Soyana (PIN collection 2334) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified, fine-grained sandstone

• Probable brackish lagoon with some access to sea
• "fine granulous sandstone" Martynov, 1932

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by M. B. Edemskij in 1929; reposited in the PIN

Primary reference: A. V. Martynov. 1932. New Permian Palaeoptera with the discussion of some problems of their evolution. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 1:1-44 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104993: authorized by Matthew Clapham, entered by Matthew Clapham on 13.02.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Reculida - Ivapteridae
Ivaptera sharovi1
Ivaptera sharovi1 Aristov 2009 winged insect
PIN 2334/209, 234
 Dicondylia - Permaeschnidae
"Permaeschna dolloi n. gen. n. sp." = Permaeschna dolloi5, "Permaeschna proxima n. sp." = Permaeschna dolloi5
"Permaeschna dolloi n. gen. n. sp." = Permaeschna dolloi5 Martynov 1931 winged insect
PIN 2334/3(2)
"Permaeschna proxima n. sp." = Permaeschna dolloi5 Martynov 1931 winged insect
PIN 2334/4
 Meganisoptera - Meganeuridae
Arctotypus sinuatus n. gen. n. sp.
Arctotypus sinuatus n. gen. n. sp. Martynov 1932 griffinfly
PIN 2334/5
 Ephemeroptera - Protereismidae
Protereismidae indet.
Protereismidae indet. Handlirsch 1919 mayfly
PIN 2334/5
 Protelytroptera -
Arctocoleus ivensis n. gen. n. sp.7
Arctocoleus ivensis n. gen. n. sp.7 Martynov 1932 winged insect
PIN 2334/55
 Glosselytrodea - Jurinidae
Jurina marginata11
Jurina marginata11 Martynov 1938 winged insect
PIN 2334/348, 349
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis6
Kamopanorpa rotundipennis6 Martynov 1932 winged insect
PIN 2334/8
 Mecoptera - Permopanorpidae
"Permopanorpa tenuis n. sp." = Permopanorpa martynovi6, "Permopanorpa angustata n. sp." = Martynopanorpa angustata6
"Permopanorpa tenuis n. sp." = Permopanorpa martynovi6 Willmann 1978 scorpionfly
PIN 2334/178
"Permopanorpa angustata n. sp." = Martynopanorpa angustata6 Martynov 1932 scorpionfly
PIN 2334/350, 255
 Mecoptera - Permochoristidae
"? Permochorista dubiosa n. sp." = Petromantis dubiosa6, Petromantis robusta6, "Permochorista maculipennis n. sp." = Petromantis maculipennis6, "Petromantis punctata" = Petromantis rossica6, "Petromantis borealis" = Petromantis rossica6, "Petrochorista dubia n. sp." = Petromantis dubia6, "Agetochorista ? fasciata" = Agetopanorpa fasciata6, "Agetochorista dubia n. sp." = Agetopanorpa dubia6, "Agetochorista ornata n. sp." = Agetopanorpa ornata6, "Agetochorista splendida" = Agetopanorpa ornata6
"? Permochorista dubiosa n. sp." = Petromantis dubiosa6 Martynov 1932 scorpionfly
PIN 2334/35
Petromantis robusta6 Martynov 1932 scorpionfly
PIN 2334/21(167)
"Permochorista maculipennis n. sp." = Petromantis maculipennis6 Martynov 1932 scorpionfly
PIN 2334/172, second specimen is PIN 2334/47; var. major n. var.
"Petromantis punctata" = Petromantis rossica6 Handlirsch 1904 scorpionfly
PIN 2334/187
"Petromantis borealis" = Petromantis rossica6 Handlirsch 1904 scorpionfly
PIN 2334/16, 67, 59, 36, 18, 40, 176, 11, 193, 34, 189, 193, 75, 71, 189, 60, 37
"Petrochorista dubia n. sp." = Petromantis dubia6 Martynov 1932 scorpionfly
PIN 2334/66; 2334/188 in Martynova, 1948
"Agetochorista ? fasciata" = Agetopanorpa fasciata6 Martynov 1932 scorpionfly
PIN 2334/10
"Agetochorista dubia n. sp." = Agetopanorpa dubia6 Martynov 1932 scorpionfly
PIN 2334/63
"Agetochorista ornata n. sp." = Agetopanorpa ornata6 Martynov 1932 scorpionfly
PIN 2334/38
"Agetochorista splendida" = Agetopanorpa ornata6 Martynov 1932 scorpionfly
PIN 2334/19
 Mecoptera - Parachoristidae
Parachorista opposita n. sp.6
Parachorista opposita n. sp.6 Martynov 1932 scorpionfly
PIN 2334/171
 Dicondylia - Permithonidae
Permosisyra latipennis n. gen. n. sp.7, Permegalomus maculipennis n. sp.7, "Permithonopsis proxima n. sp." = Permithonopsis ivensis7
Permosisyra latipennis n. gen. n. sp.7 Martynov 1932 winged insect
PIN 2334/43
Permegalomus maculipennis n. sp.7 Martynov 1932 winged insect
PIN 2334/68
"Permithonopsis proxima n. sp." = Permithonopsis ivensis7 Martynov 1932 winged insect
PIN 2334/56
 Coleoptera - Permosynidae
Permocrossos elongatus n. gen. n. sp.7
