Isady Locality (Mutovino, PIN collection 3840) (Permian of Russian Federation)

Where: Vologda, Russian Federation (60.6° N, 45.6° E: paleocoordinates 33.1° N, 36.5° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Proelginia permiana other zone, Kichuga Member (Poldarsa Formation), Wuchiapingian (259.9 - 254.2 Ma)

• Chroniosaurus levis vertebrate subzone

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine; lithified siltstone

Size class: macrofossils

Preservation: mold/impression

Primary reference: N. D. Sinitshenkova and D. S. Aristov. 2010. New Permian Stoneflies of the Family Palaeonemouridae (Insecta: Perlida = Plecoptera) from the Isady Locality. Paleontological Journal 44:49-52 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104754: authorized by Matthew Clapham, entered by Matthew Clapham on 09.02.2011, edited by Joseph Flannery-Sutherland

Creative Commons license: CC BY (attribution)

Taxonomic list

• Locality also includes flora, bivalves, ostracods, scorpions, fish scales, tetrapods
unclassified
  -
Bivalvia
 Cardiida - Opokiellidae
 Actinodontida - Palaeomutelidae
Insecta
 Eoblattida - Atactophlebiidae
Issadophlebia fusa n. gen. n. sp.9
Issadophlebia fusa n. gen. n. sp.9 Aristov 2015 insect
PIN 3840/3164
 Eoblattida - Blattogryllidae
Baharellinus porrectus n. sp.7 Aristov 2013 insect
PIN 3840/1244
Baharellinus dilaceratus n. sp.5 Aristov 2013 insect
PIN 3840/3109
 Eoblattida - Soyanopteridae
Poldarsia relictaria n. gen. n. sp.13
Poldarsia relictaria n. gen. n. sp.13 Aristov and Rasnitsyn 2011 insect
PIN 3840/793,2040 (1 km upstream from Isady)
 Eoblattida - Mesorthopteridae
Parastenaropodites exossis n. sp.8 Aristov 2014 insect
PIN 3840/1235, 3119
Parastenaropodites panneus n. sp.8 Aristov 2014 insect
PIN 3840/1233
Mesoidelia gorochovi n. sp.7 Aristov 2013 insect
PIN 3840/652
Mesoidelia procera n. sp.8 Aristov 2014 insect
PIN 3840/3112
Mesoidelia riphaea n. sp.8 Aristov 2014 insect
PIN 3840/1535
Tshermyaninus biforis n. gen. n. sp.8 Aristov 2014 insect
PIN 3840/1320
Permorthopteron foliaceus n. gen. n. sp.8 Aristov 2014 insect
PIN 3840/3108
 Eoblattida - Protophasmatidae
Permeoblatta borealis n. gen. n. sp.27
Permeoblatta borealis n. gen. n. sp.27 Rasnitsyn and Aristov 2010 insect
PIN 3840/845, 3111 (in Aristov, 2013)
 Eoblattida - Bardapteridae
Issadische maximum n. gen. n. sp.6
Issadische maximum n. gen. n. sp.6 Aristov 2013 insect
PIN 3840/2099, 3110
 Eoblattida - Megakhosaridae
Megakhosarodes tensilis n. sp.5 Aristov 2013 insect
PIN 3840/3106, 3107
Parakhosara sp.3 Storozhenko 1993 insect
PIN 3840/92
Parakhosara kopylovi n. sp.3 Aristov 2009 insect
PIN 3840/430
Parakhosara reticulata n. sp.7 Aristov 2013 insect
PIN 3840/1248
Parakhosara mutoviensis n. sp.3 Aristov 2009 insect
PIN 3840/33
Abbrevikhosara ovoidea n. gen. n. sp.7 Aristov 2013 insect
PIN 3840/1249
 Reculida - Chaulioditidae
Purtovinia ustyugensis n. gen. n. sp.7 Aristov 2013 winged insect
PIN 3840/543
Parachauliodites orthopteroides n. gen. n. sp.5 Aristov 2013 winged insect
PIN 3840/3105
Chauliodites geniatus n. sp.7 Aristov 2013 winged insect
PIN 3840/1255
Chauliodites ponomarenkoi n. sp.2 Aristov 2008 winged insect
PIN 3840/29 (1.2 km downstream)
Chauliodites circumornatus n. sp.7 Aristov 2013 winged insect
PIN 3840/1258, 1257, 1259, 1260, 1262
Chauliodites issadensis n. sp.3 Aristov 2009 winged insect
PIN 3840/91; 547, 1256 in Aristov, 2013
Permyak involucris n. gen. n. sp.7 Aristov 2013 winged insect
PIN 3840/2091
