Khermeen Tsav, Red Beds (PIN coll. 3142) [SMPE]: Middle Campanian - Late/Upper Campanian, Mongolia
collected by V. Reshetov, R. Barsbold 1972, 2008
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Ostracoda
- Podocopida
- Limnocytheridae
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Limnocythere barungoiotensis
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Ostracoda
- Podocopida
- Cyprididae
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? Eucypris elata
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Cypridea cavernosa
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Cypridea profusa
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Cypridea obliquecostae
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Cypridea fracta
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"ex. gr." | ||||||||||
Cypridea rostrata
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"ex. gr." | ||||||||||
Mongolianella cuspidigera
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Reptilia
- Ankylosauridae
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Ankylosauridae indet.
Brown 1908
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Park et al. 2021 | 1 individual | ||||||||
MPC-D 100/1359 - first discovered and uncovered by the Joint Soviet-Mongolian Paleontological Expedition team in 1972 or 1973, but only collected in 2008 by members of Korea-Mongolia International Dinosaur Expedition | ||||||||||
Saichania chulsanensis
Maryanska 1977
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1 individual | |||||||||
PIN 3142/251 | ||||||||||
Reptilia
- Protoceratopsidae
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Breviceratops kozlowskii
(Maryanska and Osmólska 1975)
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Kurzanov 1990 | 4 individuals | ||||||||
PIN 3142/1, 2, 3, 5 | ||||||||||
= Bagaceratops rozhdestvenskyi
Maryanska and Osmólska 1975
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Czepiński 2019 | |||||||||
Gobiceratops minutus n. gen., n. sp.
Alifanov 2008
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Alifanov 2008 | 1 specimen | ||||||||
synonym of Bagaceratops rozhdestvenskyi | ||||||||||
PIN 3142/299 | ||||||||||
Bagaceratops rozhdestvenskyi
Maryanska and Osmólska 1975
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1 individual | |||||||||
PIN 3142/4 (1 measurement) | ||||||||||
= Breviceratops kozlowskii
Maryanska and Osmólska 1975
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Kurzanov 1990 | |||||||||
= Platyceratops tatarinovi n. gen., n. sp.
Alifanov 2003
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Alifanov 2003 | |||||||||
synonym of Bagaceratops rozhdestvenskyi | ||||||||||
Reptilia
- Prismatoolithidae
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Protoceratopsidovum sincerum
Mikhailov 1994
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Mikhailov 1994 | |||||||||
Protoceratopsidae indet.
Granger and Gregory 1923
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Mikhailov et al. 1994 | 1 specimen | ||||||||
PIN 3142/496; = P2 Group ("?Breviceratops kozlowskii"); + P1 and P3 groups; PIN 3142/415 (group P2 eggs) | ||||||||||
= Protoceratopsidovum fluxuosum n. sp.
Mikhailov 1994
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Mikhailov 1994 | |||||||||
Reptilia
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Theropoda indet.
(Marsh 1881)
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'dinosaurs' | ||||||||||
Reptilia
- Oviraptoridae
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Oviraptor sp.
Osborn 1924
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4 specimens | |||||||||
GIN 100/30-100/33 (4 measurements) | ||||||||||
= Ingenia yanshini n. gen., n. sp.
Barsbold 1981
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Barsbold 1981 | |||||||||
recombined as Heyuannia yanshini | ||||||||||
Reptilia
- Dromaeosauridae
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Velociraptor sp.
Osborn 1924
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Reptilia
- Gobioolithidae
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Gobipterygidae indet.
Elzanowski 1974
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Mikhailov et al. 1994 | 1 individual | ||||||||
eggs | ||||||||||
= Gobioolithus minor n. gen., n. sp.
Mikhailov 1996
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Mikhailov 1996 | |||||||||
PIN 3142/422 | ||||||||||
Reptilia
- Avetheropoda
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Carnosauria indet.
