Soyana (PIN collection 117) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Yu. D. Zekkel in 1935

Primary reference: A. V. Martynov. 1938. Permian fossil Insects from the Arkhangelsk district. Part V. The family Euthygrammidae and its relationships (with the description of a new genus and family from Chekarda). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7(3):69-80 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104716: authorized by Matthew Clapham, entered by Jered Karr on 08.02.2011, edited by Matthew Clapham, Lindy Edwards and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Permothemistida - Permothemistidae
Permothemis intermedius n. sp.31, Permothemis lepidus n. sp.31
Permothemis intermedius n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2529(16), 45
Permothemis lepidus n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2554
 Palaeodictyoptera - Eukulojidae
Eukuloja atrosumma32, Eukuloja cubitalis32
Eukuloja atrosumma32 Sinitshenkova 1981 winged insect
PIN 117/24, 23(28), 47
Eukuloja cubitalis32 Martynov 1932 winged insect
PIN 117/18, 20-22, 25-27, 29-31, 34-36, 41(42), 46
 Paoliida - Paoliidae
Permotermopsis roseni34, Permotermopsis pectinata34, Stenaropodites sojanensis34, Stenaropodites mistshenkoi n. sp.34, Sojanidelia fluctuosa34, Sojanidelia makarkini34, Sojanidelia longula n. sp.34, Sojanidelia parvula34, Sojanidelia lineata34
Permotermopsis roseni34 Martynov 1937 winged insect
PIN 117/265(6)
Permotermopsis pectinata34 Martynov 1937 winged insect
PIN 117/815(831), 830(7)
Stenaropodites sojanensis34 Storozhenko 1992 winged insect
PIN 117/362(372), 374, 722, 794, 979(798), 814, 817, 835(840), 846, 860, 866, 2645, 103,104, 111, 356, 384(486), 508(9), 786, 888
Stenaropodites mistshenkoi n. sp.34 Storozhenko 1992 winged insect
PIN 117/801(4), 793, 353(836), 803
Sojanidelia fluctuosa34 Storozhenko 1992 winged insect
PIN 117/874
Sojanidelia makarkini34 Storozhenko 1992 winged insect
PIN 117/368, 1133
Sojanidelia longula n. sp.34 Storozhenko 1992 winged insect
PIN 117/814(869)
Sojanidelia parvula34 Storozhenko 1992 winged insect
PIN 117/2642
Sojanidelia lineata34 Storozhenko 1992 winged insect
PIN 117/816, 817, 861, 872
 Paoliida - Anthracoptilidae
Strephoneura pallida n. sp.25, Strephoneura vasta n. sp.25
Strephoneura pallida n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/376
Strephoneura vasta n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/90
 Orthoptera - Permelcanidae
Permelcana sojanense n. gen. n. sp.28
Permelcana sojanense n. gen. n. sp.28 Sharov 1962 grasshopper
PIN 117/346, 708
 Orthoptera - Pruvostitidae
Sylvoedischia curvovenosa n. sp.29
Sylvoedischia curvovenosa n. sp.29 Sharov 1968 grasshopper
PIN 117/426
 Orthoptera - Oedischiidae
Elcanoedischia major n. gen. n. sp.17, Elcanoedischia minor n. sp.17, "Pinegia martynovi n. sp." = Tcholmanvissia noinskii16
Elcanoedischia major n. gen. n. sp.17 Gorochov 1987 grasshopper
PIN 117/463
Elcanoedischia minor n. sp.17 Gorochov 1987 grasshopper
PIN 117/645+150, 702
"Pinegia martynovi n. sp." = Tcholmanvissia noinskii16 Zalessky 1929 grasshopper
PIN 117/258+259
 Caloneurodea - Caloneuridae
Euthygramma parallellum
Euthygramma parallellum Martynov 1928 winged insect
PIN 117/315, 316, 336, 357, 283, 160, 191, 101, 603, 182, 625
