Oeningen (Zurich collection): Sarmatian, Germany

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Arachnida - Araneae - Gnaphosidae
Macaria tenella n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Micaria tenella
Arachnida - Araneae - Theridiidae
Theridion maculipes n. sp. Heer 1864
Heer 1864 1 specimen
Theridion globulus n. sp. Heer 1864
Heer 1864 1 specimen
Theridion annulipes n. sp. Heer 1864
Heer 1864 1 specimen
Arachnida - Araneae - Araneidae
Epeira molassica n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Araneus molassicus
Arachnida - Araneae - Dictynidae
? Argyroneta longipes n. sp. Heer 1864
Heer 1864 1 specimen
Arachnida - Araneae - Thomisidae
Thomisus sulzeri n. sp. Heer 1864
Heer 1864 1 specimen
Thomisus lividus n. sp. Heer 1864
Heer 1864 1 specimen
Thomisus oeningensis n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Xysticus oeningensis
Arachnida - Araneae
Schellenbergia rotundata n. gen., n. sp. Heer 1864
Heer 1864 1 specimen
Clubiona eseri n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Araneae
Malacostraca - Isopoda - Armadillidiidae
Armadillo molassicus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Armadillidium molassicum
Insecta - Odonata
"Libellula" thetis Heer 1849
1 specimen
nomen dubium belonging to Anisoptera
larva
Insecta - Odonata - Sieblosiidae
Agrion iris n. sp. Heer 1879
Heer 1879 1 specimen
recombined as Stenolestes iris
Insecta - Odonata
Agrion (Sterope) parthenope n. sp. Heer 1849
1 specimen
recombined as Steropoides parthenope
(2 measurements)
Insecta - Ephemeroptera
Ephemera oeningensis n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Ephemeroptera
Insecta - Orthoptera - Pyrgomorphidae
Oedipoda fischeri n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Miopyrgomorpha fischeri
(1 measurement)
Insecta - Orthoptera - Acrididae
Oedipoda germari n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Acrydiana
Insecta - Orthoptera - Tetrigidae
Tetrix gracilis n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Tettigidea gracilis
Insecta - Orthoptera - Tettigoniidae
Decticus speciosus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Platycleis speciosa
Insecta - Orthoptera - Gryllidae
Gryllus troglodytes n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Grylloides
Insecta - Orthoptera - Gryllotalpidae
Gryllotalpa stricta n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Gryllotalpidae
Insecta - Dermaptera - Forficulidae
Forficula recta n. sp. Heer 1864
Heer 1864 1 specimen
Forficula minuta n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Blattodea - Rhinotermitidae
Termes hartungi n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Reticulitermes hartungi
Insecta - Blattodea - Termitidae
Termes buchii n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Termitinae
Insecta - Blattodea - Blattidae
Blattidium coloratum n. sp. Heer 1864
Heer 1864 2 specimens
recombined as Blatta colorata
from Kesselstein
Insecta - Coleoptera
Hydrobius godeti n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Coleoptera
from Libellenschicht (upper quarry)
Cistelites spectabilis n. sp. Heer 1869
Heer 1869 1 specimen
Insecta - Coleoptera - Gyrinidae
Dineutus longiventris n. sp. Heer 1862
Heer 1862 1 specimen
recombined as Dineutus (Cyclinus) longiventris
from Kesselstein
Miodineutes heeri n. sp. Hatch 1927
Hatch 1927 1 specimen
Dineutus longiventris Specimen in pl 2, fig 25 of Heer
Dineutus insignis n. sp. Heer 1862
Heer 1862 1 specimen
recombined as Miodineutes insignis
from Kesselstein
Miodineutes oeningensis n. gen., n. sp. Hatch 1927
Hatch 1927 1 specimen
