Madygen, PIN collection 2069, Dzhailoucho area: Ladinian, Kyrgyzstan

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Eoblattida - Blattogryllidae
Costatoviblatta aenigmatosa Storozhenko 1992
Storozhenko 1992 9 specimens
PIN 2069/1619, 1627, 1633, 1699, 1736, 1741, 1797, 1799, 1855
Insecta - Eoblattida - Mesorthopteridae
Parastenaropodites longiuscula Storozhenko 1996
Storozhenko 1996 6 specimens
Parastenaropodites n. gen. fluxa Storozhenko 1996
Storozhenko 1996 15 specimens
(1 measurement)
Mesoidelia faceta Storozhenko 1996
Storozhenko 1996 1 specimen
PIN 2069/1485
Parastenaropodites nervosa Storozhenko 1996
Storozhenko 1996 3 specimens
recombined as Austroidelia nervosa
Austroidelia asiatica Storozhenko 1996
Storozhenko 1996 1 specimen
Insecta - Eoblattida - Megakhosaridae
Megablattogryllus pinguis Storozhenko 1990
Storozhenko 1990 1 specimen
PIN 2069/1403
Microblattogryllus variabilis Storozhenko 1990
Storozhenko 1990 1 specimen
recombined as Protoblattogryllus variabilis
PIN 2069/1714
Megakhosarodes paulivenosus Storozhenko 1993
Storozhenko 1993 1 specimen
PIN 2069/1667
Madygenocephalus micropteron n. gen., n. sp. Aristov 2011
Aristov 2011 1 specimen
PIN 2069/1518 (2 measurements)
Insecta - Reculida - Geinitziidae
Shurabia minuta n. sp. Rasnitsyn 1982
Rasnitsyn 1982 25 specimens
PIN 2069/1742, 1678 (2 measurements)
Shurabia minutissima n. sp. Aristov 2022
Aristov 2022 1 specimen
PIN 2069/1495 (1 measurement)
Insecta - Blattaria - Caloblattinidae
Thuringoblatta sogdianensis n. sp. Vishnyakova 1998
Vishnyakova 1998 6 specimens
PIN 2069/41, 42, 88, 139, 187, 963 (1 measurement)
Insecta - Permosialidae
Permosialis triassica Novokshonov and Zhuzhgova 2004
Aristov and Rasnitsyn 2023 1 specimen
PIN 2069/3592
Insecta - Orthoptera - Xenopteridae
Ferganotriassia lata n. gen., n. sp. Gorochov 1989
Gorochov 1989 1 specimen
PIN 2069/2266 (1 measurement)
Triassoferganella angusta n. gen., n. sp. Gorochov 1989
Gorochov 1989 1 specimen
PIN 2069/2239 (1 measurement)
Pseudoferganopsis zini n. gen., n. sp. Gorochov 1989
Gorochov 1989 1 specimen
PIN 2069/2178 (1 measurement)
Ferganopterus clarus Sharov 1968
text says 121 imprints doesn't specify collection
Insecta - Orthoptera - Mesoedischiidae
Mesoedischia kirgizica n. sp. Gorochov 1987
Gorochov 1987 1 specimen
PIN 2069/2318 (1 measurement)
Mesoedischia obliqua Gorochov 1987
Gorochov 1987 1 specimen
PIN 2069/2319
Insecta - Orthoptera - Bintoniellidae
Eubintoniella ferganica n. gen., n. sp. Gorochov 1987
Gorochov 1987 1 specimen
PIN 2069/2327 (1 measurement)
Stenoshiella angusta n. gen., n. sp. Gorochov 1987
Gorochov 1987 1 specimen
PIN 2069/2325 (1 measurement)
Oshiella crassa n. gen., n. sp. Gorochov 1987
Gorochov 1987 1 specimen
recombined as Probintoniella crassa
PIN 2069/2321 (1 measurement)
Oshiella oblonga n. sp. Gorochov 1987
Gorochov 1987 1 specimen
recombined as Probintoniella oblonga
PIN 2069/2326 (1 measurement)
Probintoniella triassica n. gen., n. sp. Sharov 1968
28 specimens
PIN 2069/2320 (1 measurement)
Paroshiella alia n. gen., n. sp. Gorochov 1987
Gorochov 1987 1 specimen
PIN 2069/2324 (1 measurement)
Insecta - Orthoptera - Proparagryllacrididae
Brevibatkenella abscisa n. gen., n. sp. Gorochov 1987
Gorochov 1987 1 specimen
synonym of Gryllacrimima perfecta
PIN 2069/2240 (1 measurement)
Parafergania sharovi n. gen., n. sp. Gorochov 1987
Gorochov 1987 1 specimen
PIN 2069/2341 (1 measurement)