 Cnemidolestodea - Pinideliidae
Idelopterum samoedum n. sp.7
Idelopterum samoedum n. sp.7 Aristov 2015 winged insect
PIN 117/43
 Cnemidolestodea - Protembiidae
Tshekardomina novokshonovi n. sp.7, Sojanoraphidia rossica n. gen. n. sp.21
Tshekardomina novokshonovi n. sp.7 Aristov 2015 winged insect
PIN 117/1528
Sojanoraphidia rossica n. gen. n. sp.21 Martynova 1952 winged insect
PIN 117/2549
 Cnemidolestodea - Sylvabestiidae
Sojanopermula rasnitsyni n. sp.10, Kazanalicula reducta n. gen. n. sp.10
Sojanopermula rasnitsyni n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/301,195
Kazanalicula reducta n. gen. n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/467
 Protorthoptera - Soyaniidae
Soyania logica8
Soyania logica8 Aristov and Rasnitsyn 2008 winged insect
PIN 177/1314,2759,2803
 Grylloblattodea - Lemmatophoridae
Uraloprisca torta n. sp.7, Sylvaprisca colorata n. sp.3, Artinska sojanensis n. sp.3
Uraloprisca torta n. sp.7 Aristov 2015 ice crawler
PIN 117/335
Sylvaprisca colorata n. sp.3 Aristov 2009 ice crawler
PIN 117/855,444,1155
Artinska sojanensis n. sp.3 Aristov 2009 ice crawler
PIN 117/1008, 1060
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis n. sp.11
Camptoneurites soyanensis n. sp.11 Aristov et al. 2010 ice crawler
PIN 117/244,807,871,2660
 Grylloblattodea - Euremiscidae
Euremisca kazanica n. sp.4
Euremisca kazanica n. sp.4 Aristov 2009 ice crawler
PIN 117/1140
 Hypoperlida - Hypoperlidae
Boreopsocus danksae30, Hypoperla elegans24
Boreopsocus danksae30 Shcherbakov 1994 winged insect
Hypoperla elegans24 Martynov 1928 winged insect
PIN 117/968
 Lophioneurida - Lophioneuridae
Lophiocypha incompleta n. sp.35
Lophiocypha incompleta n. sp.35 Vishnyakova 1981 winged insect
PIN 117/2442
 Hemiptera - Boreoscytidae
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14, "Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14, "Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1854
"Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
no number given
"Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1673
 Hemiptera - Archescytinidae
Archescytinidae indet.15, Archescytina fusca15, Archescytina gunderseni15
Archescytinidae indet.15 Tillyard 1926 true bug
PIN 117/1703
Archescytina fusca15 Becker-Migdisova 1960 true bug
PIN 117/1654(1685)
Archescytina gunderseni15 Becker-Migdisova 1960 true bug
PIN 117/1991, 1855, 1931, 1798, 1934
 Hemiptera - Scytinopteridae
Scytinoptera reducta13
Scytinoptera reducta13 Martynov 1928 true bug
PIN 117/1667, 1666, 2022, 1963, 577
 Hemiptera - Prosbolopseidae
Ivaia procercopoides n. sp.15, Scytoneura elliptica15, Anaprosbole ivensis n. gen. n. sp.15, Prosbolopsis ovalis15
Ivaia procercopoides n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/1914
Scytoneura elliptica15 Martynov 1935 true bug
PIN 117/1659(83), 1711, 2040, 1817(2127),1644(2039), 2068, 1694, 1693, 1876, 2044
Anaprosbole ivensis n. gen. n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/2020
Prosbolopsis ovalis15 Martynov 1935 true bug
PIN 117/1819
 Hemiptera - Prosbolidae