Dineutus insignis specimen in pl 2, fig 28 of Heer
Insecta - Coleoptera - Dytiscidae
Dytiscus lavateri Heer 1847
Heer 1862 4 specimens
from Kesselstein
Hydaticus areolatus n. sp. Heer 1862
Heer 1862 1 specimen
replaced by Graphoderus heeri
Cybister nicoleti Heer 1862
Heer 1862 1 specimen
recombined as Megadytes nicoleti
from Kesselstein
Cybister agassizi n. sp. Heer 1862
Heer 1862 10 specimens
from Kesselstein
Cybister atavus n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
Hydroporus benzeli n. sp. Heer 1862
Heer 1862 1 specimen
recombined as Nebrioporus benzeli
Colymbetes aemulus n. sp. Heer 1862
Heer 1862 4 specimens
from Kesselstein
Insecta - Coleoptera - Carabidae
Dichirotrichus lividus Heer 1862
Heer 1862 1 specimen
from Kesselstein
Harpalus constrictus n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Harpalus stierlini n. sp. Heer 1862
Heer 1862 1 specimen
Harpalus stygius n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Harpalus tardigradus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Sinis brevicollis n. gen., n. sp. Heer 1862
Heer 1862 2 specimens
1 male, 1 female from lower quarry
Amara pinguicula n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Amara primigenia n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Amara princeps n. sp. Heer 1862
Heer 1862 2 specimens
1 from Insektenschicht (lower quarry), 1 from Kesselstein
Pterostichus minutulus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Pterostichus vetustus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Badister macrocephalus n. sp. Heer 1862
Heer 1862
"multiple examples" in Kesselstein, including var. major
Badister fragilis n. sp. Heer 1862
Heer 1862 3 specimens
from Kesselstein (2 var. paulo major)
Badister grandis n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
Nebria pluto n. sp. Heer 1862
Heer 1862 3 specimens
1 from lower quarry, 2 from Kesselstein
Calosoma nauckianum n. sp. Heer 1860
Heer 1860 1 specimen
Calosoma escheri n. sp. Heer 1860
Heer 1860 2 specimens
Calosoma escrobiculatum n. sp. Heer 1860
Heer 1860 2 specimens
Calosoma catenulatum n. sp. Heer 1860
Heer 1860 1 specimen
Kesselstein
Calosoma deplanatum n. sp. Heer 1860
Heer 1860 2 specimens
recombined as Campalita deplanatum
Glenopterus laevigatus n. gen., n. sp. Heer 1847
Heer 1847 1 specimen
(2 measurements)
Carabites laesicollis n. sp. Heer 1862
Heer 1862 1 specimen
Trechinites oblongus n. sp. Heer 1862
Heer 1862 1 specimen
from lower quarry
Trechinites clairvillii n. sp. Heer 1862
Heer 1862 1 specimen
from lower quarry
Insecta - Coleoptera - Buprestidae
Protogenia escheri n. gen., n. sp. Heer 1847
Heer 1847 1 specimen
(1 measurement)
Ancylocheira concinna n. sp. Heer 1879
Heer 1879 1 specimen
recombined as Buprestis concinna
Ancylocheira alemanica n. sp. Heer 1879
Heer 1879 1 specimen
recombined as Buprestis alemanica
Ancylochira tincta n. sp. Heer 1862
Heer 1862 2 specimens
recombined as Buprestis tincta
from Kesselstein,includes var. b. elytris majoribus
? Anthaxia doris n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Anthaxia pallida n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Buprestites agriloides n. sp. Heer 1862
Heer 1862 1 specimen
Capnodis puncticollis Heer 1847
Heer 1862 2 specimens
Capnodis antiqua Heer 1847
Heer 1862
"several" specimens
Capnodis spectabilis n. sp. Heer 1862
Heer 1862 3 specimens
from Kesselstein
Chalcophora laevigata Heer 1862
Heer 1862 8 specimens
Chalcophora pulchella n. sp. Heer 1862
Heer 1862 3 specimens
from Kesselstein
Acmaeodera antholitha n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Acmaeodera brevicollis n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Insecta - Coleoptera - Elateridae