 Reculida - Geinitziidae
Geinitzia subita n. sp.7 Aristov 2013 winged insect
PIN 3840/1239
Shurabia permiana n. sp.7 Aristov 2013 winged insect
PIN 3840/3101, 548, 549, 1237, 1240, 3100, 3103, 3104 (not 3102 = Permoshurabia sukhanica)
Permoshurabia sukhonica n. sp.12 Aristov 2022 winged insect
PIN 3840/3102
Sukhonia coriacea n. gen. n. sp.7 Aristov 2013 winged insect
PIN 3840/796
 Megasecoptera - Moravohymenidae
Issadohymen ponomarenkoi n. gen. n. sp.33
Issadohymen ponomarenkoi n. gen. n. sp.33 Sinitshenkova and Aristov 2013 winged insect
PIN 3840/799
 Dicondylia - Permosialidae
Epimastax mutovinensis n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1660, 559, 2078
 Glosselytrodea - Jurinidae
Isadelytron speciosum n. gen. n. sp.28
Isadelytron speciosum n. gen. n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/443
 Glosselytrodea - Glosselytridae
? Karajurina desperata n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/398
 Orthoptera - Proparagryllacrididae
Archifergania isadyensis n. gen. n. sp.24
Archifergania isadyensis n. gen. n. sp.24 Gorochov 2013 grasshopper
PIN 3840/1190, 319
 Orthoptera - Pruvostitidae
Suchonoedischia incompleta n. gen. n. sp.24, Vologdoptera maculosa n. gen. n. sp.3
Suchonoedischia incompleta n. gen. n. sp.24 Gorochov 2013 grasshopper
PIN 3840/663
Vologdoptera maculosa n. gen. n. sp.3 Aristov 2009 grasshopper
PIN 3840/50
 Orthoptera - Permelcanidae
Meselcana vologodica n. sp.24 Gorochov 2013 grasshopper
PIN 3840/654, 1211
 Orthoptera - Deinotitanidae
Monstrotitan monstruosa n. gen. n. sp.24
Monstrotitan monstruosa n. gen. n. sp.24 Gorochov 2013 grasshopper
PIN 3840/504, 503, 1218, 1222, 1225, 415, 655, 660, 661, 678, 1209-10, 1214-7, 1219-21, 1223
 Paoliida - Anthracoptilidae
Strephoptilus borealis n. gen. n. sp.28, Strephoptilus longus n. sp.28
Strephoptilus borealis n. gen. n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1651, 1653, 1656
Strephoptilus longus n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1652, 1654, 1655
 Plecoptera - Eustheniidae
Boreoperlidium borealis n. gen. n. sp.31
Boreoperlidium borealis n. gen. n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/604, 1609
 Plecoptera - Euxenoperlidae
Euxenoperlidae indet.31 Riek 1976 stonefly
PIN 3840/603
 Plecoptera - Palaeonemouridae
Palaeonemourisca diluta n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/638, 650, 658
Palaeonemourisca formalis n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/637
Votaknemoura admiranda n. gen. n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1607
Palaeotaeniopteryx mira n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1612
Palaeotaeniopteryx constricta n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/515
Palaeotaeniopteryx dejecta n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/636
Palaeotaeniopteryx succida n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/606, 599
Palaeotaeniopteryx triternata n. sp. Sinitshenkova and Aristov 2010 stonefly
PIN 3840/347
Palaeotaeniopteryx fixa n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/605
Palaeotaeniopteryx adjecta n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1611, 490
Palaeotaeniopteryx amissa n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1608
Palaeotaeniopteryx laeta n. sp. Sinitshenkova and Aristov 2010 stonefly
PIN 3840/344; 489, 631 in Sinitshenkova, 2013
Palaeonemoura amica n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1604
