Soyana (PIN collection 2334) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified, fine-grained sandstone

• Probable brackish lagoon with some access to sea
• "fine granulous sandstone" Martynov, 1932

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by M. B. Edemskij in 1929; reposited in the PIN

Primary reference: A. V. Martynov. 1932. New Permian Palaeoptera with the discussion of some problems of their evolution. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 1:1-44 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 104993: authorized by Matthew Clapham, entered by Matthew Clapham on 13.02.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Protelytroptera -
Arctocoleus ivensis n. gen. n. sp.7
Arctocoleus ivensis n. gen. n. sp.7 Martynov 1932 winged insect
PIN 2334/55
 Hypoperlida - Hypoperlidae
Boreopsocus danksae12 Shcherbakov 1994 winged insect
PIN 2334/395, 400, 407
 Hemiptera - Coleoscytidae
Coleoscyta rotundata n. gen. n. sp.9 Martynov 1935 planthopper
PIN 2334/107
Coleoscytodes elytrata9 Martynov 1935 planthopper
PIN 2334/91
 Hemiptera - Scytinopteridae
Scytinoptera sp.9, "Anomoscyta reducta" = Scytinoptera reducta9, "Scytinoptera ivensis n. sp." = Scytinoptera reducta9
Scytinoptera sp.9 Handlirsch 1904 true bug
PIN 2334/159
"Anomoscyta reducta" = Scytinoptera reducta9 Martynov 1928 true bug
PIN 2334/176; 88 (S. ivensis in Martynov), 124 (both in Becker-Migdisova,1948)
"Scytinoptera ivensis n. sp." = Scytinoptera reducta9 Martynov 1928 true bug
PIN 2334/100, 87
 Hemiptera - Archescytinidae
Sojanoscytina minor n. sp.8 Martynov 1933 true bug
PIN 2334/119
 Hemiptera - Boreoscytidae
"Sojanoscytina latipennis n. sp." = Boreoscyta latipennis4
"Sojanoscytina latipennis n. sp." = Boreoscyta latipennis4 Martynov 1933 true bug
PIN 2334/129
 Hemiptera - Prosbolopseidae
Permopibrocha ramosa n. gen. n. sp.9, Prosbolopsis sp.9, "Prosbolopsis ovalis n. gen. n. sp." = Prosbolopsis ovalis9, "Prosbolopsis simplex n. sp." = Prosbolopsis ovalis9
Permopibrocha ramosa n. gen. n. sp.9 Martynov 1935 true bug
PIN 2334/141
Prosbolopsis sp.9 Martynov 1935 true bug
PIN 2334/110
"Prosbolopsis ovalis n. gen. n. sp." = Prosbolopsis ovalis9 Martynov 1935 true bug
PIN 2334/130
"Prosbolopsis simplex n. sp." = Prosbolopsis ovalis9 Martynov 1935 true bug
PIN 2334/95, 99, 96
 Hemiptera - Prosbolidae
Permocicada "sp 1"9 Martynov 1928 true bug
PIN 2334/127
Permocicada simulatrix n. sp.9 Martynov 1935 true bug
PIN 2334/83
Permocicada nigronervosa3 Martynov 1928 true bug
PIN 2334/111
Permocicada elliptica n. sp.9 Martynov 1935 true bug
PIN 2334/122
"Sojanoneura aberrans n. sp." = Permocicadopsis aberrans9 Martynov 1935 true bug
PIN 2334/79
Sojanoneura sp.9 Martynov 1928 true bug
PIN 2334/78, 134, 125
Sojanoneura edemskii9 Martynov 1928 true bug
PIN 2334/89
Sojanoneura bimaculata n. sp.9 Martynov 1935 true bug
PIN 2334/150
Prosbole "sp 1"9 Handlirsch 1904 true bug
PIN 2334/155
Prosbole "sp 2"9 Handlirsch 1904 true bug
PIN 2334/137
Prosbole "sp 3"9 Handlirsch 1904 true bug
PIN 2334/94
Prosbole sojanensis n. sp.9 Martynov 1935 true bug
PIN 2334/145, 2334
Prosbole jucunda n. sp.9 Martynov 1935 true bug
PIN 2334/128
Prosbole ovalis n. sp.9 Martynov 1935 true bug
PIN 2334/163
Prosbole reducta n. sp.9 Martynov 1935 true bug
PIN 2334/109
 Caloneurodea - Caloneuridae
"Euthygramma aberrans n. sp." = Euthygramma parallellum10
"Euthygramma aberrans n. sp." = Euthygramma parallellum10 Martynov 1928 winged insect
PIN 2334/313
 Paoliida - Paoliidae
Sojanidelia maculosa13 Storozhenko 1992 winged insect
PIN 2334/277(8)
Sojanidelia nana13 Storozhenko 1992 winged insect
PIN 2334/287
 Glosselytrodea - Jurinidae
Jurina marginata11 Martynov 1938 winged insect
PIN 2334/348, 349
 Coleoptera - Permosynidae
