Soyana (PIN collection 117) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Yu. D. Zekkel in 1935

Primary reference: A. V. Martynov. 1938. Permian fossil Insects from the Arkhangelsk district. Part V. The family Euthygrammidae and its relationships (with the description of a new genus and family from Chekarda). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7(3):69-80 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104716: authorized by Matthew Clapham, entered by Jered Karr on 08.02.2011, edited by Matthew Clapham, Lindy Edwards and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hypoperlida - Hypoperlidae
Hypoperla elegans24 Martynov 1928 winged insect
PIN 117/968
Boreopsocus danksae30 Shcherbakov 1994 winged insect
 Hemiptera - Scytinopteridae
Scytinoptera reducta13 Martynov 1928 true bug
PIN 117/1667, 1666, 2022, 1963, 577
 Hemiptera - Archescytinidae
Archescytinidae indet.15 Tillyard 1926 true bug
PIN 117/1703
Archescytina fusca15 Becker-Migdisova 1960 true bug
PIN 117/1654(1685)
Archescytina gunderseni15 Becker-Migdisova 1960 true bug
PIN 117/1991, 1855, 1931, 1798, 1934
 Hemiptera - Boreoscytidae
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14, "Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14, "Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14
"Boreoscyta nefasta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1854
"Boreoscyta imperfecta n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
PIN 117/1673
"Boreoscyta mirabilis n. gen. n. sp." = Boreoscyta nefasta14 Becker-Migdisova 1949 true bug
no number given
 Hemiptera - Prosbolopseidae
Anaprosbole ivensis n. gen. n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/2020
Scytoneura elliptica15 Martynov 1935 true bug
PIN 117/1659(83), 1711, 2040, 1817(2127),1644(2039), 2068, 1694, 1693, 1876, 2044
Ivaia procercopoides n. sp.15 Becker-Migdisova 1960 true bug
PIN 117/1914
Prosbolopsis ovalis15 Martynov 1935 true bug
PIN 117/1819
 Hemiptera - Prosbolidae
Permocicada simulatrix12 Martynov 1935 true bug
PIN 117/2049
Permocicada nigronervosa12 Martynov 1928 true bug
PIN 117/2016(2019), 1702, 1648, (2046, 2070(2057) (var. longifurcata))
Permocicada triramosa12 Becker-Migdisova 1940 true bug
PIN 117/1709
Permocicada elliptica12 Martynov 1935 true bug
PIN 117/2135
Permocicada integra12 Becker-Migdisova 1940 true bug
PIN 117/1705, 1662
Permocicada idelensis12 Zalessky 1929 true bug
PIN 117/1655, 1701(2063), 2060, 1696
Permocicadopsis angustata12 Martynov 1935 true bug
PIN 117/2131; 1665, 1679(2136) (var. typica); 1642(var. brevifurcillata); 1647(var. devia); 2132(var. remota)
Permocicadopsis aberrans12 Martynov 1935 true bug
PIN 117/1684, 1656, 1651
Sojanoneura "sp. 1 nov"12 Martynov 1928 true bug
PIN 117/2065
Sojanoneura edemskii12 Martynov 1928 true bug
PIN 117/1675(2051), 2037(1691), 1646(2059), 2134(2062)
Sojanoneura bimaculata12 Martynov 1935 true bug
PIN 117/2009, 1695, 2059, 1713, 2058
Sojanoneura elytrata12 Martynov 1928 true bug
PIN 117/1654(1641)
Sojanoneura triangularis12 Martynov 1935 true bug
PIN 117/1660, 1697(2014)
Sojanoneura kazanensis12 Zalessky 1929 true bug
PIN 117/2064(1652)
Sojanoneura tincta n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2129
Prosbole dilatata12 Martynov 1935 true bug
PIN 117/2050
Prosbole zekkeli n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/2047(2139)