Palaeonemoura issadensis n. sp. Sinitshenkova and Aristov 2010 stonefly
PIN 3840/346; 630 in Sinitshenkova, 2013
Palaeonemoura antrorsa n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/495
 Plecoptera -
Mirumoperla multinerva n. gen. n. sp.31
Mirumoperla multinerva n. gen. n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1614
 Plecoptera - Palaeoperlidae
Properla issadensis n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/425
Kargaloperla furcata n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1606
Kargaloperla decipiens n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/1605
 Plecoptera - Tshekardoperlidae
Issadoperla permiana n. gen. n. sp.31
Issadoperla permiana n. gen. n. sp.31 Sinitshenkova 2013 stonefly
PIN 3840/497
 Dicondylia -
Kitshuga ryzhkovae n. gen. n. sp.4
Kitshuga ryzhkovae n. gen. n. sp.4 Aristov 2012 winged insect
PIN 3840/542
 Dicondylia - Phyloblattidae
Phyloblattidae indet.37 Schneider 1983 winged insect
PIN 3840/859
 Blattodea - Mutoviidae
Mutovia intercalaria n. gen. n. sp.36
Mutovia intercalaria n. gen. n. sp.36 Vršanský and Aristov 2012 cockroach
PIN 3840/57
 Dicondylia -
Permofossilis commasticatus n. gen. n. sp.5 Aristov 2013 winged insect
PIN 3840/683, 1236
 Dicondylia - Permophasmatidae
Isadyphasma suchonae n. sp.24 Gorochov 2013 winged insect
PIN 3840/1228
Isadyphasma bashkuevi n. gen. n. sp.24 Gorochov 2013 winged insect
PIN 3840/507, 505, 1227, 1229
Isadyphasma deminutum n. sp.24 Gorochov 2013 winged insect
PIN 3840/1231
 Embioptera - Alexarasniidae
Alexarasniidae indet.23 Gorochov 2011 webspinners
PIN 3840/1232
Alexarasnia limbata n. sp.10 Aristov 2017 webspinners
PIN 3840/3113
Alexarasnia rossica n. gen. n. sp.23 Gorochov 2011 webspinners
PIN 3840/63
 Grylloblattodea - Tunguskapteridae
"Isadia bipectinata n. gen. n. sp." = Issadiella bipectinata3
"Isadia bipectinata n. gen. n. sp." = Issadiella bipectinata3 Aristov 2009 ice crawler
PIN 3840/301
 Grylloblattodea - Liomopteridae
Liomopterites suhonensis n. sp.3 Aristov 2009 ice crawler
PIN 3840/96
Falseshurabia transitoria n. gen. n. sp.7 Aristov 2013 ice crawler
PIN 3840/1240
Sylvaella semicolorata n. sp.7 Aristov 2013 ice crawler
PIN 3840/1246
Liomofrater circumcisus n. gen. n. sp.7 Aristov 2013 ice crawler
PIN 3840/1347
 Grylloblattodea -
Kenguronympha lenta n. gen. n. sp.7, Issadonympha oculea n. gen. n. sp.7, Sukhonoptera insolita n. gen. n. sp.3
Kenguronympha lenta n. gen. n. sp.7 Aristov 2013 ice crawler
PIN 3840/1647
Issadonympha oculea n. gen. n. sp.7 Aristov 2013 ice crawler
PIN 3840/553
Sukhonoptera insolita n. gen. n. sp.3 Aristov 2009 ice crawler
PIN 3840/51
 Cnemidolestodea - Sylvabestiidae
"Mezenalicula conjuncta n. sp." = Ivkinus conjunctus14
"Mezenalicula conjuncta n. sp." = Ivkinus conjunctus14 Aristov and Storozhenko 2011 winged insect
PIN 3840/1243
 Cnemidolestodea - Issatermidae
Issatermes parvus n. gen. n. sp.11
Issatermes parvus n. gen. n. sp.11 Aristov 2018 winged insect
PIN 3840/3114
 Caloneurodea - Caloneuridae
Isadistica issada n. gen. n. sp.28, Isadistica longa n. sp.28
Isadistica issada n. gen. n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1701
Isadistica longa n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/684
 Miomoptera - Palaeomanteidae
? Tridelopterum indebitum n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1029
 Hypoperlida - Hypoperlidae
Idelopsocus mutovinus n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1650
 Dicondylia - Permembiidae