Elater (Alaus) spectabilis n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Alaus spectabilis
Insecta - Coleoptera - Lampyridae
Lampyris orciluca n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Lampyris
Insecta - Coleoptera - Cantharidae
Telephorus macilentus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Cantharis (Cantharis) macilenta
Insecta - Coleoptera - Byrrhidae
Byrrhus oeningensis Heer 1847
Heer 1862
several from Kesselstein
Insecta - Coleoptera - Lymexylidae
Hylecoetus cylindricus n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Hylecoetus
Insecta - Coleoptera - Tenebrionidae
Tagenopsis brevicornis n. gen., n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Coleoptera - Meloidae
Zonites vetusta n. sp. Heer 1864
Heer 1864 1 specimen
Lytta aesculapii Heer 1847
Heer 1847 1 specimen
Insecta - Coleoptera - Trogossitidae
Gymnochila obesa n. sp. Heer 1862
Heer 1862 1 specimen
recombined as Gymnocheilis obesa
from Kesselstein
Trogosita amissa n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein and Insektenschicht (lower quarry)
Trogosita sculpturata n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Trogosita assimilis n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Trogosita koellikeri (Heer 1847)
Heer 1862 1 specimen
Trogosita longicollis n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Trogosita bella n. sp. Heer 1862
Heer 1862 1 specimen
Insecta - Coleoptera - Attelabidae
Attelabus durus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Phialodes durus
Insecta - Coleoptera - Curculionidae
Lixus oeningensis n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Curculionites
Naupactus crassirostris n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Entiminae
Sitona atavina n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Entiminae
Brachycerus nanus n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Curculionites
Cleonus speciosus n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Cleonini
Insecta - Coleoptera - Brentidae
Miocenocylas heeri n. gen., n. sp. Legalov 2018
Legalov 2018 1 specimen
Apion antiquum n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Apionidae
Antliarhinites gracilis n. gen., n. sp. Heer 1864
Heer 1864 1 specimen
Rhynchites dionysus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Omphalapion dionysus
Insecta - Coleoptera - Cerambycidae
Lamia antiqua n. sp. Heer 1879
Heer 1879 1 specimen
Acanthoderus sepultus n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Acanthoderes
Acanthoderus lepidus n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Acanthoderes
Callidium procerum n. sp. Heer 1864
Heer 1864 1 specimen
Callidium escheri n. sp. Heer 1864
Heer 1864 1 specimen
Clytus pulcher n. sp. Heer 1864
Heer 1864 1 specimen
Prionus spectabilis n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Ergates spectabilis
Prionus polyphemus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Ergates polyphemus
Insecta - Coleoptera - Chrysomelidae
Lema vetusta n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Crioceris vetusta
Gonioctena japeti n. sp. Heer 1847
Heer 1847 1 specimen
(2 measurements)
Galeruca gemmifera n. sp. Heer 1879
Heer 1879 1 specimen
Galeruca buchi n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Coleoptera - Coccinellidae
Coccinella colorata n. sp. Heer 1864
Heer 1864 1 specimen
Coccinella decempustulata n. sp. Heer 1879
Heer 1879 1 specimen
replaced by Coccinella heeri
Written "Coccinella 10-pustulata"
Coccinella spectabilis n. sp. Heer 1864
Heer 1864 1 specimen