Prosbole curvimarginata n. sp.12 Becker-Migdisova 1940 true bug
PIN 117/1687
 Dicondylia - Lophioneuridae
Lophiocypha incompleta n. sp.35 Vishnyakova 1981 winged insect
PIN 117/2442
 Dicondylia - Permosialidae
Permosialis quadriramosa n. sp.21 Martynova 1952 winged insect
PIN 117/1444(2652)
Permosialis latiformis n. sp.21 Martynova 1952 winged insect
PIN 117/1153, 2653
Permosialis bifasciata21 Martynov 1932 winged insect
PIN 117/1688(2656)
Permosialis fasciata n. sp.21 Martynova 1952 winged insect
PIN 117/2651
Permosialis perfecta n. sp.21 Martynova 1952 winged insect
PIN 117/1772
Permosialis defurcata n. sp.21 Martynova 1952 winged insect
PIN 117/1316
 Cnemidolestodea - Sylvabestiidae
Sojanopermula rasnitsyni n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/301,195
Kazanalicula reducta n. gen. n. sp.10 Aristov and Storozhenko 2011 winged insect
PIN 117/467
 Cnemidolestodea - Protembiidae
Sojanoraphidia rossica n. gen. n. sp.21 Martynova 1952 winged insect
PIN 117/2549
Tshekardomina novokshonovi n. sp.7 Aristov 2015 winged insect
PIN 117/1528
 Cnemidolestodea - Pinideliidae
Idelopterum samoedum n. sp.7 Aristov 2015 winged insect
PIN 117/43
 Caloneurodea - Caloneuridae
Euthygramma parallellum Martynov 1928 winged insect
PIN 117/315, 316, 336, 357, 283, 160, 191, 101, 603, 182, 625
 Protorthoptera - Soyaniidae
Soyania logica8 Aristov and Rasnitsyn 2008 winged insect
PIN 177/1314,2759,2803
 Orthoptera - Pruvostitidae
Sylvoedischia curvovenosa n. sp.29 Sharov 1968 grasshopper
PIN 117/426
 Orthoptera - Oedischiidae
Elcanoedischia major n. gen. n. sp.17, Elcanoedischia minor n. sp.17, "Pinegia martynovi n. sp." = Tcholmanvissia noinskii16
Elcanoedischia major n. gen. n. sp.17 Gorochov 1987 grasshopper
PIN 117/463
Elcanoedischia minor n. sp.17 Gorochov 1987 grasshopper
PIN 117/645+150, 702
"Pinegia martynovi n. sp." = Tcholmanvissia noinskii16 Zalessky 1929 grasshopper
PIN 117/258+259
 Orthoptera - Permelcanidae
Permelcana sojanense n. gen. n. sp.28
Permelcana sojanense n. gen. n. sp.28 Sharov 1962 grasshopper
PIN 117/346, 708
 Paoliida - Paoliidae
Stenaropodites sojanensis34 Storozhenko 1992 winged insect
PIN 117/362(372), 374, 722, 794, 979(798), 814, 817, 835(840), 846, 860, 866, 2645, 103,104, 111, 356, 384(486), 508(9), 786, 888
Stenaropodites mistshenkoi n. sp.34 Storozhenko 1992 winged insect
PIN 117/801(4), 793, 353(836), 803
Sojanidelia fluctuosa34 Storozhenko 1992 winged insect
PIN 117/874
Sojanidelia parvula34 Storozhenko 1992 winged insect
PIN 117/2642
Sojanidelia lineata34 Storozhenko 1992 winged insect
PIN 117/816, 817, 861, 872
Sojanidelia longula n. sp.34 Storozhenko 1992 winged insect
PIN 117/814(869)
Sojanidelia makarkini34 Storozhenko 1992 winged insect
PIN 117/368, 1133
 Paoliida - Anthracoptilidae
Strephoneura pallida n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/376
Strephoneura vasta n. sp.25 Novokshonov and Aristov 2004 winged insect
PIN 117/90
 Grylloblattodea - Euremiscidae
Euremisca kazanica n. sp.2 Aristov 2009 ice crawler
PIN 117/1140
 Grylloblattodea - Camptoneuritidae
Camptoneurites soyanensis n. sp.11 Aristov et al. 2010 ice crawler
PIN 117/244,807,871,2660
 Glosselytrodea - Jurinidae
Jurina marginata18 Martynov 1938 winged insect
PIN 117/584
 Coleoptera - Permocupedidae
Permocupes semenovi26 Martynov 1932 beetle