Huene 1920
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Reptilia
- Oblongoolithidae
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Oblongoolithus glaber n. gen., n. sp.
Mikhailov 1996
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Mikhailov 1996 | 1 specimen | ||||||||
PIN 3142/500-1 | ||||||||||
Reptilia
- Crocodylia
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Crocodylia indet.
(Owen 1842)
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Gubin 1999 | |||||||||
'crocodiles' | ||||||||||
Reptilia
- Testudines
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Testudines indet.
Batsch 1788
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'tortoises' | ||||||||||
Reptilia
- Testudines
- Lindholmemydidae
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Mongolemys barsboldi n. sp.
Sukhanov and Narmandakh 1974
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Danilov 2003 | 1 specimen | ||||||||
recombined as Gravemys barsboldi | ||||||||||
IG unnumbered | ||||||||||
Reptilia
- Polyglyphanodontidae
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Tchingisaurus multivagus n. gen., n. sp.
Alifanov 1993
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Alifanov 1993 | 1 specimen | ||||||||
PIN 3142/309 (holotype mandible) | ||||||||||
Gobinatus arenosus n. gen., n. sp.
Alifanov 1993
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Alifanov 1993 | 1 specimen | ||||||||
PIN 3142/308 (holotype skull) | ||||||||||
Reptilia
- Priscagamidae
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Chamaeleognathus iordanskyi n. gen., n. sp.
Alifanov 1996
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Alifanov 1996 | 2 specimens | ||||||||
PIN 3142/345 (holotype skull) | ||||||||||
Morunasius modestus n. gen., n. sp.
Alifanov 1996
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Alifanov 1996 | 1 specimen | ||||||||
PIN 3142/317 (holotype fragmentary skull and vertebrae) | ||||||||||
Phrynosomimus asper n. gen., n. sp.
Alifanov 1996
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Alifanov 1996 | 1 specimen | ||||||||
PIN 3142/318 (holotype skull) | ||||||||||
Reptilia
- Hoplocercidae
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Gladidenagama semiplena n. gen., n. sp.
Alifanov 1996
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Alifanov 1996 | 1 specimen | ||||||||
PIN 3142/319 (holotype skull) | ||||||||||
Reptilia
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Prodenteia ministra n. gen., n. sp.
Alifanov 1993
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Alifanov 1993 | 1 specimen | ||||||||
PIN 3142/324 (holotype partial skull) | ||||||||||
Mongolochamops facilis n. sp.
Alifanov 1988
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Alifanov 1988 | 1 specimen | ||||||||
recombined as Altanteius facilis | ||||||||||
PIN 3142/306 (holotype partial snout) | ||||||||||
Parameiva oculea n. gen., n. sp.
Alifanov 1993
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Alifanov 1993 | 1 specimen | ||||||||
PIN 3142/310 (holotype partial skull) | ||||||||||
Mammalia
- Zalambdalestidae
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Barunlestes butleri
Kielan-Jaworowska 1975
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Kielan-Jaworowska and Trofimov 1980 | 1 specimen | ||||||||
PIN 3142/710 (skull) | ||||||||||
Mammalia
- Asioryctidae
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Asioryctes nemegtensis
Kielan-Jaworowska 1975
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Mammalia
- Deltatheroida
- Deltatheridiidae
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Deltatheridium pretrituberculare
Gregory and Simpson 1926
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Mammalia
- Multituberculata
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Nemegtbaatar gobiensis
Kielan-Jaworowska 1974
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Chulsanbaatar vulgaris
Kielan-Jaworowska 1974
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Amphibia
- Temnospondyli
- Gobiatidae
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Gobiates leptocolaptus
(Borsuk-Bialynicka 1978)
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Gubin 1999 | 1 individual | ||||||||
Gobiates khermeentsavi n. gen., n. sp.