replaced by Coccinella ponomarenkoi
Coccinella amabilis n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Coleoptera - Nitidulidae
Nitidula maculigera n. sp. Heer 1862
Heer 1862 1 specimen
Nitidula ancora n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Nitidula pallida n. sp. Heer 1862
Heer 1862 1 specimen
Insecta - Coleoptera - Histeridae
Hister marmoratus n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Hister antiquus n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Hister aemulus n. sp. Heer 1862
Heer 1862
several from Kesselstein
Hister mastodontis n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Hister maculigerus n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
Hister morosus n. sp. Heer 1862
Heer 1862 3 specimens
from Insektenschicht (lower quarry)
Hister coprolithorum n. sp. Heer 1862
Heer 1862
numerous specimens from Kesselstein
Hister vetustus n. sp. Heer 1862
Heer 1862
several from Kesselstein
Insecta - Coleoptera - Hydrophilidae
Hydrobius couloni n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Hydrous ovalis n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Hydrophilini
from Kesselstein
Hydrous rehmanni (Heer 1847)
Heer 1862 9 specimens
synonym of Hydrochara noachica
Hydrophilus spectabilis Heer 1847
Heer 1847 1 specimen
Hydrophilus giganteus n. sp. Heer 1862
Heer 1862 11 specimens
synonym of Hydrophilus spectabilis
from Kesselstein; ETHZ NO-Z51.3-39-4:4-Pl.I143b
Hydrobiomorpha heeri n. sp. Fikáček and Schmied 2013
Fikáček and Schmied 2013 1 specimen
ETHZ NO-Z51.3-39-4:4-Pl. I 149b (label reads Hydrophilus ovalis)
Hydrous braunii (Heer 1847)
Heer 1862 4 specimens
recombined as Hydrobiomorpha braunii
from Kesselstein and Insektenschicht (lower quarry)
Hydrous escheri n. sp. Heer 1862
Heer 1862 1 specimen
synonym of Hydrobiomorpha braunii
from Insektenschicht (lower quarry); ETHZ NO-Z51.3-39-4:4-Pl.I148
Hydrophilopsis elongata n. gen., n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Hydrophilini
from Insektenschicht (lower quarry)
Escheria bella n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Hydrophilidae
Insecta - Coleoptera - Staphylinidae
Silpha tricostata (Heer 1847)
Heer 1862 1 specimen
original and current combination Peltis tricostata
from Kesselstein
Staphylinus atavus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Scaphisoma gracile n. sp. Heer 1862
Heer 1862 3 specimens
2 from Kesselstein, 1 from from Insektenschicht (lower quarry)
Scaphidium deletum Heer 1847
Heer 1862 1 specimen
Lathrobium oeningense n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Protactus minor n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Bledius speciosus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Oxytelus proaevus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
? Oxyporus seuberti n. sp. Heer 1862
Heer 1862 1 specimen
Insecta - Coleoptera - Glaphyridae
Glaphyrus antiquus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Insecta - Coleoptera
Copris subterranea n. sp. Heer 1862
Heer 1862 2 specimens
nomen dubium belonging to Scarabaeoidea
from Kesselstein
Insecta - Coleoptera - Hybosoridae
Hybosorus lividus n. sp. Heer 1862
Heer 1862 1 specimen
Insecta - Coleoptera - Scarabaeidae
Scarabaeus proserpinae n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Dynastinae
from Kesselstien
Onthophagus bisontinus n. sp. Heer 1862
Heer 1862 1 specimen
from Insektenschicht (lower quarry)
Onthophagus prodromus n. sp. Heer 1862
Heer 1862
several from Kesselstein
Onthophagus crassus n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