PIN 117/399(408)
 Mecoptera - Permochoristidae
Petromantis sp.23, ? Petromantis sp.23, "Petromantis robusta" = Petromantis robusta20, "Petromantis variolosa" = Petromantis robusta20, "Petromantis rossica" = Petromantis rossica20, "Petromantis borealis" = Petromantis rossica20, "Permochorista maculipennis" = Petromantis maculipennis20, "Permochorista venosa" = Petromantis maculipennis20, "Petrochorista dubia" = Petromantis dubia20, Petrochorista ivae n. sp.20, Petrochorista sojanensis20, "Agetochorista letopalae" = Agetopanorpa letopalae20, "Agetochorista ornata" = Agetopanorpa ornata20, "Agetochorista splendida" = Agetopanorpa ornata20, "Agetochorista dubia" = Agetopanorpa dubia20, "Agetochorista fasciata" = Agetopanorpa fasciata20
Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/2176, 1629-1851
? Petromantis sp.23 Handlirsch 1904 scorpionfly
PIN 117/276
"Petromantis robusta" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1446
"Petromantis variolosa" = Petromantis robusta20 Martynov 1932 scorpionfly
PIN 117/1420
"Petromantis rossica" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1321, 1336, 1358, 1361, 1392, 1397, 1406, 1439, 2133
"Petromantis borealis" = Petromantis rossica20 Handlirsch 1904 scorpionfly
PIN 117/1328, 1344, 1351, 1364, 1389, 1394, 1405, 1412, 1416, 1418, 1441, 2724, 2731
"Permochorista maculipennis" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1332, 1334, 1340, 1367, 1376, 1379, 1393, 1403, 1411, 1433, 1440, 2687; var. major 117/1355
"Permochorista venosa" = Petromantis maculipennis20 Martynov 1932 scorpionfly
PIN 117/1111, 1448
"Petrochorista dubia" = Petromantis dubia20 Martynov 1932 scorpionfly
PIN 117/1320
Petrochorista ivae n. sp.20 Martynova 1948 scorpionfly
PIN 117/1341, 1407
Petrochorista sojanensis20 Martynova 1948 scorpionfly
PIN 117/1404, 2685
"Agetochorista letopalae" = Agetopanorpa letopalae20 Martynova 1948 scorpionfly
PIN 117/1339, 1414
"Agetochorista ornata" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1335, 1347, 1368, 1373, 1377, 1380, 1381, 1383, 1395, 1415, 1423, 1426, 1428, 1429, 1437, 1445
"Agetochorista splendida" = Agetopanorpa ornata20 Martynov 1932 scorpionfly
PIN 117/1333, 1342(1390), 1398; var. differens 117/1436
"Agetochorista dubia" = Agetopanorpa dubia20 Martynov 1932 scorpionfly
PIN 117/1401
"Agetochorista fasciata" = Agetopanorpa fasciata20 Martynov 1932 scorpionfly
PIN 117/1325, 1343, 2745
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata20
"Permopanorpa angustata" = Martynopanorpa angustata20 Martynov 1932 scorpionfly
PIN 117/2695
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis20 Martynov 1932 winged insect
PIN 117/2695
 Raphidioptera - Parasialidae
Parasialis dissedens27 Ponomarenko 1977 snakefly
PIN 117/2146, 2335, 2600, 26220
Sojanasialis longipennis n. gen. n. sp.27 Ponomarenko 1977 snakefly
PIN 117/2459
 Dicondylia - Permithonidae
Permithonopsis cellulosa n. sp.21, "Permithonopsis ivensis" = Permithonopsis ivensis21, "Permithonopsis proxima" = Permithonopsis ivensis21, Permithonopsis longifurcata n. sp.21, "Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21, Palaemerobius avus n. sp.21
Permithonopsis cellulosa n. sp.21 Martynova 1952 winged insect
PIN 117/2533, 2560
"Permithonopsis ivensis" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2537(2598)
