Soyana (PIN collection 2334): Roadian, Russian Federation
collected by M. B. Edemskij 1929

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Protelytroptera
Arctocoleus ivensis n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2334/55 (1 measurement)
Insecta - Hypoperlida - Hypoperlidae
Boreopsocus danksae Shcherbakov 1994
Shcherbakov 1994 3 specimens
PIN 2334/395, 400, 407
Insecta - Hemiptera - Coleoscytidae
Coleoscyta rotundata n. gen., n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/107 (1 measurement)
Coleoscytodes elytrata Martynov 1935
Martynov 1935 1 individual
PIN 2334/91 (1 measurement)
Insecta - Hemiptera - Scytinopteridae
Scytinoptera sp. Handlirsch 1904
Martynov 1935 1 individual
PIN 2334/159
Anomoscyta reducta Martynov 1928
Martynov 1935 3 individuals
recombined as Scytinoptera reducta
PIN 2334/176; 88 (S. ivensis in Martynov), 124 (both in Becker-Migdisova,1948) (1 measurement)
Scytinoptera ivensis n. sp. Martynov 1935
Martynov 1935 2 individuals
synonym of Scytinoptera reducta
PIN 2334/100, 87 (2 measurements)
Insecta - Hemiptera - Archescytinidae
Sojanoscytina minor n. sp. Martynov 1933
Martynov 1933 1 specimen
PIN 2334/119 (1 measurement)
Insecta - Hemiptera - Boreoscytidae
Sojanoscytina latipennis n. sp. Martynov 1933
Becker-Migdisova 1949 1 individual
recombined as Boreoscyta latipennis
PIN 2334/129 (1 measurement)
Insecta - Hemiptera - Prosbolopseidae
Permopibrocha ramosa n. gen., n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/141 (1 measurement)
Prosbolopsis sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/110 (1 measurement)
Prosbolopsis ovalis n. gen., n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/130 (1 measurement)
Prosbolopsis simplex n. sp. Martynov 1935
Martynov 1935 3 individuals
synonym of Prosbolopsis ovalis
PIN 2334/95, 99, 96 (1 measurement)
Insecta - Hemiptera - Prosbolidae
Permocicada informal sp 1
Martynov 1935 1 individual
PIN 2334/127 (1 measurement)
Permocicada simulatrix n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/83 (1 measurement)
Permocicada nigronervosa Martynov 1928
Becker-Migdisova 1940 1 specimen
PIN 2334/111 (1 measurement)
Permocicada elliptica n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/122 (1 measurement)
Sojanoneura aberrans n. sp. Martynov 1935
Martynov 1935 1 individual
recombined as Permocicadopsis aberrans
PIN 2334/79 (1 measurement)
Sojanoneura sp. Martynov 1928
Martynov 1935 3 individuals
PIN 2334/78, 134, 125 (3 measurements)
Sojanoneura edemskii Martynov 1928
Martynov 1935 1 individual
PIN 2334/89 (1 measurement)
Sojanoneura bimaculata n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/150 (1 measurement)
Prosbole informal sp 1
Martynov 1935 1 individual
PIN 2334/155 (1 measurement)
Prosbole informal sp 2
Martynov 1935 1 individual
PIN 2334/137 (1 measurement)
Prosbole informal sp 3
Martynov 1935 1 individual
PIN 2334/94
Prosbole sojanensis n. sp. Martynov 1935
Martynov 1935 2 individuals
PIN 2334/145, 2334 (2 measurements)
Prosbole jucunda n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/128 (1 measurement)
Prosbole ovalis n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/163 (1 measurement)
Prosbole reducta n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2334/109 (1 measurement)
Insecta - Caloneurodea - Caloneuridae
Euthygramma aberrans n. sp. Martynov 1938
Martynov 1938 1 individual
synonym of Euthygramma parallellum
PIN 2334/313 (1 measurement)
Insecta - Paoliida - Paoliidae
Sojanidelia maculosa Storozhenko 1992
Storozhenko 1992 1 specimen
PIN 2334/277(8)
Sojanidelia nana Storozhenko 1992
Storozhenko 1992 1 specimen
PIN 2334/287
Insecta - Glosselytrodea - Jurinidae