Gymnopleurus rotundatus n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
Gymnopleurus sisyphus n. sp. Heer 1847
Heer 1847 1 specimen
(1 measurement)
Copris druidum n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Anomala fugax n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Lepitrix germanica n. sp. Heer 1862
Heer 1862 1 specimen
Serica minutula n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
Oniticellus amplicollis n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Scarabaeides
from Kesselstein
Aphodius brevipennis n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Trichius aedilis n. sp. Heer 1862
Heer 1862 1 specimen
recombined as Gnorimus aedilis
from Kesselstein
Trichius lugubris n. sp. Heer 1862
Heer 1862 1 specimen
Trichius unifasciatus n. sp. Heer 1862
Heer 1862 2 specimens
from Kesselstein
? Trichius rotundatus n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Valgus oeningensis n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Insecta - Coleoptera
Gymnopleurus deperditus n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Scarabaeoidea
from Kesselstein
Onthophagus urus Heer 1847
Heer 1862
nomen dubium belonging to Scarabaeoidea
"several" specimens
Insecta - Coleoptera - Geotrupidae
Geotrupes germari n. sp. Heer 1862
Heer 1862 1 specimen
from Kesselstein
Insecta - Coleoptera
Hydrophilus stenopterus n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Coleoptera
from Kesselstein
Helophorus magnus n. sp. Heer 1862
Heer 1862 1 specimen
nomen dubium belonging to Coleoptera
from Kesselstein
Insecta - Diptera - Chironomidae
Chironomus gaudini n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Diptera - Tabanidae
? Hexatoma oeningensis n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Heptatoma oeningensis
Insecta - Diptera - Syrphidae
Syrphus bremii n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Syrphiae
Syrphus schellenbergi n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Syrphiae
Insecta - Diptera - Oestridae
Dipterites obovatus n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Adipterites obovatus
Insecta - Diptera - Bibionidae
Dilophus pumilio n. sp. Skartveit and Pika 2014
Skartveit and Pika 2014 1 specimen
ETHZ Pl. I 886 (2 measurements)
Bibio crassinervis n. sp. Skartveit and Pika 2014
Skartveit and Pika 2014 6 specimens
ETHZ Pl. I 851, Pl.I 851(2), Pl. I 862, Pl. I 875, Pl. I 888
Bibio moestus Heer 1849
Skartveit and Pika 2014 3 specimens
ETHZ Pl. I 855, 857(x2)
Bibio brevis Heer 1849
Skartveit and Pika 2014 11 specimens
Some in lot Pl. I 856; ETHZ Pl.I 865, Pl. I 887
Bibio pulchellus Heer 1849
Skartveit and Pika 2014 3 specimens
ETHZ Pl. I 854
Bibio fusiformis Heer 1849
Skartveit and Pika 2014 7 specimens
ETHZ Pl. I 858 (x2), 861 (x6)
Bibio angustatus Heer 1849
Skartveit and Pika 2014 5 specimens
ETHZ Pl. I 851
Plecia macilenta n. sp. Skartveit and Pika 2014
Skartveit and Pika 2014 2 specimens
ETHZ Pl. I 853, 853 female (1 measurement)
Protomyia jucunda Heer 1849
7 specimens
recombined as Plecia jucunda
ETHZ Pl. I 871, 853(x2), 856, 861, 867, 869 (some labeled by Brongniart fide Skartveit & Pika 2014)
Plecia hilaris Heer 1849
Skartveit and Pika 2014 1 specimen
ETHZ Pl. I 850
Bibio elongatus Heer 1849
Skartveit and Pika 2014 1 specimen
nomen dubium belonging to Bibionidae
ETHZ S.N. 189 Pl. I 863 (labeled by Heer)
Protomyia speciosa n. sp. Heer 1864
Heer 1864 3 specimens
synonym of Penthetria amoena
ETHZ Pl. I 873; also Pl. I 872, 876 as P. amoena in Skartveit & Pika, 2014
Penthetria brevicollis (Heer 1849)
Skartveit and Pika 2014 1 specimen
ETHZ Pl. I 870
Insecta - Diptera - Mycetophilidae