Insecta - Orthoptera - Permelcanidae
Meselcana madygenica n. gen., n. sp. Sharov 1968
100 specimens
PIN 2069/2409 (1 measurement)
Insecta - Orthoptera - Gryllavidae
Gryllavus madygenicus Sharov 1968
1 specimen
PIN 2069/2429
Insecta - Orthoptera - Haglidae
Tinnihagla handlirschi n. sp. Gorochov 1986
Gorochov 1986 1 specimen
PIN 2069/2418 (1 measurement)
Proisfaroptera martynovi n. sp. Sharov 1968
1 specimen
PIN 2069/2440, 2360 (1 measurement)
Protshorkuphlebia triassica Sharov 1968
1 specimen
PIN 2069/2352
Cantohagla gracilis n. gen., n. sp. Gorochov 1986
Gorochov 1986 1 specimen
PIN 2069/2442 (1 measurement)
Maragella reducta n. gen., n. sp. Gorochov 1986
Gorochov 1986 1 specimen
PIN 2069/2374 (1 measurement)
Insecta - Orthoptera - Locustavidae
Brevilocustavus microscopicus n. sp. Gorochov 2005
Gorochov 2005 1 specimen
PIN 2069/2329 (1 measurement)
Insecta - Plecoptera - Perlariopseidae
Dicronemoura dira n. sp. Sinitshenkova 1987
Sinitshenkova 1987 2 specimens
PIN 2069/1909, 1906
Dicronemoura acaulis Sinitshenkova 1987
Sinitshenkova 1987 3 specimens
PIN 2069/1905, 1908, 1910
Fritaniopsis remota Sinitshenkova 1987
Sinitshenkova 1987 1 specimen
PIN 2069/1914
Tritaniella synneura Sinitshenkova 1987
Sinitshenkova 1987 3 specimens
PIN 2069/1902, 1906, 1913
Tritaniella mera Sinitshenkova 1987
Sinitshenkova 1987 2 specimens
PIN 2069/1907, 1915
Ramonemoura constricta n. gen., n. sp. Sinitshenkova 1987
Sinitshenkova 1987 1 specimen
PIN 2069/1899 (1 measurement)
Insecta - Grylloblattodea - Madygenophlebiidae
Madygenophlebia nana n. sp. Storozhenko 1992
Storozhenko 1992 2 specimens
PIN 2069/1640,1609 (2 measurements)
Insecta - Grylloblattodea - Gorochoviidae
Gorochoviella n. gen. conjuncta Storozhenko 1994
Storozhenko 1994 1 specimen
PIN 2069/1846?
Gorochovia bifurca Storozhenko 1994
Storozhenko 1994 1 specimen
PIN 2069/1846?
Gorochovia minuta n. sp. Storozhenko 1994
Storozhenko 1994 11 specimens
(2 measurements)
Gorochovia n. gen. individua Storozhenko 1994
Storozhenko 1994 5 specimens
PIN 2069/1404,1590,1772,1783,1884
Pseudoliomopterites lucidus n. sp. Storozhenko 1994
Storozhenko 1994 1 specimen
PIN 2069/1482 (1 measurement)
Pseudoliomopterites obscurus n. gen., n. sp. Storozhenko 1994
Storozhenko 1994 1 specimen
PIN 2069/1805 (1 measurement)
Insecta - Hemiptera - Maguviopseidae
Maguviopsis kotchnevi Becker-Migdisova 1953
Shcherbakov 2011 1 specimen
PIN 2069/1724
Insecta - Hemiptera - Mesojabloniidae
Mesojablonia kukalovae n. gen., n. sp. Storozhenko 1992
Storozhenko 1992 1 specimen
PIN 2069/1677 (1 measurement)
Insecta - Hemiptera - Paraknightiidae
Triknightia mira n. gen., n. sp. Shcherbakov 2021
Shcherbakov 2021 1 specimen
PIN 2069/3278 (2 measurements)
Insecta - Hemiptera - Pincombeidae
Madygenopsyllidium djailautshoense n. gen., n. sp. Becker-Migdisova 1985
Becker-Migdisova 1985 1 specimen
PIN 2069/3204
Insecta - Hymenoptera - Xyelidae
Triassoxyela grandipennis n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
recombined as Asioxyela grandipennis
PIN 2069/3 (1 measurement)
Asioxyela parvula n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
PIN 2069/6 (1 measurement)
Asioxyela smilodon n. gen., n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
PIN 2069/5 (2 measurements)
Euryxyela euryptera n. gen., n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
PIN 2069/8 (1 measurement)
Triassoxyela orycta Rasnitsyn 1969