Spinar and Tatarinov 1986
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Spinar and Tatarinov 1986 | 3 individuals | ||||||||
three partial skulls (PIN 3142/1, 2, and 3) | ||||||||||
see common names |
Geography
Country: | Mongolia | State/province: | Omnogov |
Coordinates: | 43.3° North, 99.6° East (view map) | ||
Paleocoordinates: | 40.3° North, 89.2° East | ||
Basis of coordinate: | estimated from map | ||
Geographic resolution: | outcrop |
Time
Period: | Cretaceous | Epoch: | Late/Upper Cretaceous |
Stage: | Campanian | 10 m.y. bin: | Cretaceous 7 |
*Period: | Late/Upper Cretaceous - Late/Upper Cretaceous | *Epoch: | Senonian |
*International age/stage: | Campanian - Maastrichtian | ||
Key time interval: | Middle Campanian - Late/Upper Campanian | ||
Age range of interval: | 83.50000 - 70.60000 m.y. ago | ||
* legacy (obsolete) database fields |
Stratigraphy
Formation: | Baruungoyot | ||||
Stratigraphic resolution: | group of beds | ||||
Stratigraphy comments: Red Beds of Khermin Tsav |
Lithology and environment
Primary lithology: | not reported |
Environment: | terrestrial indet. |
Taphonomy
Modes of preservation: | body |
Size of fossils: | macrofossils |
Preservation of anatomical detail: | excellent |
Collection methods and comments
Collection excludes: | some macrofossils | ||
Collection methods: | surface (in situ),field collection | ||
Reason for describing collection: | taxonomic analysis | ||
Collectors: | V. Reshetov, R. Barsbold | Collection dates: | 1972, 2008 |
Collection method comments: A broad collection that might include the collection #1157, but necessary because of nomenclatural variation from authors to authors, and also because the stratigraphic position of Gobiates khermeentsavi in the locality section is unknown. Distinct from PMPE and other collections from the same general site. Collected via Joint Soviet-Mongolian Paleontological Expedition |
Metadata
Also known as: | Khermin Tsav; Khermin-Tsab; Hermiin Tsav, Soviet-Mongolian Paleontological Expedition | ||
Database number: | 11582 | ||
Authorizer: | E. Fara, M. Carrano, P. Mannion, R. Benson | Enterer: | E. Fara, M. Carrano, R. Benson, P. Mannion |
Modifier: | M. Carrano | Research group: | vertebrate |
Collections that are a subset of this one: | |||
Created: | 2001-08-22 11:26:41 | Last modified: | 2023-11-03 14:50:03 |
Access level: | group members | Released: | 2001-11-22 11:26:41 |
Creative Commons license: | CC BY |
Reference information
Primary reference:
59136. | R. Gradzinski, Z. Kielan-Jaworowska, and T. Maryanska. 1977. Upper Cretaceous Djadokhta, Barun Goyot and Nemegt formations of Mongolia, including remarks on previous subdivisions. Acta Geologica Polonica 27(3):281-318 [M. Carrano/M. Carrano] |
Secondary references:
37919 | V. R. Alifanov. 1988. The new lizards (Lacertilia: Teiidae) from the Upper Cretaceous of Mongolia. In F. Kurochkin (ed.), Fossil reptiles and birds of Mongolia 90-100 [R. Benson/R. Benson] | |
37931 | V. R. Alifanov. 1989. [New priscagamas (Lacertilia) from the Upper Cretaceous of Mongolia and their systematic position in the Iguania]. Paleontologiceskij Zhurnal 4:73-87 [R. Benson/R. Benson] | |