Permocrossos elongatus n. gen. n. sp.7
Permocrossos elongatus n. gen. n. sp.7 Martynov 1932 beetle
PIN 2334/51
 Mecoptera - Parachoristidae
Parachorista opposita n. sp.6 Martynov 1932 scorpionfly
PIN 2334/171
 Mecoptera - Permochoristidae
Petromantis robusta6, "Petromantis borealis" = Petromantis rossica6, "Petromantis punctata" = Petromantis rossica6, "Permochorista maculipennis n. sp." = Petromantis maculipennis6, "Petrochorista dubia n. sp." = Petromantis dubia6, "? Permochorista dubiosa n. sp." = Petromantis dubiosa6, "Agetochorista ornata n. sp." = Agetopanorpa ornata6, "Agetochorista splendida" = Agetopanorpa ornata6, "Agetochorista dubia n. sp." = Agetopanorpa dubia6, "Agetochorista ? fasciata" = Agetopanorpa fasciata6
Petromantis robusta6 Martynov 1932 scorpionfly
PIN 2334/21(167)
"Petromantis borealis" = Petromantis rossica6 Handlirsch 1904 scorpionfly
PIN 2334/16, 67, 59, 36, 18, 40, 176, 11, 193, 34, 189, 193, 75, 71, 189, 60, 37
"Petromantis punctata" = Petromantis rossica6 Handlirsch 1904 scorpionfly
PIN 2334/187
"Permochorista maculipennis n. sp." = Petromantis maculipennis6 Martynov 1932 scorpionfly
PIN 2334/172, second specimen is PIN 2334/47; var. major n. var.
"Petrochorista dubia n. sp." = Petromantis dubia6 Martynov 1932 scorpionfly
PIN 2334/66; 2334/188 in Martynova, 1948
"? Permochorista dubiosa n. sp." = Petromantis dubiosa6 Martynov 1932 scorpionfly
PIN 2334/35
"Agetochorista ornata n. sp." = Agetopanorpa ornata6 Martynov 1932 scorpionfly
PIN 2334/38
"Agetochorista splendida" = Agetopanorpa ornata6 Martynov 1932 scorpionfly
PIN 2334/19
"Agetochorista dubia n. sp." = Agetopanorpa dubia6 Martynov 1932 scorpionfly
PIN 2334/63
"Agetochorista ? fasciata" = Agetopanorpa fasciata6 Martynov 1932 scorpionfly
PIN 2334/10
 Mecoptera - Permopanorpidae
"Permopanorpa angustata n. sp." = Martynopanorpa angustata6, "Permopanorpa tenuis n. sp." = Permopanorpa martynovi6
"Permopanorpa angustata n. sp." = Martynopanorpa angustata6 Martynov 1932 scorpionfly
PIN 2334/350, 255
"Permopanorpa tenuis n. sp." = Permopanorpa martynovi6 Willmann 1978 scorpionfly
PIN 2334/178
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis6 Martynov 1932 winged insect
PIN 2334/8
 Dicondylia - Permithonidae
"Permithonopsis proxima n. sp." = Permithonopsis ivensis7, Permegalomus maculipennis n. sp.7, Permosisyra latipennis n. gen. n. sp.7
"Permithonopsis proxima n. sp." = Permithonopsis ivensis7 Martynov 1932 winged insect
PIN 2334/56
Permegalomus maculipennis n. sp.7 Martynov 1932 winged insect
PIN 2334/68
Permosisyra latipennis n. gen. n. sp.7 Martynov 1932 winged insect
PIN 2334/43
 Palaeodictyoptera - Eukulojidae
"Kuloja paurovenosa n. sp." = Eukuloja cubitalis
"Kuloja paurovenosa n. sp." = Eukuloja cubitalis Martynov 1932 winged insect
PIN 2334/1
 Palaeodictyoptera - Permothemistidae
"Palaeothemis libelluloides n. gen. n. sp." = Permothemis libelluloides
"Palaeothemis libelluloides n. gen. n. sp." = Permothemis libelluloides Martynov 1932 winged insect
PIN 2334/7
 Reculida - Ivapteridae
Ivaptera sharovi1 Aristov 2009 winged insect
PIN 2334/209, 234
 Ephemeroptera - Protereismidae
Protereismidae indet. Handlirsch 1919 mayfly
PIN 2334/5
 Dicondylia - Permaeschnidae
"Permaeschna dolloi n. gen. n. sp." = Permaeschna dolloi5, "Permaeschna proxima n. sp." = Permaeschna dolloi5
"Permaeschna dolloi n. gen. n. sp." = Permaeschna dolloi5 Martynov 1931 winged insect
PIN 2334/3(2)
"Permaeschna proxima n. sp." = Permaeschna dolloi5 Martynov 1931 winged insect
PIN 2334/4
 Meganisoptera - Meganeuridae
Arctotypus sinuatus n. gen. n. sp.
Arctotypus sinuatus n. gen. n. sp. Martynov 1932 griffinfly
PIN 2334/5
 Eoblattida - Idelinellidae
Permostriga sp.2 Novokshonov 1999 insect
PIN 2334/411 (different from Bekker-Migdisova specimen?)