Rasnitsyn 1969 1 specimen
PIN 2069/4 (paratype of Triassoxyela kirghizica Rasnitsyn, 1964)
Triassoxyela kirgizica n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
PIN 2069/1 (1 measurement)
Xyelinus angustiradius n. gen., n. sp. Rasnitsyn 1964
Rasnitsyn 1964 2 specimens
PIN 2069/10, 11 (1 measurement)
Asioxyela antiqua n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
recombined as Leioxyela antiqua
PIN 2069/7 (1 measurement)
Euryxyela lata n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
recombined as Leioxyela lata
PIN 2069/9 (1 measurement)
Triassoxyela striata n. sp. Rasnitsyn 1964
Rasnitsyn 1964 1 specimen
recombined as Xiphoxyela striata
PIN 2069/2 (2 measurements)
Insecta - Mecoptera - Parachoristidae
Parachorista sana Novokshonov 2001
Novokshonov 2001 1 specimen
PIN 2069/3442 (1 measurement)
Insecta - Mecoptera - Mesopsychidae
Prochoristella ignara Novokshonov 1997
Novokshonov 1997 1 specimen
PIN 2069/3394
Insecta - Mecoptera - Orthophlebiidae
Choristopanorpa opinata Novokshonov 2001
Novokshonov 2001 1 specimen
PIN 2069/3460
Insecta - Mecoptera - Permochoristidae
Mecolusor confusicius n. gen., n. sp. Novokshonov 2001
Novokshonov 2001 1 specimen
PIN 2069/3391 (1 measurement)
Mesageta rieki Novokshonov 1997
Novokshonov 1997 1 specimen
PIN 2069/3525
Insecta - Coleoptera - Triaplidae
Catinoides rotundatus n. gen., n. sp. Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1358 (1 measurement)
Triassocatinius brachynotus n. sp. Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1291 (1069 is error) (1 measurement)
Triaplus laticoxa Ponomarenko 1977
Arnoldi et al. 1977 2 specimens
PIN 2069/1228, 1294
Avocatinus elongatus Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1300
Insecta - Coleoptera - Ademosynidae
Cephalosyne capitata Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1342
Dolichosyne confragosa n. gen., n. sp. Ponomarenko 1969
Ponomarenko 1969 2 specimens
PIN 2069/1336, 1340 (1 measurement)
Ademosyne kirghizica n. sp. Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1369 (1 measurement)
Insecta - Coleoptera - Triadocupedidae
Cupesia monilicornia n. gen., n. sp. Ponomarenko 1966
Ponomarenko 1966 1 specimen
PIN 2069/1115 (2 measurements)
Triadocupes ferghanensis n. gen., n. sp. Ponomarenko 1966
Ponomarenko 1966 7 specimens
PIN 2069/1098 (2 measurements)
Triadocupes latus n. sp. Ponomarenko 1966
Ponomarenko 1966 4 specimens
PIN 2069/1101 (2 measurements)
Triadocupes ellipticus n. sp. Ponomarenko 1966
Ponomarenko 1966 7 specimens
PIN 2069/1111 (2 measurements)
Platycupes dolichocerus n. gen., n. sp. Ponomarenko 1966
Ponomarenko 1966 1 specimen
PIN 2069/1107 (2 measurements)
Platycupes reticulatus n. sp. Ponomarenko 1966
Ponomarenko 1966 4 specimens
PIN 2069/1063 (2 measurements)
Platycupes pusillus n. sp. Ponomarenko 1966
Ponomarenko 1966 17 specimens
PIN 2069/1099 (2 measurements)
Insecta - Coleoptera
Rhabdocupes minor n. sp. Ponomarenko 1966
Ponomarenko 1966 7 specimens
PIN 2069/1039 (2 measurements)
Rhabdocupes longus n. gen., n. sp. Ponomarenko 1966
Ponomarenko 1966 4 specimens
PIN 2069/1041 (2 measurements)
Insecta - Coleoptera - Phoroschizidae
Lethocoleus sternalis n. gen., n. sp. Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1273 (1 measurement)
Schizophoroides rugosus n. gen., n. sp. Ponomarenko 1969
Ponomarenko 1969 1 specimen
PIN 2069/1290 (1 measurement)
Insecta - Coleoptera - Tricoleidae
