Chapada do Araripe, Romualdo Member (general) (Cretaceous of Brazil)

Also known as Araripe Plateau

Where: Ceará, Brazil (7.2° S, 39.3° W: paleocoordinates 10.9° S, 9.4° W)

• coordinate estimated from map

• basin-level geographic resolution

When: Romualdo Member (Santana Formation), Early/Lower Albian (112.0 - 109.0 Ma)

• In recent stratigraphic schemes (e.g. Martill 2007) the Santana Formation and the "Romualdo Member" are essentially synonyms

•Santana Formation suggested to be early Albian in age by Heimhofer & Hochuli (2010) on the basis of palynomorphs

• member-level stratigraphic resolution

Environment/lithology: lagoonal or restricted shallow subtidal; concretionary, calcareous shale

• Limestone concretions in black shales

Size class: macrofossils

Preservation: concretion, soft parts

Reposited in the AMNH, BSP, SMNS

Collection methods: quarrying, chemical

Primary reference: L. I. Price. 1971. A presença de Pterosauria no Cretáceo Inferior da Chapada do Araripe, Brasil. Anais da Academia Brasileira de Ciências 43:451-461 [R. Butler/R. Butler]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 92290: authorized by Richard Butler, entered by Richard Butler on 05.11.2009, edited by Roger Benson, Philip Mannion, Mark Bell, Matthew Carrano and Jonathan Tennant

Creative Commons license: CC BY (attribution)