Permocrossos elongatus n. gen. n. sp.7 Martynov 1932 beetle
PIN 2334/51
 Hypoperlida - Hypoperlidae
Boreopsocus danksae12
Boreopsocus danksae12 Shcherbakov 1994 winged insect
PIN 2334/395, 400, 407
 Hemiptera - Archescytinidae
Sojanoscytina minor n. sp.8
Sojanoscytina minor n. sp.8 Martynov 1933 true bug
PIN 2334/119
 Hemiptera - Coleoscytidae
Coleoscytodes elytrata9, Coleoscyta rotundata n. gen. n. sp.9
Coleoscytodes elytrata9 Martynov 1935 planthopper
PIN 2334/91
Coleoscyta rotundata n. gen. n. sp.9 Martynov 1935 planthopper
PIN 2334/107
 Hemiptera - Scytinopteridae
Scytinoptera sp.9, "Anomoscyta reducta" = Scytinoptera reducta9, "Scytinoptera ivensis n. sp." = Scytinoptera reducta9
Scytinoptera sp.9 Handlirsch 1904 true bug
PIN 2334/159
"Anomoscyta reducta" = Scytinoptera reducta9 Martynov 1928 true bug
PIN 2334/176; 88 (S. ivensis in Martynov), 124 (both in Becker-Migdisova,1948)
"Scytinoptera ivensis n. sp." = Scytinoptera reducta9 Martynov 1928 true bug
PIN 2334/100, 87
 Hemiptera - Boreoscytidae
"Sojanoscytina latipennis n. sp." = Boreoscyta latipennis4
"Sojanoscytina latipennis n. sp." = Boreoscyta latipennis4 Martynov 1933 true bug
PIN 2334/129
 Hemiptera - Prosbolidae
Sojanoneura sp.9, Sojanoneura bimaculata n. sp.9, Sojanoneura edemskii9, Prosbole "sp 1"9, Prosbole "sp 2"9, Prosbole "sp 3"9, Prosbole ovalis n. sp.9, Prosbole reducta n. sp.9, Prosbole jucunda n. sp.9, Prosbole sojanensis n. sp.9, "Sojanoneura aberrans n. sp." = Permocicadopsis aberrans9, Permocicada "sp 1"9, Permocicada simulatrix n. sp.9, Permocicada nigronervosa3, Permocicada elliptica n. sp.9
Sojanoneura sp.9 Martynov 1928 true bug
PIN 2334/78, 134, 125
Sojanoneura bimaculata n. sp.9 Martynov 1935 true bug
PIN 2334/150
Sojanoneura edemskii9 Martynov 1928 true bug
PIN 2334/89
Prosbole "sp 1"9 Handlirsch 1904 true bug
PIN 2334/155
Prosbole "sp 2"9 Handlirsch 1904 true bug
PIN 2334/137
Prosbole "sp 3"9 Handlirsch 1904 true bug
PIN 2334/94
Prosbole ovalis n. sp.9 Martynov 1935 true bug
PIN 2334/163
Prosbole reducta n. sp.9 Martynov 1935 true bug
PIN 2334/109
Prosbole jucunda n. sp.9 Martynov 1935 true bug
PIN 2334/128
Prosbole sojanensis n. sp.9 Martynov 1935 true bug
PIN 2334/145, 2334
"Sojanoneura aberrans n. sp." = Permocicadopsis aberrans9 Martynov 1935 true bug
PIN 2334/79
Permocicada "sp 1"9 Martynov 1928 true bug
PIN 2334/127
Permocicada simulatrix n. sp.9 Martynov 1935 true bug
PIN 2334/83
Permocicada nigronervosa3 Martynov 1928 true bug
PIN 2334/111
Permocicada elliptica n. sp.9 Martynov 1935 true bug
PIN 2334/122
 Hemiptera - Prosbolopseidae
Permopibrocha ramosa n. gen. n. sp.9, Prosbolopsis sp.9, "Prosbolopsis ovalis n. gen. n. sp." = Prosbolopsis ovalis9, "Prosbolopsis simplex n. sp." = Prosbolopsis ovalis9
Permopibrocha ramosa n. gen. n. sp.9 Martynov 1935 true bug
PIN 2334/141
Prosbolopsis sp.9 Martynov 1935 true bug
PIN 2334/110
"Prosbolopsis ovalis n. gen. n. sp." = Prosbolopsis ovalis9 Martynov 1935 true bug
PIN 2334/130
"Prosbolopsis simplex n. sp." = Prosbolopsis ovalis9 Martynov 1935 true bug
PIN 2334/95, 99, 96
 Paoliida - Paoliidae
Sojanidelia nana13, Sojanidelia maculosa13
Sojanidelia nana13 Storozhenko 1992 winged insect
PIN 2334/287
Sojanidelia maculosa13 Storozhenko 1992 winged insect
PIN 2334/277(8)
 Caloneurodea - Caloneuridae
"Euthygramma aberrans n. sp." = Euthygramma parallellum10
"Euthygramma aberrans n. sp." = Euthygramma parallellum10 Martynov 1928 winged insect
PIN 2334/313
 Palaeodictyoptera - Eukulojidae
"Kuloja paurovenosa n. sp." = Eukuloja cubitalis
"Kuloja paurovenosa n. sp." = Eukuloja cubitalis Martynov 1932 winged insect
PIN 2334/1
 Permothemistida - Permothemistidae
"Palaeothemis libelluloides n. gen. n. sp." = Permothemis libelluloides
"Palaeothemis libelluloides n. gen. n. sp." = Permothemis libelluloides Martynov 1932 winged insect
PIN 2334/7
 Eoblattida - Idelinellidae
Permostriga sp.2
Permostriga sp.2 Novokshonov 1999 insect
PIN 2334/411 (different from Bekker-Migdisova specimen?)
1Aristov 2008; 2Aristov and Rasnitsyn 2012; 3Becker-Migdisova 1940; 4Becker-Migdisova 1949; 5Martynov 1931; 6Martynov 1932; 7Martynov 1932; 8Martynov 1933; 9Martynov 1935; 10Martynov 1938; 11Martynov 1938; 12Shcherbakov 1994; 13Storozhenko 1992