38090 | V. R. Alifanov. 1993. New lizards of the family Macrocephalosauridae (Sauria) from the Upper Cretaceous of Mongolia, critical remarks on the systematics of the Teiidae. Paleontological Journal 27(1):70-90 [R. Benson/R. Benson] | |
38117 | V. R. Alifanov. 1996. Lizards of the families Priscagamidae and Hoplocercidae (Sauria, Iguania): phylogenetic position and new representatives from the Late Cretaceous of Mongolia. Paleontological Journal 30(4):466-483 [R. Benson/R. Benson] | |
38112 | V. R. Alifanov. 2000. The fossil record of Cretaceous lizards from Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 368-389 [R. Benson/R. Benson/M. Carrano] | |
11744 | ETE | V. R. Alifanov. 2003. Two new dinosaurs of the infraorder Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People's Republic. Paleontological Journal 37(5):524-534 [M. Carrano/M. Carrano/M. Carrano] |
29118 | ETE | V. R. Alifanov. 2008. The tiny horned dinosaur Gobiceratops minutus gen. et sp. nov. (Bagaceratopidae, Neoceratopsia) from the Upper Cretaceous of Mongolia. Paleontological Journal 42(6):621-633 [M. Carrano/M. Carrano/R. Butler] |
71135 | V. M. Arbour and P. J. Currie. 2011. Tail and pelvis pathologies of ankylosaurian dinosaurs. Historical Biology 23(4):375-390 [M. Carrano/M. Carrano] | |
66054 | V. M. Arbour, P. J. Currie, and D. Badamgarav. 2014. The ankylosaurid dinosaurs of the Upper Cretaceous Baruungoyot and Nemegt formations of Mongolia. Zoological Journal of the Linnean Society 172(3):631-652 [G. Lloyd/G. Lloyd/M. Carrano] | |
64040 | V. M. Arbour and P. J. Currie. 2016. Systematics, phylogeny and palaeobiogeography of the ankylosaurid dinosaurs. Journal of Systematic Palaeontology 14(5):385-444 [G. Lloyd/G. Lloyd/M. Carrano] | |
29489 | ETE | R. Barsbold. 1981. Bezzubyye khishchnyye dinozavry Mongolii [Toothless carnivorous dinosaurs of Mongolia]. Sovmestnaia Sovetsko-Mongol’skaia Paleontologicheskaia Ekspeditsiia Trudy 15:28-39 [M. Carrano/M. Carrano] |
15004 | ETE | R. Barsbold. 1983. Khishchnye dinosavry mela Mongoliy [Carnivorous dinosaurs from the Cretaceous of Mongolia]. Transactions of the Joint Soviet-Mongolian Paleontological Expedition 19:1-117 [M. Carrano/M. Carrano] |
86269 | Ł Czepiński. 2019. Ontogeny and variation of a protoceratopsid dinosaur Bagaceratops rozhdestvenskyi from the Late Cretaceous of the Gobi Desert. Historical Biology 32(10):1394-1421 [M. Carrano/M. Carrano] | |
39832 | I. Danilov. 2003. Gravemys Sukhanov and Narmandakh, 1983 (Testudinoidea: Lindholmemydidae) from the Late Cretaceous of Asia: new data. PaleoBios 23(3):9-19 [M. Carrano/M. Carrano] | |
4173 | 5% 1280 | Y. M. Gubin. 1999. Gobiatidy (Anura) iz verkhnemelovogo mestonakhozhdeniya Khermin-Tsav; Gobi, Mongoliya (= Gobiatids (Anura) from the Upper Cretaceous locality Khermeen-Tsav (Gobi Desert, Mongolia)). Paleontologicheskii Zhurnal 1:76-87 [E. Fara/E. Fara/J. Alroy] |
39263 | Z. Kielan-Jaworowska and B. A. Trofimov. 1980. Cranial morphology of the Cretaceous eutherian mammal Barunlestes. Acta Palaeontologica Polonica 25(2):167-185 [R. Benson/R. Benson] | |
69233 | E. N. Kurochkin and R. Barsbold. 2000. The Russian-Mongolian expeditions and research in vertebrate palaeontology. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 235-255 [M. Carrano/M. Carrano] | |