Issapaloptera infracta n. gen. n. sp.28, Neembia ampla n. gen. n. sp.28
Issapaloptera infracta n. gen. n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1667
Neembia ampla n. gen. n. sp.28 Rasnitsyn and Aristov 2013 winged insect
PIN 3840/1670
 Mecoptera - Permotanyderidae
Rudimentanus rieki n. gen. n. sp.18, Chimaeropteron antiquum n. gen. n. sp.18
Rudimentanus rieki n. gen. n. sp.18 Bashkuev 2022 scorpionfly
PIN 3840/2592, 1010C, 2593
Chimaeropteron antiquum n. gen. n. sp.18 Bashkuev 2022 scorpionfly
PIN 3840/2594
 Mecoptera - Mesopsychidae
Permopsyche sp.15 Bashkuev 2011 scorpionfly
PIN 3840/1556
Permopsyche issadensis n. gen. n. sp.15 Bashkuev 2011 scorpionfly
Permopsyche rasnitsyni n. sp.15 Bashkuev 2011 scorpionfly
PIN 3840/1351,733,1363,1371
 Mecoptera - Nedubroviidae
Nedubrovia shcherbakovi n. gen. n. sp.16
Nedubrovia shcherbakovi n. gen. n. sp.16 Bashkuev 2011 scorpionfly
 Mecoptera - Permochoristidae
Tatarakara variomaculata n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/1427 (holotype)
Permeca pygmaea n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/420, 1419, 1420
Permeca media n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/588, 752, 1418
Mesochorista generalis n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/1421, 79, 83, 407, 509, 755, 758, 1423-1426, 1493, 786, 1422
Neudolbenus giganteus n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/1391
Neudolbenus kopylovi n. gen. n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/745, 241, 1390
Agetopanorpa danili n. sp.17 Bashkuev 2013 scorpionfly
PIN 3840/1527, 709, 1532
 Trichoptera - Cladochoristidae
Cladochorista issadica n. sp.35 Sukatsheva and Aristov 2013 caddisfly
PIN 3840/192
Cladochorista ? issadica35 Sukatsheva and Aristov 2013 caddisfly
PIN 3840/1284
 Raphidioptera - Nanosialidae
Hymega rasnitsyni n. gen. n. sp.29, Nanosialis bashkuevi n. sp.29, Nanosialis ponomarenkoi n. gen. n. sp.29, Lydasialis micheneri n. gen. n. sp.29, Raphisialis martynovi n. gen. n. sp.29
Hymega rasnitsyni n. gen. n. sp.29 Shcherbakov 2013 snakefly
PIN 3840/2600
Nanosialis bashkuevi n. sp.29 Shcherbakov 2013 snakefly
PIN 3840/2633
Nanosialis ponomarenkoi n. gen. n. sp.29 Shcherbakov 2013 snakefly
PIN 3840/2603A, 2604A
Lydasialis micheneri n. gen. n. sp.29 Shcherbakov 2013 snakefly
PIN 3840/2602, 2601
Raphisialis martynovi n. gen. n. sp.29 Shcherbakov 2013 snakefly
PIN 3840/2009
 Dicondylia -
Coleoptera indet.19 Linnaeus 1758 beetle
PIN 3840/1280, 2027, 2042 (beetle hindwings)
 Coleoptera - Permocupedidae
Protocupoides esini n. sp.26 Ponomarenko 2013 beetle
PIN 3840/28, 437, 1282
 Coleoptera - Taldycupedidae
Taldycupedidae indet.19, "Simmondsia permiana n. sp." = Proterocupes permiana26, Taldycupes cellulosus n. sp.26
Taldycupedidae indet.19 Rohdendorf 1961 beetle
PIN 3840/688
"Simmondsia permiana n. sp." = Proterocupes permiana26 Ponomarenko 2013 beetle
PIN 3840/1602, 688, 1279, 1703
Taldycupes cellulosus n. sp.26 Ponomarenko 2013 beetle
PIN 3840/438, 436, 686
 Coleoptera - Asiocoleidae
Bicoleus laticella n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1203
Tetracoleus tricoleoides n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1202, 1275
 Coleoptera - Schizocoleidae
Karakanocoleus mutovinensis n. sp.26 Ponomarenko 2013 beetle
PIN 3840/439
Uskatocoleus sukhonensis n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1273, 1206, 1269, 1270, 1601
Uskatocoleus convexus n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1205, 1204, 1207
