Soyana (PIN collection 2455): Roadian, Russian Federation
collected by Gundersen 1930

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Insecta - Protelytroptera
Arctocoleus ivensis Martynov 1932
Martynov 1932 1 individual
PIN 2455/33 (1 measurement)
Insecta - Miomoptera - Palaeomanteidae
Archisialis praecox n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/31 (1 measurement)
Insecta - Hypoperlida - Hypoperlidae
Boreopsocus danksae Shcherbakov 1994
Shcherbakov 1994 1 individual
PIN 2455/258
Insecta - Hemiptera - Coleoscytidae
Coleoscyta rotundata Martynov 1935
Becker-Migdisova 1960 1 specimen
PIN 2455/310 (C. venosa in Martynov, 1935) (1 measurement)
Coleoscytodes venosa n. gen., n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/299 (310 is Coleoscyta rotundata) (1 measurement)
Insecta - Hemiptera - Scytinopteridae
Scytinoptera sp. Handlirsch 1904
Martynov 1935 1 individual
PIN 2455/59
Scytinoptera reducta (Martynov 1928)
Becker-Migdisova 1948 1 specimen
PIN 2455/176A
Insecta - Hemiptera - Archescytinidae
Permoscytina tetraclada n. sp. Martynov 1933
Martynov 1933 1 individual
recombined as Archescytina tetraclada
PIN 2455/45 (1 measurement)
Ivascytina difficilis n. gen., n. sp. Martynov 1933
Martynov 1933 1 individual
recombined as Sojanoscytina difficilis
PIN 2455/44 (1 measurement)
Insecta - Hemiptera - Prosbolopseidae
Scytoneura elliptica n. gen., n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/267 (1 measurement)
Ivaia indistincta n. gen., n. sp. Becker-Migdisova 1960
Becker-Migdisova 1960 2 specimens
PIN 2455/346, x (2 measurements)
Prosbolopsis simplex Martynov 1935
Martynov 1935 1 individual
synonym of Prosbolopsis ovalis
PIN 2455/304 (1 measurement)
Insecta - Hemiptera - Prosbolidae
Permocicada informal sp 2
Martynov 1935 1 individual
PIN 2455/55 (1 measurement)
Permocicada subtruncata n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/189 (1 measurement)
Permocicada nigronervosa Martynov 1928
Becker-Migdisova 1940 1 individual
PIN 2455/224 (1 measurement)
Permocicada borealis n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/60 (1 measurement)
Permocicada angustata n. sp. Martynov 1935
Martynov 1935 1 individual
recombined as Permocicadopsis angustata
PIN 2455/302 (1 measurement)
Sojanoneura bimaculata Martynov 1935
Martynov 1935 2 individuals
PIN 2455/170, 213 (2 measurements)
Sojanoneura dubiosa n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/62 (1 measurement)
Sojanoneura furcatula n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/229 (1 measurement)
Sojanoneura triangularis n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/65 (1 measurement)
Sojanoneura signata n. sp. Martynov 1935
Martynov 1935 2 individuals
PIN 2455/57, 58 (2 measurements)
Prosbole informal sp 3
Martynov 1935 2 individuals
PIN 2455/56, 53
Prosbole informal sp 4
Martynov 1935 1 individual
PIN 2455/52
Prosbole dilatata n. sp. Martynov 1935
Martynov 1935 1 individual
PIN 2455/292 (1 measurement)
Insecta - Paoliida - Anthracoptilidae
Strephoneura tenebrosa n. sp. Novokshonov and Aristov 2004
Novokshonov and Aristov 2004 2 individuals
PIN 2455/339, 340 (1 measurement)
Insecta - Mecoptera - Permocentropidae
Permocentropus philopotamoides n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/146 (1 measurement)
Insecta - Mecoptera - Permochoristidae
Petromantis sp. Handlirsch 1904
Martynov 1932 1 individual
PIN 2455/330 (1 measurement)
Petromantis rossica Handlirsch 1904
Martynova 1948 5 specimens
PIN 2455/172, 244, 245, 246, 3032
Petromantis borealis Martynov 1928
Martynov 1932 6 individuals
synonym of Petromantis rossica
PIN 2445/174, 120, 348, 167; 2445/86, 175 in Martynova, 1948 (1 measurement)
Petromantis punctata n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Petromantis rossica
PIN 2455/29 (1 measurement)
Permochorista venosa n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Petromantis maculipennis
PIN 2455/329 (1 measurement)
Permochorista venosa Martynov 1932
Martynova 1948 1 specimen
synonym of Petromantis maculipennis
PIN 2455/157
Permochorista nigromaculata n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Petromantis maculipennis
PIN 2455/332 (1 measurement)
? Permochorista stigmata n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Petromantis maculipennis
PIN 2455/185 (1 measurement)
Petrochorista distincta n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/331 (1 measurement)
Permochorista sp. Tillyard 1918
Martynov 1932 1 individual
synonym of Mesochorista
PIN 2455/268 (1 measurement)
Agetochorista ornata Martynov 1932
Martynov 1932 1 individual
recombined as Agetopanorpa ornata
PIN 2455/147
Agetochorista umbrata n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Agetopanorpa ornata
PIN 2455/43 (1 measurement)
Agetochorista splendida n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Agetopanorpa ornata
PIN 2445/343 (1 measurement)
Agetochorista reducta n. sp. Martynov 1932
Martynov 1932 1 individual
synonym of Agetopanorpa dubia
PIN 2445/10 (1 measurement)
Agetochorista sp. Martynov 1932
Martynov 1932 1 individual
synonym of Agetopanorpa
(1 measurement)
Agetochorista fasciata n. sp. Martynov 1932
Martynov 1932 1 individual
recombined as Agetopanorpa fasciata
PIN 2455/27 (1 measurement)
Oochorista gunderseni n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2445/347 (1 measurement)
Insecta - Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/87 (2 measurements)
Insecta - Permithonidae
Permithonopsis sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/117 (1 measurement)
Permithonopsis ivensis n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/37 (1 measurement)
Eopsychops sojanensis n. gen., n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/184 (1 measurement)
Permosisyra punctata n. sp. Martynov 1932
Martynov 1932 1 individual
PIN 2455/9 (1 measurement)
Insecta - Megasecoptera - Protohymenidae
Ivahymen constrictus n. gen., n. sp. Martynov 1932
1 individual
PIN 2455/5 (1 measurement)
Insecta - Palaeodictyoptera - Eukulojidae
Kuloja cubitalis n. sp. Martynov 1932
3 individuals
recombined as Eukuloja cubitalis
PIN 2455/3, 4; 2455/291 in Sinitshenkova 1981 (2 measurements)
Insecta - Permagrionidae
Permolestes gracilis n. gen., n. sp. Martynov 1932
1 individual
PIN 2455/1,2 One specimen (2 measurements)
see common names