Permocicadopsis angustata12, Permocicadopsis aberrans12, Permocicada nigronervosa12, Permocicada integra12, Permocicada triramosa12, Permocicada idelensis12, Permocicada elliptica12, Permocicada simulatrix12, Prosbole zekkeli n. sp.12, Prosbole dilatata12, Prosbole curvimarginata n. sp.12, Sojanoneura "sp. 1 nov"12, Sojanoneura kazanensis12, Sojanoneura tincta n. sp.12, Sojanoneura triangularis12, Sojanoneura bimaculata12, Sojanoneura edemskii12, Sojanoneura elytrata12
Permocicadopsis angustata12 Martynov 1935 true bug
PIN 117/2131; 1665, 1679(2136) (var. typica); 1642(var. brevifurcillata); 1647(var. devia); 2132(var. remota)
Permocicadopsis aberrans12 Martynov 1935 true bug
PIN 117/1684, 1656, 1651
Permocicada nigronervosa12 Martynov 1928 true bug
PIN 117/2016(2019), 1702, 1648, (2046, 2070(2057) (var. longifurcata))
Permocicada integra12 Becker-Migdisova 1940 true bug
PIN 117/1705, 1662
Permocicada triramosa12 Becker-Migdisova 1940 true bug
PIN 117/1709
Permocicada idelensis12 Zalessky 1929 true bug
PIN 117/1655, 1701(2063), 2060, 1696
Permocicada elliptica12 Martynov 1935 true bug
PIN 117/2135
Permocicada simulatrix12 Martynov 1935 true bug
PIN 117/2049
Prosbole zekkeli n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2047(2139)
Prosbole dilatata12 Martynov 1935 true bug
PIN 117/2050
Prosbole curvimarginata n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/1687
Sojanoneura "sp. 1 nov"12 Martynov 1928 true bug
PIN 117/2065
Sojanoneura kazanensis12 Zalessky 1929 true bug
PIN 117/2064(1652)
Sojanoneura tincta n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2129
Sojanoneura triangularis12 Martynov 1935 true bug
PIN 117/1660, 1697(2014)
Sojanoneura bimaculata12 Martynov 1935 true bug
PIN 117/2009, 1695, 2059, 1713, 2058
Sojanoneura edemskii12 Martynov 1928 true bug
PIN 117/1675(2051), 2037(1691), 1646(2059), 2134(2062)
Sojanoneura elytrata12 Martynov 1928 true bug
PIN 117/1654(1641)
 Glosselytrodea - Jurinidae
Jurina marginata18
Jurina marginata18 Martynov 1938 winged insect
PIN 117/584
 Dicondylia - Permosialidae
Permosialis perfecta n. sp.21, Permosialis latiformis n. sp.21, "Permosialis fasciata n. sp." = Permosialis paucinervis21, "Permosialis defurcata n. sp." = Permosialis paucinervis21, "Permosialis bifasciata" = Permosialis paucinervis21, "Permosialis quadriramosa n. sp." = Palaeomantopsis furcatella21
Permosialis perfecta n. sp.21 Martynova 1952 winged insect
PIN 117/1772
Permosialis latiformis n. sp.21 Martynova 1952 winged insect
PIN 117/1153, 2653
"Permosialis fasciata n. sp." = Permosialis paucinervis21 Martynov 1928 winged insect
PIN 117/2651
"Permosialis defurcata n. sp." = Permosialis paucinervis21 Martynov 1928 winged insect
PIN 117/1316
"Permosialis bifasciata" = Permosialis paucinervis21 Martynov 1928 winged insect
PIN 117/1688(2656)
"Permosialis quadriramosa n. sp." = Palaeomantopsis furcatella21 Martynov 1928 winged insect
PIN 117/1444(2652)
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis20
Kamopanorpa rotundipennis20 Martynov 1932 winged insect
PIN 117/2695
 Mecoptera - Permochoristidae
"Agetochorista dubia" = Agetopanorpa dubia20, "Agetochorista fasciata" = Agetopanorpa fasciata20, "Agetochorista ornata" = Agetopanorpa ornata20, "Agetochorista splendida" = Agetopanorpa ornata20, "Agetochorista letopalae" = Agetopanorpa letopalae20, Petromantis sp.23, ? Petromantis sp.23, "Petrochorista dubia" = Petromantis dubia20, "Petromantis rossica" = Petromantis rossica20, "Petromantis borealis" = Petromantis rossica20, "Petromantis robusta" = Petromantis robusta20, "Petromantis variolosa" = Petromantis robusta20, "Permochorista maculipennis" = Petromantis maculipennis20, "Permochorista venosa" = Petromantis maculipennis20, Petrochorista ivae n. sp.20, Petrochorista sojanensis20