Uskatocoleus micron n. sp.26 Ponomarenko 2013 beetle
PIN 3840/417
Pseudochrysomelites medialis n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1671
Pseudochrysomelites sphenoidalis n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1268
Pseudochrysomelites longus n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1208
Pseudochrysomelites bashkuevi n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1276
Erunakicupes grossus n. sp.26 Ponomarenko 2013 beetle
PIN 3840/440
Schizocoleus depressus n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1278
 Coleoptera - Rhombocoleidae
Rossocoleus aristovi n. sp.26 Ponomarenko 2013 beetle
PIN 3840/1274
 Dicondylia - Kennedyidae
Kennedya suchonensis n. sp.20 Felker 2021 winged insect
PIN 3840/2605, 1306, 1699, 2607, 5003
 Dicondylia - Voltzialestidae
Prototerskeja dubia n. gen. n. sp.21, Permosticta elegans n. gen. n. sp.21, Permosticta parva n. sp.21
Prototerskeja dubia n. gen. n. sp.21 Felker 2023 winged insect
PIN 3840/5001, 1690, 1691, 2095, 5011
Permosticta elegans n. gen. n. sp.21 Felker 2023 winged insect
PIN 3840/1685, 2619, 5004, 1687, 1689, 1692, 2608, 2609, 2611, 2613, 2614, 2617, 2622, 5002, 5006, 5008
Permosticta parva n. sp.21 Felker 2023 winged insect
PIN 3840/314, 1688
 Dicondylia - Progoneuridae
Issadoneura marilevorum n. gen. n. sp.21
Issadoneura marilevorum n. gen. n. sp.21 Felker 2023 winged insect
PIN 3840/5509, 1686, 1696, 2093, 2610, 2612
 Dicondylia - Permagrionidae
Triadolestes mutovkensis n. sp.21 Felker 2023 winged insect
PIN 3840/1040
 Ephemerida - Misthodotidae
Misthodotes dubius n. sp.30 Sinitshenkova 2013 mayfly
PIN 3840/1706
Misthodotes tshernovae n. sp.34 Sinitshenkova and Vassilenko 2012 mayfly
PIN 3840/1534
 Ephemerida - Protereismatidae
Ponalex maximus n. gen. n. sp.32, Alexandrinia ipsa n. sp.30, Alexandrinia gigantea n. gen. n. sp.34, Alexandrinia vitta n. sp.30
Ponalex maximus n. gen. n. sp.32 Sinitshenkova and Aristov 2012 mayfly
PIN 3840/875
Alexandrinia ipsa n. sp.30 Sinitshenkova 2013 mayfly
PIN 3840/1708
Alexandrinia gigantea n. gen. n. sp.34 Sinitshenkova and Vassilenko 2012 mayfly
PIN 3840/874, 876
Alexandrinia vitta n. sp.30 Sinitshenkova 2013 mayfly
PIN 3840/1707
Arachnida
 Scorpiones -
Scorpiones indet.22 Latreille 1817 scorpion
PIN 3840/986, 987, 2083
Ostracoda
  -
Myriapoda
 Scolopendromorpha -
Permocryptops shelleyi n. gen. n. sp.25
Permocryptops shelleyi n. gen. n. sp.25 Khramov et al. 2018 centipede
PIN 3840/3165
Osteichthyes
 Eurynotoidiformes - Eurynotoidiidae
Eurynotoidiidae indet.1 Minikh and Minikh 1990 ray-finned fish
Isadia aristoviensis1 Minikh 1990 ray-finned fish
Isadia suchonensis1 Minikh 1986 ray-finned fish
 Elonichthyiformes - Varialepididae
Varialepis ex gr. stanislavi1 Minikh 2009 ray-finned fish
 Elonichthyiformes - Eigiliidae
Strelnia certa1 Minich 1995 ray-finned fish
Amphibia
 Temnospondyli - Dvinosauridae
Dvinosaurus aff. primus1 Amalitzki 1921 tetrapod
Osteichthyes
 Therapsida - Gorgonopidae
Gorgonopidae indet.1 Lydekker 1890 therapsid
Reptilia
 Procolophonomorpha - Pareiasauridae
Pareiasauridae indet.1 Lydekker 1889 parareptile
Osteichthyes
 Seymouriamorpha - Kotlassidae
Microphon arcanus1 Bulanov 2003 tetrapod
 Tetrapoda - Chroniosuchidae
Chroniosaurus levis1 Golubev 1998 tetrapod
Actinopteri
 Scanilepiformes - Evenkiidae
Toyemia sp.1 Minikh 1990
Toyemia tverdochlebovi1 Minikh 1990
Osteichthyes
  - Discordichthyidae
Mutovinia stella1 Minich 1992
Geryonichthys sp.1 Minikh 1998