Tricoleodes acutus n. gen., n. sp. Ponomarenko 1969
Ponomarenko 1969 2 specimens
PIN 2069/1194, 1196 (1 measurement)
Insecta - Coleoptera - Cupedidae
Mesocupoides proporeius Ponomarenko 1969
Ponomarenko 1969 2 specimens
misspelling Kirghizocupes proporeius
PIN 2069/1053, 1034
Kirghizocupes cellulosus n. gen., n. sp. Ponomarenko 1966
Ponomarenko 1966 6 specimens
PIN 2069/1071 (2 measurements)
Pterocupes leptocerus n. sp. Ponomarenko 1966
Ponomarenko 1966 1 specimen
PIN 2069/1126 (2 measurements)
Pterocupes antennatus n. gen., n. sp. Ponomarenko 1966
Ponomarenko 1966 3 specimens
PIN 2069/1076 (2 measurements)
Insecta - Coleoptera - Ommatidae
Notocupoides fasciatus n. sp. Ponomarenko 1966
Ponomarenko 1966 5 specimens
PIN 2069/1072 (2 measurements)
Notocupoides capitatus n. sp. Ponomarenko 1966
Ponomarenko 1966 2 specimens
PIN 2069/1121 (2 measurements)
Insecta - Coleoptera - Obrieniidae
Obrienia illaetabilis Zherikhin and Gratshev 1993
Zherikhin and Gratshev 1993 1 specimen
PIN 2069/3550 (2 measurements)
Insecta - Paurophlebiidae
Cladophlebia parvula Pritykina 1981
Pritykina 1981 2 specimens
PIN 2069/29, 38
Paurophlebia angusta Pritykina 1981
Pritykina 1981 1 specimen
PIN 2069/31
Insecta - Kennedyidae
Kennedya carpenteri Pritykina 1981
Pritykina 1981 1 specimen
PIN 2069/37
Insecta - Batkeniidae
Triassoneura primitiva n. sp. Pritykina 1981
Pritykina 1981 1 specimen
recombined as Paratriassoneura primitiva
PIN 2069/18 (1 measurement)
Insecta - Protomyrmeleontidae
Ferganagrion kirghiziensis n. gen., n. sp. Nel et al. 2005
Nel et al. 2005 1 specimen
PIN 2069/3266
Insecta - Diptera - Limoniidae
Mabelysia charlesi n. gen., n. sp. Shcherbakov 1995
Shcherbakov et al. 1995 1 specimen
PIN 2069/3541 (1 measurement)
Insecta - Diptera - Gnomuscidae
Gnomusca renyxa n. sp. Shcherbakov 1995
Shcherbakov et al. 1995 1 specimen
PIN 2069/3756 (1 measurement)
Insecta - Diptera - Protorhyphidae
Vymrhyphus triassicus n. sp. Blagoderov 1995
Shcherbakov et al. 1995 1 specimen
PIN 2069/3757 (1 measurement)
Insecta - Diptera - Hennigmatidae
Anemeca liya n. gen., n. sp. Shcherbakov 1995
Shcherbakov et al. 1995 1 specimen
PIN 2068/3354 (1 measurement)
see common names

Geography
Country:Kyrgyzstan State/province:Osh
Coordinates: 40.1° North, 70.2° East (view map)
Paleocoordinates:36.5° North, 60.1° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Middle Triassic
Stage:Ladinian 10 m.y. bin:Triassic 2
Key time interval:Ladinian
Age range of interval:242.00000 - 237.00000 m.y. ago
Stratigraphy
Formation:Madygen
Stratigraphic resolution:group of beds
Stratigraphy comments: Aristov (2020): The age of the Madygen locality is usually reported as the Ladinian or Carnian of the Middle or Upper Triassic (Shcherbakov, 2008). This concept emerged, because I.A. Dobruskina (1995) did not discriminate between Ladinian and Carnian floras. Our data show that only three of the six families found at Madygen are known from younger sediments (Fig. 1), and therefore we can confirm the assumption of the Ladinian age of the Madygen locality (A.G. Ponomarenko, personal communication, 2018).
Lithology and environment
Primary lithology: lithified "siliciclastic"
Environment:lacustrine - small Tectonic setting:rift
Geology comments: Oxbow lake; Madygen basin; tectonically-induced intermontane basin. Complex facies pattern consistent with active tectonics in upland region.