"Agetochorista dubia" = Agetopanorpa dubia20 Martynov 1932 scorpionfly
PIN 117/1401
"Agetochorista fasciata" = Agetopanorpa fasciata20 Martynov 1932 scorpionfly
PIN 117/1325, 1343, 2745
"Agetochorista ornata" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1335, 1347, 1368, 1373, 1377, 1380, 1381, 1383, 1395, 1415, 1423, 1426, 1428, 1429, 1437, 1445
"Agetochorista splendida" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1333, 1342(1390), 1398; var. differens 117/1436
"Agetochorista letopalae" = Agetopanorpa letopalae20 Martynova 1948 scorpionfly
PIN 117/1339, 1414
Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/2176, 1629-1851
? Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/276
"Petrochorista dubia" = Petromantis dubia20 Martynov 1932 scorpionfly
PIN 117/1320
"Petromantis rossica" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1321, 1336, 1358, 1361, 1392, 1397, 1406, 1439, 2133
"Petromantis borealis" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1328, 1344, 1351, 1364, 1389, 1394, 1405, 1412, 1416, 1418, 1441, 2724, 2731
"Petromantis robusta" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1446
"Petromantis variolosa" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1420
"Permochorista maculipennis" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1332, 1334, 1340, 1367, 1376, 1379, 1393, 1403, 1411, 1433, 1440, 2687; var. major 117/1355
"Permochorista venosa" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1111, 1448
Petrochorista ivae n. sp.20 Martynova 1948 scorpionfly
PIN 117/1341, 1407
Petrochorista sojanensis20 Martynova 1948 scorpionfly
PIN 117/1404, 2685
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata20
"Permopanorpa angustata" = Martynopanorpa angustata20 Martynov 1932 scorpionfly
PIN 117/2695
 Coleoptera - Permocupedidae
Permocupes semenovi26
Permocupes semenovi26 Martynov 1932 beetle
PIN 117/399(408)
 Raphidioptera - Parasialidae
Parasialis dissedens27, Sojanasialis longipennis n. gen. n. sp.27
Parasialis dissedens27 Ponomarenko 1977 snakefly
PIN 117/2146, 2335, 2600, 26220
Sojanasialis longipennis n. gen. n. sp.27 Ponomarenko 1977 snakefly
PIN 117/2459
 Dicondylia - Permithonidae
"Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21, Permithonopsis cellulosa n. sp.21, Permithonopsis longifurcata n. sp.21, "Permithonopsis ivensis" = Permithonopsis ivensis21, "Permithonopsis proxima" = Permithonopsis ivensis21, Palaemerobius avus n. sp.21
"Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21 Martynova 1952 winged insect
PIN 117/2513
Permithonopsis cellulosa n. sp.21 Martynova 1952 winged insect
PIN 117/2533, 2560
Permithonopsis longifurcata n. sp.21 Martynova 1952 winged insect
PIN 117/2515
"Permithonopsis ivensis" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2537(2598)
"Permithonopsis proxima" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2524
Palaemerobius avus n. sp.21 Martynova 1952 winged insect