Mycetophila pusillima n. sp. Heer 1864
Heer 1864 1 specimen
Mycetophila orci n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Diptera - Cecidomyiidae
Cecidomyia bremii n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Cecidomyiinae
Insecta - Diptera - Sciaridae
Sciara deleta n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Lepidoptera
Bombycites buchii n. sp. Heer 1864
Heer 1864 1 specimen
"Bombycites" oeningensis n. sp. Heer 1849
1 specimen
pupa
Insecta - Hymenoptera - Sphecidae
Ammophila annosa n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Ammophila
Ammophila inferna n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Hymenoptera - Apidae
Bombus atavus n. sp. Heer 1867
Heer 1867 1 specimen
recombined as Xylocopa atavus
Bombus jurinei n. sp. Heer 1864
Heer 1867 2 specimens
recombined as Xylocopa jurinei
ETH Pl. I, 682a (type in Heer, 1865), 682b (specimen in heer, 1867) (4 measurements)
Xylocopa senilis Heer 1849
Zeuner and Manning 1976 2 specimens
ETH 21a(b), doutbfully 21c (2 measurements)
Bombus grandaevus Heer 1849
Heer 1867 1 specimen
nomen dubium belonging to Apiariae
Bombus abavus n. sp. Heer 1867
Zeuner and Manning 1976 3 specimens
Holotype (lost), ETH Pl. I, 808 (examined by Cockerell), SN.201 (2 measurements)
Anthophorites longaeva Heer 1867
Zeuner and Manning 1976 3 specimens
ETH Pl. I, 683, 712, 809 (not Heer's types) (3 measurements)
Anthophorites mellona n. gen., n. sp. Heer 1849
1 specimen
(1 measurement)
Anthophorites veterana Heer 1849
Zeuner and Manning 1976 1 specimen
ETH Pl. I, 684
Insecta - Hymenoptera
Andrena primaeva n. sp. Cockerell 1909
Cockerell 1909 1 specimen
SN.202
Insecta - Hymenoptera - Megachilidae
Megachilidae indet. Latreille 1802
Zeuner and Manning 1976 4 specimens
ETH Pl. I, 687; Pl. I, 688(689); Pl. I, 718
Apis adamitica n. sp. Heer 1864
Heer 1864 2 specimens
recombined as Lithurge adamitica
ETH V.S. 4 (Heer's holotype); SN.200 (apparently different from Heer specimens) in Zeuner & Manning, 1976
Osmia nigra n. sp. Zeuner and Manning 1976
Zeuner and Manning 1976 2 specimens
ETH Pl. I, 719; Pl. I, 685 (1 measurement)
Insecta - Hymenoptera - Formicidae
Ponera veneraria n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Poneropsis veneraria
Poneropsis stygia n. sp. Heer 1867
Heer 1867 4 specimens
(1 measurement)
Ponera globosa n. sp. Heer 1849
1 specimen
recombined as Poneropsis globosa
(1 measurement)
Imhoffia pallida n. sp. Heer 1864
Heer 1867 1 specimen
Formica orbata Heer 1849
1 specimen
Formica lignitum Germar 1837
3 specimens
recombined as Camponotus lignitus
Formica heraclea Heer 1849
1 specimen
recombined as Camponotus heracleus
Insecta - Hymenoptera - Vespidae
Polistes kirbyanus n. sp. Cockerell 1915
Cockerell 1915 1 specimen
ETH Pl. I, 681; Osmia kirbyana in Heer's label (unpublished) (2 measurements)
Polistes primitiva n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Polistes
Insecta - Hymenoptera - Scoliidae
Scolia saussureana n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Hymenoptera - Ichneumonidae
Ichneumonites bellus n. sp. Heer 1867
Heer 1867 2 specimens
Ichneumon infernalis n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Hymenoptera
Pteromalites oeningensis n. gen., n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Chalcidoidea
Insecta - Hemiptera - Cercopidae
Cercopis herrichi n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Cercopis
Cercopis hageni n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Cercopis
Cercopis germari n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Hemiptera - Gerridae
Limnobates prodromus n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Halobatinae