Taxonomic list

Chondrichthyes
 Hybodontiformes - Hybodontidae
Tribodus limae n. gen. n. sp.2
Tribodus limae n. gen. n. sp.2 Brito and Ferreira 1989 elasmobranch
Holotype: GP/2T-2 IG-USP
 Rajiformes - Rhinobatidae
Rhinobatos beurleni n. sp.20 Silva Santos 1968 guitarfish
Holotype: No. 2510 D e V; Paratype: No. 917-P, D e V
Actinopteri
 Pycnodontiformes - Pycnodontidae
Microdon penalvai n. sp.21 Silva Santos 1970
Holotype: No. 918, DGM - DNPM
 Lepisosteiformes -
Lepidotes wenzae3 Brito and Gallo 2003 gar
Holotype: MNHN-BCE 387
 Lepisosteiformes - Obaichthyidae
Obaichthys ? laevis29 Wenz and Brito 1992 gar
DGM 1155
Obaichthys decoratus n. gen. n. sp.29 Wenz and Brito 1992 gar
DGM 1336
 Halecomorphi - Ionoscopidae
Oshunia brevis30 Wenz and Kellner 1986
CD-P-31
Reptilia
 Testudines -
Brasilemys josai n. gen. n. sp.16
Brasilemys josai n. gen. n. sp.16 Lapparent de Broin 2000 sideneck turtle
Museu de Geologia de Barcelona, Catalunya, Spain, number MGB 37911. A carapace with a partial skull, two hyoid branches, left lower jaw; axis and 3d cervical vertebrae in the carapace
 Pterosauria -
Pterodactyloidea indet.19 pterosaur
DNPM D6M 488-LE, wing bones. AMNH 22569, pelvis. Geol. Inst. Univ. Amsterdam 4895, two cervical vertebrae. Gigase collection V-201, 10 articulated posterior cervical and anterior dorsal vertebrae. BSP 1987 I 65, right scapulocoracoid. SMNS 55408, humerus. SMNS 82001, ulna. SMNS 81976, humerus and ulna/radius. SMNS 55412, 55415, phalanges. SMNS 80437
Pterodactyloidea indet.15 pterosaur
BSP 1982 I 91, 1982 I 93
Santanadactylus pricei n. sp.27 Wellnhofer 1985 pterosaur
BSP 1980 I 122 (holotype), partial left forelimb. Paratypes: BSP 1980 I 43, 1980 I 120, MZ A/III 522, forelimb material. BSP 1987 I 1, partial left forelimb
cf. Santanadactylus pricei22 Wellnhofer 1985 pterosaur
SMNS 55883, humerus, 55410, ulna, 55411, ulna and radius
Araripesaurus castilhoi n. gen. n. sp. Price 1971 pterosaur
529-R, partial forelimb
 Pterosauria - Pteranodontidae
Santanadactylus spixi n. sp.27 Wellnhofer 1985 pterosaur
BSP 1980 I 121, partial forelimb
Santanadactylus brasilensis n. gen. n. sp.4 Buisonjé 1980 pterosaur
Geol. Inst. Univ. Amsterdam 4894 (holotype), humerus/scapula/coracoid
 Pterosauria -
Araripedactylus dehmi n. gen. n. sp.26 Wellnhofer 1977 pterosaur
BSPG 1975 I 166, proximal phalanx, right wing-finger
Azhdarchoidea indet.8 pterosaur
SMNK PAL 3985, MPSC R 859 (prob. different taxa)
 Pterosauria - Tapejaridae
Tapejara wellnhoferi n. gen. n. sp.13 Kellner 1989 pterosaur
Blittersdorff collection, CD-R-080, partial skull
Thalassodrominae indet.1 Kellner and Campos 2007 pterosaur
AMNH 22567, three posteriormost cervical vertebrae, first seven dorsals, (fused in to a notarium), right and left scapulocoracoids, a partial sternum, incomplete right humerus, part of the wing finger, incomplete right femur and parts of the right tibia/fibula
 Pterosauria - Thalassodromidae
Tupuxuara longicristatus n. gen. n. sp.14
Tupuxuara longicristatus n. gen. n. sp.14 Kellner and Campos 1988 pterosaur
 Pterosauria - Dsungaripteridae
Banguela oberlii n. gen. n. sp.9
Banguela oberlii n. gen. n. sp.9 Headden and Campos 2014 pterosaur
SAO 251093, mandible
 Pterosauria -
Brasileodactylus sp.23, Brasileodactylus araripensis n. gen. n. sp.11, "cf. Criorhynchus mesembrinus" = Tropeognathus mesembrinus22, "Tropeognathus mesembrinus n. gen. n. sp." = Tropeognathus mesembrinus28, Cearadactylus atrox n. gen. n. sp.17, Anhanguera sp.25, "Santanadactylus araripensis n. sp." = Anhanguera araripensis27, "Santanadactylus cf. araripensis" = Anhanguera araripensis27, Anhanguera blittersdorffi n. gen. n. sp.5, "cf. Coloborhynchus spielbergi" = Anhanguera spielbergi22, "Coloborhynchus spielbergi n. sp." = Anhanguera spielbergi24, "Araripesaurus santanae n. sp." = Anhanguera santanae27, "? Cearadactylus ligabuei n. sp." = Anhanguera ligabuei6, "Tropeognathus robustus n. sp." = Anhanguera robustus28
Brasileodactylus sp.23 Kellner 1984 pterosaur
AMNH 24444, skull and partial left wing
Brasileodactylus araripensis n. gen. n. sp.11 Kellner 1984 pterosaur
MN 4804–V, anterior part of mandible
"cf. Criorhynchus mesembrinus" = Tropeognathus mesembrinus22 Wellnhofer 1987 pterosaur
SMNS 56994, mandible
"Tropeognathus mesembrinus n. gen. n. sp." = Tropeognathus mesembrinus28 Wellnhofer 1987 pterosaur
BSP 1987 I 46, skull and lower jaw
Cearadactylus atrox n. gen. n. sp.17 Leonardi and Borgomanero 1985 pterosaur
F-PV-93 (holotype), incomplete skull with mandibles
Anhanguera sp.25 Campos and Kellner 1985 pterosaur
SAO 200602, mandible
"Santanadactylus araripensis n. sp." = Anhanguera araripensis27 Wellnhofer 1985 pterosaur
BSP 1982 I 89, skull, lower jaws, partial forelimb. BSP 1987 I 66, right metacarpal. SMNS 55409, humerus. SAO 16494, skull
"Santanadactylus cf. araripensis" = Anhanguera araripensis27 Wellnhofer 1985 pterosaur
BSP 1982 I 92, forelimb material
Anhanguera blittersdorffi n. gen. n. sp.5 Campos and Kellner 1985 pterosaur
Blittersdorff collection, complete skull
"cf. Coloborhynchus spielbergi" = Anhanguera spielbergi22 Veldmeijer 2003 pterosaur
SMNS 55413, ulna
"Coloborhynchus spielbergi n. sp." = Anhanguera spielbergi24 Veldmeijer 2003 pterosaur
RGM 401 880, skull and postcrania
"Araripesaurus santanae n. sp." = Anhanguera santanae27 Wellnhofer 1985 pterosaur
BSP 1982 I 90, partial skull and partial forelimb
"? Cearadactylus ligabuei n. sp." = Anhanguera ligabuei6 Dalla Vecchia 1993 pterosaur
CSRL 12692/12713 (holotype), anterior part of a skull
"Tropeognathus robustus n. sp." = Anhanguera robustus28 Wellnhofer 1987 pterosaur
BSP 1987 I 47, lower jaw
 Pterosauria - Ornithocheiridae
Barbosania gracilirostris n. gen. n. sp.7
Barbosania gracilirostris n. gen. n. sp.7 Elgin and Frey 2011 pterosaur
MNHS/00/85, skull and partial postcranium
 Loricata -
Araripesuchus gomesii10 Price 1959 crocodilian
AMNH 24450 - complete articulated skeleton
 Loricata - Peirosauridae
Caririsuchus camposi n. gen. n. sp.12
Caririsuchus camposi n. gen. n. sp.12 Kellner 1987 crocodilian
CD-R-041 & DGM 1468 R - partial skeleton
Santanaclupea silvasantosi
  -
Santanaclupea silvasantosi n. gen. n. sp.18
Santanaclupea silvasantosi n. gen. n. sp.18 Maisey 1993
AMNH 12790, complete specimen; AMNH 12789