PIN 117/2532(2592), 2540
 Dicondylia - Permagrionidae
Epilestes angustapterix n. sp.22, Solikamptilon pectinatus n. sp.22
Epilestes angustapterix n. sp.22 Nel et al. 2012 winged insect
PIN 117/63
Solikamptilon pectinatus n. sp.22 Nel et al. 2012 winged insect
PIN 117/13
 Dicondylia - Permepallagidae
Permepallage angustissima n. gen. n. sp.19
Permepallage angustissima n. gen. n. sp.19 Martynov 1938 winged insect
PIN 117/65 (says Tshekarda in Nel et al. 2012)
 Dicondylia - Kennedyidae
Kennedya ivensis n. sp.22
Kennedya ivensis n. sp.22 Nel et al. 2012 winged insect
PIN 117/15; 117/14(128) in Felker 2023
 Reculida - Ivapteridae
Ivaptera sharovi n. gen. n. sp.2
Ivaptera sharovi n. gen. n. sp.2 Aristov 2009 winged insect
PIN 117/852, 140, 1085, 113, 123, 139, 224, 284, 322, 339, 379, 539, 565, 639, 783, 785, 825, 865, 873, 887, 897,945,950,964,979,990,992,994,995,999,1004,1005,1007,1009,1014,1019,1029,1038,1049,1050,1061,1071,1077,1081; PIN 117/477, 1134 in Aristov, 2015
 Reculida - Geinitziidae
Geinitzia ima n. sp.7, Stegopterum primus n. sp.7, Permoshurabia mesenensis n. gen. n. sp.4, Shurabia inferior n. sp.7
Geinitzia ima n. sp.7 Aristov 2015 winged insect
PIN 117/805
Stegopterum primus n. sp.7 Aristov 2015 winged insect
PIN 117/765
Permoshurabia mesenensis n. gen. n. sp.4 Aristov 2009 winged insect
PIN 117/806, 102, 2774
Shurabia inferior n. sp.7 Aristov 2015 winged insect
PIN 117/282
 Eoblattida - Bardapteridae
Olgaephilus emendatus33, Letopala costalis n. gen. n. sp.9
Olgaephilus emendatus33 Storozhenko 1990 insect
PIN 117/844, 383, 842
Letopala costalis n. gen. n. sp.9 Aristov and Rasnitsyn 2011 insect
PIN 117/617
 Eoblattida - Mesorthopteridae
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1, Parastenaropodites aquilonius n. sp.6
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1 Aristov 2008 insect
PIN 117/144, 176, 740, 975, 982, 1082
Parastenaropodites aquilonius n. sp.6 Aristov 2014 insect
PIN 117/223
 Eoblattida - Blattogryllidae
Permoblattogryllus praecox n. gen. n. sp.5
Permoblattogryllus praecox n. gen. n. sp.5 Aristov 2011 insect
PIN 117/83
 Eoblattida - Megakhosaridae
Ivakhosara pectinimedia n. gen. n. sp.1, Megakhosarella paula n. sp.7, "Megakhosarina minuscula n. sp." = Blattokhosara minuscula1
Ivakhosara pectinimedia n. gen. n. sp.1 Aristov 2008 insect
PIN 117/2488
Megakhosarella paula n. sp.7 Aristov 2015 insect
PIN 117/996
"Megakhosarina minuscula n. sp." = Blattokhosara minuscula1 Aristov 2008 insect
PIN 117/86; 117/820 in Storozhenko & Aristov, 2014
1Aristov 2008; 2Aristov 2009; 3Aristov 2009; 4Aristov 2009; 5Aristov 2011; 6Aristov 2014; 7Aristov 2015; 8Aristov and Rasnitsyn 2008; 9Aristov and Rasnitsyn 2011; 10Aristov and Storozhenko 2011; 11Aristov et al. 2010; 12Becker-Migdisova 1940; 13Becker-Migdisova 1948; 14Becker-Migdisova 1949; 15Becker-Migdisova 1960; 16Béthoux and Nel 2002; 17Gorochov 1987; 18Martynov 1938; 19Martynov 1938; 20Martynova 1948; 21Martynova 1952; 22Nel et al. 2012; 23Novokshonov 1994; 24Novokshonov 2001; 25Novokshonov and Aristov 2004; 26Ponomarenko 1963; 27Ponomarenko 1977; 28Sharov 1962; 29Sharov 1968; 30Shcherbakov 1994; 31Sinitshenkova 1980; 32Sinitshenkova 1981; 33Storozhenko 1990; 34Storozhenko 1992; 35Vishnyakova 1981