Taphonomy
Modes of preservation:mold/impression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Metadata
Database number:106090
Authorizer:M. Clapham Enterer:M. Clapham, L. Edwards, J. Karr, A. Delelli
Modifier:M. Clapham Research group:paleoentomology
Created:2011-03-02 09:55:17 Last modified:2020-10-10 22:02:07
Access level:the public Released:2011-03-02 09:55:17
Creative Commons license:CC BY
Reference information

Primary reference:

31339. A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [K. Beard/K. Beard/J. Karr]

Secondary references:

35387 D. S. Aristov. 2011. New and little known Blattogryllidae (Insecta: Grylloblattida) from the Permian and Triassic of Russia, Kazakhstan, and Kyrgyzstan. Paleontological Journal 45:65-72 [M. Clapham/M. Clapham]
81963 D. S. Aristov. 2022. New Geinitziidae (Insecta, Reculida) from the Permian and Triassic of Eurasia. Paleontological Journal 56:263-267 [M. Clapham/M. Clapham]
85832 D. S. Aristov and A. P. Rasnitsyn. 2023. New and little known Permosialidae (Insecta: Palaeomanteida) from the Upper Permian of Mongolia and the Middle or Upper Triassic of Kyrgyzstan. Paleontological Journal 57:422-431 [M. Clapham/M. Clapham]
39413 L. V. Arnoldi, V. V. Zherikhin, L. M. Nikritin and A. G. Ponomarenko. 1977. Mezozoiskie zhestkokryiye. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 161:1-204 [M. Clapham/M. Clapham/M. Clapham]
41079 E. E. Becker-Migdisova. 1985. Iskopaemye Nasekomye Psillomorfy. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 206:1-92 [M. Clapham/M. Clapham/M. Clapham]
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39480 A. V. Gorochov. 1987. Novye iskopaemye prjamokrylyx semejstv Bintoniellidae, Mesoedischiidae fam. n. and Pseudelcanidae fam. n. (Orthoptera, Ensifera) iz permskix i triasovyx otlozzenij SSSR. Vestnik Zoologii (1)18-23 [M. Clapham/J. Karr/M. Clapham]
37411 A. V. Gorochov. 1987. New fossil Orthopterans of the families Adumbratomrphidae fam. n, Pruvostitidae and Proparagryllacrididae (Orthoptera, Ensifera) from Perm and Triassic deposits of the USSR. Vestnik Zoologii 1987(4):20-28 [M. Clapham/J. Karr]
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8350ETE L. N. Pritykina. 1981. Noviye Triasoviye Strekozy Sredney Azii [New Triassic dragonflies from Central Asia]. Trudy Paleontologicheskogo instituta akademii nauk SSSR 183:5-42 [C. Labandeira/A. Spencer-Lee/S. Bruning]
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77262 D. E. Shcherbakov. 2021. New Curvicubitidae and Paraknightiidae (Homoptera: Cicadomorpha) from the Triassic of Central Asia. Russian Entomological Journal 30:129-134 [M. Clapham/M. Clapham]
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37000 S. Storozhenko. 1994. New Triassic grylloblattids from Kirghizia (Insecta, Grylloblattida). Spixiana 17(1):27-35 [M. Clapham/M. Clapham]
36999 S. Storozhenko. 1996. New Triassic Mesorthopteridae (Insecta, Plecopteroidea, Grylloblattida). Spixiana 19(1):115-127 [M. Clapham/M. Clapham]
37718 S. Y. Storozhenko. 1990. New Permian and Mesozoic Insects (Insecta, Grylloblattida: Blattogryllidae, Geinitziidae) from Asia. Paleontological Journal 24(4):53-61 [M. Clapham/J. Karr]
42412 S. Y. Storozhenko. 1992. New Mesozoic Grylloblattid insects (Grylloblattida) from Central Asia. Paleontological Journal 26(1):85-95 [M. Clapham/J. Karr]
37007 S. Y. Storozhenko. 1992. A new family of Triassic grylloblattids from central Asia (Insecta, Grylloblattida). Spixiana 15(1):67-73 [M. Clapham/M. Clapham/J. Karr]
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37089 V. V. Zherikhin and V. G. Gratshev. 1993. Obrieniidae, fam. nov., the oldest Mesozoic weevils (Coleoptera, Curculionoidea). Paleontological Journal 27(1A):50-68 [M. Clapham/J. Karr/M. Clapham]