon spurius n. gen. n. sp." = Eumysodon spurius6 Argyropulo 1939 mouse
PIN, no. 210/264 - type
"Eumysodon orlovi n. sp." = Eumysodon spurius6 Argyropulo 1939 mouse
PIN no. 210/264a - type
"Aralomys gigas n. sp." = Tachyoryctoides4 mouse
PIN, no. 210/263 - type; referred specimen: PIN, no. 210/772
Tachyoryctoides obrutschewi6 Bohlin 1937 mouse
 Rodentia - Cricetidae
Eumyarion tremulus3, Argyromys woodi n. sp.6, "Argyromys aralensis n. sp." = Argyromys aralensis6, "Protalactaga borissiaki n. sp." = Argyromys aralensis6, Eucricetodon aff. asiaticus6, Eucricetodon aff. caducus6
Eumyarion tremulus3 Lopatin 1996 mouse
PIN, no. 210/764
Argyromys woodi n. sp.6 mouse
PIN no. 210/262 (type), 765 and 766
"Argyromys aralensis n. sp." = Argyromys aralensis6 Argyropulo 1939 mouse
PIN, no. 210/261 - type
"Protalactaga borissiaki n. sp." = Argyromys aralensis6 Argyropulo 1939 mouse
PIN no. 210/260 - type
Eucricetodon aff. asiaticus6 Matthew and Granger 1923 mouse
Eucricetodon aff. caducus6 Shevyreva 1967 mouse
 Rodentia - Castoridae
Agnotocastor sp.6, Steneofiber sp.6, "Capatanka schokensis" = Propalaeocastor schokensis6, "Steneofiber kumbulakensis" = Propalaeocastor kumbulakensis6
Agnotocastor sp.6 Stirton 1935 beaver
Steneofiber sp.6 Geoffroy 1833 beaver
"Capatanka schokensis" = Propalaeocastor schokensis6 Bendukidze 1993 beaver
"Steneofiber kumbulakensis" = Propalaeocastor kumbulakensis6 Lytschev 1970 beaver
PIN no. 210/767-769
 Lagomorpha - Leporidae
Desmatolagus aff. schargaltensis6, "Agispelagus simplex n. gen. n. sp." = Desmatolagus simplex1
"Agispelagus simplex n. gen. n. sp." = Desmatolagus simplex1 Argyropulo 1940 rabbit
PIN, no. 210/609 - type
 Lagomorpha - Ochotonidae
Sinolagomys pachygnathus1 Li and Qiu 1980 pika
PIN, no. 210/773
 Lipotyphla - Erinaceidae
Amphechinus sp.3 Aymard 1850 hedgehog
PIN, no. 210/265 and 737
Amphechinus akespensis n. sp.2 Lopatin 1999 hedgehog
PIN, no. 210/735 - type; referred material: PIN, no. 210/736 and 210/774
Amphechinus minimus6 Bohlin 1942 hedgehog
Palaeoscaptor acridens6 Matthew and Granger 1924 hedgehog
 Perissodactyla - Paraceratheriidae
Paraceratherium sp.6 Forster Cooper 1911 odd-toed ungulate
Paraceratherium prohorovi6 odd-toed ungulate
 Perissodactyla - Rhinocerotidae
Gigantamynodon akespensis n. sp.5 Bayshashov 1993 rhinoceros
KAN PK 11/6 - type
Protaceratherium sp.6 Abel 1910 rhinoceros
 Perissodactyla - Eggysodontidae
Eggysodon sp.6 Roman 1910 odd-toed ungulate
 Ungulata - Lophiomerycidae
Lophiomeryx sp.6 Pomel 1853 ruminant
 Ungulata -
Prodremotherium sp.6 Filhol 1877 ruminant
 Ungulata - Palaeomerycidae
Amphitragulus sp.6 Pomel 1847 ruminant
 Ungulata - Moschidae
Micromeryx sp.6 Lartet 1851 musk deer
 Ungulata - Archaeomerycidae
Miomeryx sp.6 Matthew and Granger 1925 ruminant