Insecta - Hemiptera - Belostomatidae
Belostoma speciosum n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Belostomatidae
Insecta - Hemiptera - Corixidae
Corisa fasciolata Heer 1853
Heer 1853 1 specimen
nomen dubium belonging to Corixidae
from the Libellen-schicht
Insecta - Hemiptera - Reduviidae
Harpactor maculipes n. sp. Heer 1853
Heer 1853 1 specimen
lower quarry
Harpactor obsoletus n. sp. Heer 1853
Heer 1853 1 specimen
from the Kesselstein
Insecta - Hemiptera - Nabidae
Prostemma oeningensis n. sp. Heer 1853
Heer 1853 1 specimen
lower quarry
Insecta - Hemiptera - Tingidae
Monanthia wollastoni n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Dictyla wollastoni
Insecta - Hemiptera - Alydidae
Nabis gracillima n. sp. Heer 1864
Heer 1864 1 specimen
recombined as Licremytra gracillima
Alydus pulchellus n. sp. Heer 1853
Heer 1853 1 specimen
nomen dubium belonging to Alydidae
lower quarry
Insecta - Hemiptera - Scutelleridae
Pachycoris escheri Heer 1853
Heer 1853 1 specimen
Pachycoris burmeisteri n. sp. Heer 1864
Heer 1864 1 specimen
Pachycoris protogaeus n. sp. Heer 1853
Heer 1853 1 specimen
lower quarry
Insecta - Hemiptera - Pentatomidae
Eurydema arcuata n. sp. Heer 1853
Heer 1853 2 specimens
from the Kesselstein
Eurydema impudica Heer 1853
Heer 1853 3 specimens
Pentatoma pictum n. sp. Heer 1864
Heer 1864 1 specimen
Halys bruckmanni Heer 1853
Heer 1853 1 specimen
from the Kesselstein
Halys spectabilis n. sp. Heer 1864
Heer 1864 1 specimen
Insecta - Hemiptera - Cydnidae
Cydnopsis deleta n. sp. Heer 1853
Heer 1853 1 specimen
lower quarry
Cydnopsis exilis n. sp. Heer 1853
Heer 1853 1 specimen
Cydnopsis pygmaea n. sp. Heer 1853
Heer 1853 1 specimen
upper quarry
Cydnus oeningensis Heer 1853
Heer 1853 1 specimen
lower quarry
Insecta - Hemiptera - Rhyparochromidae
Pachymerus obsoletus n. sp. Heer 1853
Heer 1853 1 specimen
lower quarry
Insecta - Hemiptera - Lygaeidae
Cephalocoris pilosus n. gen., n. sp. Heer 1853
Heer 1853 1 specimen
lower quarry
Insecta - Hemiptera - Heterogastridae
Heterogaster tristis n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Heterogaster
Insecta - Thysanoptera
Thrips annosa n. sp. Heer 1864
Heer 1864 1 specimen
nomen dubium belonging to Thysanoptera
Thrips oeningensis n. sp. Heer 1864
Heer 1864 1 specimen
nomen nudum belonging to Thysanoptera
see common names

Geography
Country:Germany State/province:Baden-Württemberg
Coordinates: 47.7° North, 8.9° East (view map)
Paleocoordinates:47.9° North, 7.8° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
Stage:Serravallian 10 m.y. bin:Cenozoic 5
Key time interval:Sarmatian Mammal zone: MN 7
Age range of interval:12.70000 - 11.60800 m.y. ago
Stratigraphy
Formation:Upper Freshwater-Molasse Member:Upper Oehningen beds
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology: lithified "limestone"
Environment:lacustrine - large Tectonic setting:foreland basin
Geology comments: "Rising into massive mountains, as in the well-known Righi and Rossberg, they attain a thickness of more than 6000 feet. While they include proofs of the presenceof the sea, they have preserved with marvellous perfection a large number of the plants which clothed the Alps, and of the insects which flitted through the woodlands. They form part of a great series of deposits which have been termed “Molasse” by the Swiss geologists." Geikie, 1882
"...deposited in the Northern Alpine Foreland basin..." Uhl et al., 2006
Taphonomy
Modes of preservation:mold/impression,adpression
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:survey of museum collection