Jurina marginata Martynov 1938
Martynov 1938 2 specimens
PIN 2334/348, 349
Insecta - Coleoptera - Permosynidae
Permocrossos elongatus n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2334/51 (1 measurement)
Insecta - Mecoptera - Parachoristidae
Parachorista opposita n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2334/171 (1 measurement)
Insecta - Mecoptera - Permochoristidae
Petromantis robusta Martynov 1932
Martynov 1932 1 individual
PIN 2334/21(167) (1 measurement)
Petromantis borealis Martynov 1928
Martynov 1932 17 individuals
synonym of Petromantis rossica
PIN 2334/16, 67, 59, 36, 18, 40, 176, 11, 193, 34, 189, 193, 75, 71, 189, 60, 37
Petromantis punctata Martynov 1932
Martynov 1932 1 individual
synonym of Petromantis rossica
PIN 2334/187
Permochorista maculipennis n. sp. Martynov 1932
Martynov 1932 2 individuals
recombined as Petromantis maculipennis
PIN 2334/172, second specimen is PIN 2334/47; var. major n. var. (2 measurements)
Petrochorista dubia n. sp. Martynov 1932
Martynov 1932 2 individuals
recombined as Petromantis dubia
PIN 2334/66; 2334/188 in Martynova, 1948 (1 measurement)
? Permochorista dubiosa n. sp. Martynov 1932
Martynov 1932 1 individual
recombined as Petromantis dubiosa
PIN 2334/35 (1 measurement)
Agetochorista ornata n. sp. Martynov 1932
Martynov 1932 1 individual
recombined as Agetopanorpa ornata
PIN 2334/38 (1 measurement)
Agetochorista splendida Martynov 1932
Martynov 1932 1 individual
synonym of Agetopanorpa ornata
PIN 2334/19 (1 measurement)
Agetochorista dubia n. sp. Martynov 1932
Martynov 1932 1 individual
recombined as Agetopanorpa dubia
PIN 2334/63 (1 measurement)
Agetochorista ? fasciata Martynov 1932
Martynov 1932 1 individual
recombined as Agetopanorpa fasciata
PIN 2334/10 (1 measurement)
Insecta - Mecoptera - Permopanorpidae
Permopanorpa angustata n. sp. Martynov 1932
Martynov 1932 2 individuals
recombined as Martynopanorpa angustata
PIN 2334/350, 255 (2 measurements)
Permopanorpa tenuis n. sp. Martynov 1932
Martynov 1932 1 individual
replaced by Permopanorpa martynovi
PIN 2334/178 (1 measurement)
Insecta - Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis Martynov 1932
Martynov 1932 1 individual
PIN 2334/8
Insecta - Permithonidae
Permithonopsis proxima n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Permithonopsis ivensis
PIN 2334/56 (1 measurement)
Permegalomus maculipennis n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2334/68 (1 measurement)
Permosisyra latipennis n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2334/43 (1 measurement)
Insecta - Palaeodictyoptera - Eukulojidae
Kuloja paurovenosa n. sp. Martynov 1932
1 individual
synonym of Eukuloja cubitalis
PIN 2334/1 (1 measurement)
Insecta - Palaeodictyoptera - Permothemistidae
Palaeothemis libelluloides n. gen., n. sp. Martynov 1932
1 individual
recombined as Permothemis libelluloides
PIN 2334/7 (1 measurement)
Insecta - Reculida - Ivapteridae
Ivaptera sharovi Aristov 2009
Aristov 2008 2 individuals
PIN 2334/209, 234
Insecta - Ephemeroptera - Protereismidae
Protereismidae indet. Handlirsch 1919
1 individual
PIN 2334/5
Insecta - Permaeschnidae
Permaeschna dolloi n. gen., n. sp. Martynov 1931
Martynov 1931 1 specimen
PIN 2334/3(2) (1 measurement)
Permaeschna proxima n. sp. Martynov 1931
Martynov 1931 1 specimen
synonym of Permaeschna dolloi
PIN 2334/4 (1 measurement)
Insecta - Meganisoptera - Meganeuridae
Arctotypus sinuatus n. gen., n. sp. Martynov 1932
1 individual
PIN 2334/5 (1 measurement)
Insecta - Eoblattida - Idelinellidae
Permostriga sp. Novokshonov 1999
Aristov and Rasnitsyn 2012 1 specimen
PIN 2334/411 (different from Bekker-Migdisova specimen?)
see common names