"Permithonopsis proxima" = Permithonopsis ivensis21 Martynov 1932 winged insect
PIN 117/2524
Permithonopsis longifurcata n. sp.21 Martynova 1952 winged insect
PIN 117/2515
"Bianchia spectabilis n. gen. n. sp." = Permegalomus spectabilis21 Martynova 1952 winged insect
PIN 117/2513
Palaemerobius avus n. sp.21 Martynova 1952 winged insect
PIN 117/2532(2592), 2540
 Palaeodictyoptera - Eukulojidae
Eukuloja atrosumma32 Sinitshenkova 1981 winged insect
PIN 117/24, 23(28), 47
Eukuloja cubitalis32 Martynov 1932 winged insect
PIN 117/18, 20-22, 25-27, 29-31, 34-36, 41(42), 46
 Palaeodictyoptera - Permothemistidae
Permothemis lepidus n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2554
Permothemis intermedius n. sp.31 Sinitshenkova 1980 winged insect
PIN 117/2529(16), 45
 Reculida - Geinitziidae
Permoshurabia mesenensis n. gen. n. sp.2 Aristov 2009 winged insect
PIN 117/806, 102, 2774
Shurabia inferior n. sp.7 Aristov 2015 winged insect
PIN 117/282
Geinitzia ima n. sp.7 Aristov 2015 winged insect
PIN 117/805
Stegopterum primus n. sp.7 Aristov 2015 winged insect
PIN 117/765
 Reculida - Ivapteridae
Ivaptera sharovi n. gen. n. sp.3
Ivaptera sharovi n. gen. n. sp.3 Aristov 2009 winged insect
PIN 117/852, 140, 1085, 113, 123, 139, 224, 284, 322, 339, 379, 539, 565, 639, 783, 785, 825, 865, 873, 887, 897,945,950,964,979,990,992,994,995,999,1004,1005,1007,1009,1014,1019,1029,1038,1049,1050,1061,1071,1077,1081; PIN 117/477, 1134 in Aristov, 2015
 Reculida - Lemmatophoridae
Uraloprisca torta n. sp.7 Aristov 2015 winged insect
PIN 117/335
Artinska sojanensis n. sp.4 Aristov 2009 winged insect
PIN 117/1008, 1060
Sylvaprisca colorata n. sp.4 Aristov 2009 winged insect
PIN 117/855,444,1155
 Dicondylia - Permepallagidae
Permepallage angustissima n. gen. n. sp.19
Permepallage angustissima n. gen. n. sp.19 Martynov 1938 winged insect
PIN 117/65 (says Tshekarda in Nel et al. 2012)
 Dicondylia - Kennedyidae
Kennedya ivensis n. sp.22 Nel et al. 2012 winged insect
PIN 117/15
 Dicondylia - Permagrionidae
Epilestes angustapterix n. sp.22 Nel et al. 2012 winged insect
PIN 117/63
Solikamptilon pectinatus n. sp.22 Nel et al. 2012 winged insect
PIN 117/13
 Eoblattida - Permotermopsidae
Permotermopsis pectinata34 Martynov 1937 insect
PIN 117/815(831), 830(7)
Permotermopsis roseni34 Martynov 1937 insect
PIN 117/265(6)
 Eoblattida - Mesorthopteridae
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1, Parastenaropodites aquilonius n. sp.6
"Megakhosarina intricata n. sp." = Parastenaropodites intricatus1 Aristov 2008 insect
PIN 117/144, 176, 740, 975, 982, 1082
Parastenaropodites aquilonius n. sp.6 Aristov 2014 insect
PIN 117/223
 Eoblattida - Megakhosaridae
"Megakhosarina minuscula n. sp." = Blattokhosara minuscula1, Megakhosarella paula n. sp.7, Ivakhosara pectinimedia n. gen. n. sp.1
"Megakhosarina minuscula n. sp." = Blattokhosara minuscula1 Aristov 2008 insect
PIN 117/86; 117/820 in Storozhenko & Aristov, 2014
Megakhosarella paula n. sp.7 Aristov 2015 insect
PIN 117/996
Ivakhosara pectinimedia n. gen. n. sp.1 Aristov 2008 insect
PIN 117/2488
 Eoblattida - Bardapteridae
Letopala costalis n. gen. n. sp.9 Aristov and Rasnitsyn 2011 insect
PIN 117/617
Olgaephilus emendatus33 Storozhenko 1990 insect
PIN 117/844, 383, 842
 Eoblattida - Blattogryllidae
Permoblattogryllus praecox n. gen. n. sp.5 Aristov 2011 insect
PIN 117/83