Reason for describing collection:taxonomic analysis
Collection method comments: Collection of Zurich University
Metadata
Database number:117252
Authorizer:M. Clapham Enterer:M. Clapham, A. Delelli, J. Karr
Modifier:M. Clapham Research group:paleoentomology
Created:2011-09-18 05:12:10 Last modified:2013-11-25 13:02:06
Access level:the public Released:2011-09-18 05:12:10
Creative Commons license:CC BY
Reference information

Primary reference:
37344. O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]

Secondary references:

44456 T. D. A. Cockerell. 1909. Some European fossil bees. The Entomologist 42:313-317 [M. Clapham/M. Clapham]
40480 T. D. A. Cockerell. 1915. Miocene fossil insects. Proceedings of the Academy of Natural Sciences of Philadelphia 66:634-648 [M. Clapham/M. Clapham/M. Clapham]
46637 M. Fikáček and H. Schmied. 2013. Insect fauna of the late Miocene locality of Öhningen (Germany) less diverse than reported: an example of the Hydrophilid beetles (Coleoptera). Journal of Paleontology 87:427-443 [M. Clapham/M. Clapham/M. Clapham]
50838 M. H. Hatch. 1927. A revision of fossil Gyrinidae. Bulletin of the Brooklyn Entomological Society 22:89-96 [M. Clapham/M. Clapham]
38897 O. Heer. 1847. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Erster Theil: Käfer 1-229 [M. Clapham/M. Clapham]
39159 O. Heer. 1853. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Dritte Theil: Rhynchoten 1-138 [M. Clapham/M. Clapham/M. Uhen]
50522 O. Heer. 1860. Ueber die fossilen Calosomen. Programm der Eidgenössisches Polytechnischen Schule für das Schuljahr 1860/61 1-10 [M. Clapham/M. Clapham]
49902 O. Heer. 1862. Beiträge zur Insektenfauna Oeningens: Coleoptera. Geodephagen, Hydrocanthariden, Gyriniden, Brachelytren, Clavicornen, Lamellicornen und Buprestiden. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem 16(2):1-90 [M. Clapham/M. Clapham]
50652 O. Heer. 1864. Ueber die fossilen Kakerlaken. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zürich 9:273-302 [M. Clapham/M. Clapham]
47508 O. Heer. 1864. Die Urwelt der Schweiz 1-622 [M. Clapham/M. Clapham/M. Clapham]
38709 O. Heer. 1867. Fossile Hymenopteren aus Oeningen und Radoboj. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft für die Gesammten Naturwissenschaften 22:1-42 [M. Clapham/M. Clapham/M. Clapham]
48996 O. Heer. 1869. Contributions to the fossil flora of north Greenland, being a description of the plants collected by Mr. Edward Whymper during the summer of 1867. Philosophical Transactions of the Royal Society of London 159:445-488 [M. Clapham/M. Clapham]
50579 O. Heer. 1879. Die Urwelt der Schweiz, Zweite Auflage 1-713 [M. Clapham/M. Clapham]
44111 A. A. Legalov. 2013. New and little known weevils (Coleoptera: Curculionoidea) from the Paleogene and Neogene. Historical Biology 25:59-80 [M. Clapham/M. Clapham]
64986 A. A. Legalov. 2018. A new tribe of the subfamily Cyladinae (Coleoptera, Brentidae) in the Miocene of Germany. Paleontological Journal 52:35-38 [M. Clapham/M. Clapham]
36994 A. Nel and J. C. Paicheler. 1994. Les Lestoidea (Odonata, Zygoptera) fossiles: Un inventaire critique. Annales de Paléontologie 80(1):1-59 [M. Clapham/J. Karr/J. Karr]
53927 J. Skartveit and M. Pika. 2014. Revision of Bibionidae (Diptera) named by Oswald Heer from the Miocene of Öhningen, southern Germany. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 87:103-134 [M. Clapham/M. Clapham]
5180ETE 892F. E. Zeuner and F. J. Manning. 1976. A monograph on fossil bees (Hymenoptera: Apoidea). Bulletin of the British Museum (Natural History), Geology 27(3):149-268 [C. Labandeira/C. Labandeira/M. Clapham]