Geography
Country:Russian Federation State/province:Arkangelsk
Coordinates: 65.6° North, 43.7° East (view map)
Paleocoordinates:33.2° North, 33.1° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian Epoch:Guadalupian
Stage:Roadian 10 m.y. bin:Permian 3
Key time interval:Roadian
Age range of interval:272.30000 - 268.80000 m.y. ago
Stratigraphy
Formation:Iva-Gora Beds
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology:fine lithified sandstone
Lithology description: "fine granulous sandstone" Martynov, 1932
Environment:lagoonal Tectonic setting:foreland basin
Geology comments: Probable brackish lagoon with some access to sea
Taphonomy
Modes of preservation:mold/impression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:survey of museum collection
Collection size:297
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:M. B. Edemskij Collection dates:1929
Metadata
Database number:104993
Authorizer:M. Clapham Enterer:M. Clapham, J. Karr
Modifier:M. Clapham Research group:paleoentomology
Created:2011-02-13 04:33:37 Last modified:2013-11-25 13:02:06
Access level:the public Released:2011-02-13 04:33:37
Creative Commons license:CC BY
Reference information

Primary reference:

36456. A. V. Martynov. 1932. New Permian Palaeoptera with the discussion of some problems of their evolution. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 1:1-44 [M. Clapham/J. Karr/M. Clapham]

Secondary references:

34929 D. S. Aristov. 2008. New Grylloblattids of the Family Megakhosaridae (Insecta: Grylloblattida) from the Permian of Russia. Paleontological Journal 42(3):269-272 [M. Clapham/J. Karr]
39962 D. S. Aristov and A. P. Rasnitsyn. 2012. Revision of the family Idelinellidae, with a review of the Permian Eoblattida (Insecta). Paleontological Journal 46:49-60 [M. Clapham/M. Clapham]
39941 E. E. Becker-Migdisova. 1940. Iskopaemye permskie cikady semejstva Prosbolidae s reki sojany. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 11(2):1-95 [M. Clapham/J. Karr/M. Clapham]
48456 E. E. Becker-Migdisova. 1948. Permskie tsikady semeystva Scytinopteridae s reki Soyany. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 15:1-42 [M. Clapham/M. Clapham]
37431 E. E. Becker-Migdisova. 1949. Novoe permskoe semeistvo Boreoscytidae i vopros o filogenii predkov Homoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 20:171-182 [M. Clapham/J. Karr/M. Clapham]
44086 A. V. Martynov. 1931. On some new remarkable Odonata from the Permian of Archangelsk district. Bulletin de l'Académie des Sciences de l'URSS, Classe des Sciences Mathématiques et Naturelles (1)141-147 [M. Clapham/J. Karr/M. Clapham]
37589 A. V. Martynov. 1932. Permian fossil Insects from the Arkhangelsk district. Part I. The order Mecoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 2:23-62 [M. Clapham/J. Karr/M. Clapham]
37594 A. V. Martynov. 1932. Permian fossil Insects from the Arkhangelsk district. Part II. Neuroptera, Megaloptera and Coleoptera, with the description of two new beetles from Tikhie Gory. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 2:63-96 [M. Clapham/J. Karr/M. Clapham]
47241 A. V. Martynov. 1933. On the Permian family Archescytinidae (Homoptera) and its relationships. Bulletin de l'Academie des sciences de l'URSS, Leningrad 1933:883-894 [M. Clapham/J. Karr]
37595 A. V. Martynov. 1935. Permian fossil Insects from the Arkhangelsk district. Part 5. Homoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 4:1-35 [M. Clapham/J. Karr/M. Clapham]
37611 A. V. Martynov. 1938. Permian fossil Insects from the Arkhangelsk district. Part V. The family Euthygrammidae and its relationships (with the description of a new genus and family from Chekarda). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7(3):69-80 [M. Clapham/J. Karr/M. Clapham]
53604 A. V. Martynov. 1938. O novom Permskom otryade pryamokrylykh nasekomykh Glosselytrodea. Izvestiya Akademii Nauk SSSR 1938:187-206 [M. Clapham/M. Clapham]
35137 A. Nel, G. Fleck, R. Garrouste, G. Gand, J. Lapeyrie, S. M. Bybee, and J. Prokop. 2009. Revision of Permo-Carboniferous griffenflies (Insecta: Odonatoptera: Meganisoptera) based upon new species and redescription of selected poorly known taxa from Eurasia. Palaeontographica Abteilung A 289(4-6):89-121 [M. Clapham/J. Karr]
39083 D. E. Shcherbakov. 1994. A new genus of the Paleozoic order Hypoperlida. Russian Entomological Journal 3:33-36 [M. Clapham/M. Clapham]
50618 S. Storozhenko. 1992. Permian fossil insects of north-east Europe: new and little-known Ideliidae (Insecta, Plecopteroidea, Grylloblattida). Entomologica Fennica 3:21-39 [M. Clapham/M. Clapham/M. Clapham]