86270 | S. M. Kurzanov. 1990. A new Late Cretaceous protoceratopsid genus from Mongolia. Paleontological Journal 1990(4):85-91 [M. Carrano/M. Carrano] | |
14217 | ETE | K. Mikhailov, K. Sabath, and S. Kurzanov. 1994. Eggs and nests from the Cretaceous of Mongolia. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge 88-115 [M. Carrano/M. Carrano/M. Carrano] |
44374 | K. E. Mikhailov. 1994. Theropod and protoceratopsian dinosaur eggs from the Cretaceous of Mongolia and Kazakhstan. Paleontological Journal 28:101-120 [J. Alroy/A. Garcia Selles/M. Carrano] | |
69240 | K. E. Mikhailov. 1996. Bird eggs in the Upper Cretaceous of Mongolia. Paleontological Journal 30(1):114-116 [M. Carrano/M. Carrano/M. Carrano] | |
69241 | K. E. Mikhailov. 1996. New genera of fossil eggs from the Upper Cretaceous of Mongolia. Paleontological Journal 30(2):246-248 [M. Carrano/M. Carrano] | |
18353 | ETE | K. E. Mikhailov. 2000. Eggs and eggshells of dinosaurs and birds from the Cretaceous of Mongolia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 560-572 [M. Carrano/M. Carrano/M. Carrano] |
75923 | J.-Y. Park, Y.-N. Lee, P. J. Currie, M. J. Ryan, P. Bell, R. Sissons, E. B. Koppelhus, R. Barsbold, S. Lee and S.-H. Kim. 2021. A new ankylosaurid skeleton from the Upper Cretaceous Baruungoyot Formation of Mongolia: its implications for ankylosaurid postcranial evolution. Scientific Reports 11:4101:1-10 [P. Mannion/P. Mannion] | |
82395 | J.-Y. Park, Y.-N. Lee, Y. Kobayashi, L. L. Jacobs, R. Barsbold, H.-J. Lee, N. Kim, K.-Y. Song, and M. J. Polcyn. 2021. A new ankylosaurid from the Upper Cretaceous Nemegt Formation of Mongolia and implications for paleoecology of armoured dinosaurs. Scientific Reports 11:22928 [P. Mannion/P. Mannion] | |
69215 | M. A. Shishkin. 2000. Mesozoic amphibians from Mongolia and the Central Asiatic republics. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 297-308 [M. Carrano/M. Carrano] | |
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30994 | Z. V. Spinar and L. P. Tatarinov. 1986. A new genus and species of discoglossid frog from the Upper Cretaceous of the Gobi Desert. Journal of Vertebrate Paleontology 6(2):113-122 [J. Alroy/J. Alroy] | |
41695 | V. B. Sukhanov. 2000. Mesozoic turtles of Middle and Central Asia. In M. J. Benton, M. A. Shishkin, D. M. Unwin, & E N. Kurichkin (eds.), The Age of Dinosaurs in Russia and Mongolia 309-367 [R. Benson/R. Benson/M. Carrano] | |
52600 | T. A. Tumanova. 1977. Novyye dannyye ob ankilozavre Tarchia gigantea [New data on the ankylosaur Tarchia gigantea]. Paleontologicheskii Zhurnal 1977(4):92-100 [M. Carrano/M. Carrano/M. Carrano] | |
52601 | T. A. Tumanova. 1978. New data on the ankylosaur Tarchia gigantea. Paleontological Journal 1977(4):480-486 [M. Carrano/M. Carrano/M. Carrano] | |
26796 | ETE | T. A. Tumanova. 1987. Pantsirnyye dinozavry Mongolii [The armored dinosaurs of Mongolia]. Trudy Sovmestnaya Sovetsko-Mongol'skaya Paleontologicheskaya Ekspeditsiya 32:1-80 [M. Carrano/M. Carrano/M. Carrano] |
3793 | R. J. Zakrzewski. 1985. A New Species of Woodrat (Cricetidae) from the Pleistocene (Irvingtonian) of South Dakota. Journal of Mammalogy 66(4):770-773 [J. Alroy/J. Alroy/J. Ju] |