Geography
Country:Russian Federation State/province:Arkangelsk
Coordinates: 65.6° North, 43.7° East (view map)
Paleocoordinates:33.2° North, 33.1° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Permian Epoch:Guadalupian
Stage:Roadian 10 m.y. bin:Permian 3
Key time interval:Roadian
Age range of interval:272.30000 - 268.80000 m.y. ago
Stratigraphy
Formation:Iva-Gora Beds
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology:fine lithified sandstone
Environment:lagoonal Tectonic setting:foreland basin
Geology comments: Probable brackish lagoon with some access to sea
Taphonomy
Modes of preservation:mold/impression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:survey of museum collection
Collection size:229
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Collectors:Gundersen Collection dates:1930
Metadata
Also known as:Iva-gora
Database number:111187
Authorizer:M. Clapham Enterer:J. Karr, M. Clapham
Modifier:M. Clapham Research group:paleoentomology
Created:2011-06-20 15:14:37 Last modified:2015-09-23 20:43:43
Access level:the public Released:2011-06-20 15:14:37
Creative Commons license:CC BY
Reference information

Primary reference:

36456. A. V. Martynov. 1932. New Permian Palaeoptera with the discussion of some problems of their evolution. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 1:1-44 [M. Clapham/J. Karr/M. Clapham]

Secondary references:

39941 E. E. Becker-Migdisova. 1940. Iskopaemye permskie cikady semejstva Prosbolidae s reki sojany. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 11(2):1-95 [M. Clapham/J. Karr/M. Clapham]
48456 E. E. Becker-Migdisova. 1948. Permskie tsikady semeystva Scytinopteridae s reki Soyany. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 15:1-42 [M. Clapham/M. Clapham]
37942 E. E. Becker-Migdisova. 1960. New Permian Homoptera from European USSR. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 76:1-111 [M. Clapham/J. Karr/M. Clapham]
37589 A. V. Martynov. 1932. Permian fossil Insects from the Arkhangelsk district. Part I. The order Mecoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 2:23-62 [M. Clapham/J. Karr/M. Clapham]
37594 A. V. Martynov. 1932. Permian fossil Insects from the Arkhangelsk district. Part II. Neuroptera, Megaloptera and Coleoptera, with the description of two new beetles from Tikhie Gory. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 2:63-96 [M. Clapham/J. Karr/M. Clapham]
47241 A. V. Martynov. 1933. On the Permian family Archescytinidae (Homoptera) and its relationships. Bulletin de l'Academie des sciences de l'URSS, Leningrad 1933:883-894 [M. Clapham/J. Karr]
37595 A. V. Martynov. 1935. Permian fossil Insects from the Arkhangelsk district. Part 5. Homoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 4:1-35 [M. Clapham/J. Karr/M. Clapham]
50889 O. M. Martynova. 1948. Materialy po evolyutsii Mecoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 14:1-76 [M. Clapham/M. Clapham/M. Clapham]
37700 V. G. Novokshonov and D. S. Aristov. 2004. New Taxa of Hypoperlids (Insecta: Hypoperlida) from the Upper Permian of the Arkhangelsk Region. Paleontological Journal 38(1):60-66 [M. Clapham/J. Karr]
39083 D. E. Shcherbakov. 1994. A new genus of the Paleozoic order Hypoperlida. Russian Entomological Journal 3:33-36 [M. Clapham/M. Clapham]
41068 N. D. Sinitshenkova. 1981. Svoeobraznye "dvukrylye" nasekomye iz Permi Arkhangel'skoy oblasti i Urala (Hypoperlida, Eukulojidae). Byulleten Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Geologicheskiy 56:116-126 [M. Clapham/M. Clapham/M. Clapham]