PBDB Collection

Kityak (PIN collection 1366) (Permian of Russian Federation)

Where: Kirov, Russian Federation (56.3° N, 50.7° E: paleocoordinates 28.5° N, 43.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Belebeevo Formation, Roadian (272.3 - 268.8 Ma)

Environment/lithology: lacustrine; lithified siliciclastic sediments

Size class: macrofossils

Preservation: mold/impression

Primary reference: D. S. Aristov. 2008. New Grylloblattida (Insecta) from the Middle and Upper Permian of Russia. Far Eastern Entomologist 188:1-7 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 105058: authorized by Matthew Clapham, entered by Matthew Clapham on 14.02.2011, edited by Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Eoblattida - Megakhosaridae	Microkhosara fragilis n. gen. n. sp.¹⁶ Microkhosara fragilis n. gen. n. sp.¹⁶ Storozhenko 1993 insect PIN 1366/172
Eoblattida - Atactophlebiidae	Malmyzhia kazanica n. gen. n. sp.⁴ Malmyzhia kazanica n. gen. n. sp.⁴ Aristov 2015 insect PIN 1366/246
Dicondylia - Permagrionidae	Sushkinia angulata n. sp.⁸, Solikamptilon aequus n. sp.⁸ Sushkinia angulata n. sp.⁸ Felker 2023 winged insect PIN 1366/1, 10, 7(11) Solikamptilon aequus n. sp.⁸ Felker 2023 winged insect PIN 1366/697, 2
Dicondylia - Progoneuridae	Progoneura kityakensis n. sp.⁷ Progoneura kityakensis n. sp.⁷ Felker 2020 winged insect PIN 1366/385
Dicondylia - Permosialidae	Permosialis sp.⁵, Permosialis ualentovae n. sp.¹¹, Permosialis belebei n. sp.⁵ Permosialis sp.⁵ Martynov 1928 winged insect PIN 1366/382 (formerly paratype hindwing of P. ualentovae) Permosialis ualentovae n. sp.¹¹ Novokshonov and Zhuzhgova 2004 winged insect PIN 1366/349 (not paratypes 379, 463, 382) Permosialis belebei n. sp.⁵ Aristov and Rasnitsyn 2022 winged insect PIN 1366/379, 463 (formerly paratypes of Permosialis ualentovae)
Grylloblattodea - Liomopteridae	Protomia proteus n. gen. n. sp.¹, Liomopterella kitiakensis n. sp.¹, Liomopterella acuminata n. sp.¹, Liomopterella kirovensis n. sp. Protomia proteus n. gen. n. sp.¹ Aristov 2004 ice crawler PIN 1366/226, 124, 153, 155, 156, 161 Liomopterella kitiakensis n. sp.¹ Aristov 2004 ice crawler PIN 1366/121, 259 Liomopterella acuminata n. sp.¹ Aristov 2004 ice crawler PIN 1366/260, 177 Liomopterella kirovensis n. sp. Aristov 2008 ice crawler PIN 1366/124, 184, 250, 266
Grylloblattodea - Lemmatophoridae	Artinska novella n. sp.³ Artinska novella n. sp.³ Aristov 2015 ice crawler PIN 1366/172a
Orthoptera - Permelcanidae	"Permelcana vjushkovi n. sp." = Permelcana sojanense¹⁴ "Permelcana vjushkovi n. sp." = Permelcana sojanense¹⁴ Sharov 1962 grasshopper PIN 1366/290
Orthoptera - Pruvostitidae	Kitjakoedischia pruvostitoides n. gen. n. sp.⁹ Kitjakoedischia pruvostitoides n. gen. n. sp.⁹ Gorochov 1987 grasshopper PIN 1366/285
Cnemidolestodea - Protembiidae	Kirovopteron luteus n. gen. n. sp.³ Kirovopteron luteus n. gen. n. sp.³ Aristov 2015 winged insect PIN 1366/654
Cnemidolestodea - Sylvabestiidae	Kityakia attrita n. gen. n. sp.² Kityakia attrita n. gen. n. sp.² Aristov 2014 winged insect PIN 1366/695, 498, 568
Paoliida - Paoliidae	Idelina mala n. sp.³, Stenaropodites magna n. sp.¹⁵ Idelina mala n. sp.³ Aristov 2015 winged insect PIN 1366/122 Stenaropodites magna n. sp.¹⁵ Storozhenko 1992 winged insect PIN 1366/239
Dicondylia - Spiloblattinidae	Kitjakia longifurcata n. gen. n. sp.¹⁷ Kitjakia longifurcata n. gen. n. sp.¹⁷ Vishnyakova 1993 winged insect PIN 1366/19
Dicondylia -	"Belebey mutilus n. gen. n. sp." = Belebeus mutilus⁴ "Belebey mutilus n. gen. n. sp." = Belebeus mutilus⁴ Aristov 2015 winged insect PIN 1366/404
Mecoptera - Permochoristidae	Malmyzha incompta n. gen. n. sp.¹⁰, "Agetochorista obsoleta n. sp." = Agetopanorpa obsoleta¹⁰, Parageta kitjakensis n. sp.¹⁰ Malmyzha incompta n. gen. n. sp.¹⁰ Novokshonov 1995 scorpionfly PIN 1366/414, 401, 403, 430, 437, 443, 449, 415 "Agetochorista obsoleta n. sp." = Agetopanorpa obsoleta¹⁰ Novokshonov 1995 scorpionfly PIN 1366/393 Parageta kitjakensis n. sp.¹⁰ Novokshonov 1995 scorpionfly PIN 1366/441
Coleoptera - Permocupedidae	"Kaltanicupes major n. sp." = Uralocupes major¹², Kaltanicupes acutus n. sp.¹², Kaltanicupes kitjakensis n. sp.¹² "Kaltanicupes major n. sp." = Uralocupes major¹² Ponomarenko 1963 beetle PIN 1366/376 Kaltanicupes acutus n. sp.¹² Ponomarenko 1963 beetle PIN 1366/377 Kaltanicupes kitjakensis n. sp.¹² Ponomarenko 1963 beetle PIN 1366/378
Coleoptera - Rhombocoleidae	Rossocoleus altus n. sp.¹³ Rossocoleus altus n. sp.¹³ Ponomarenko 2004 beetle PIN 1366/371, 372
Reculida - Geinitziidae	Permovalia abortiva n. gen. n. sp.³ Permovalia abortiva n. gen. n. sp.³ Aristov 2015 winged insect PIN 1366/287, 330
Reculida - Chaulioditidae	Miralioma monstrosa n. gen. n. sp.¹ Miralioma monstrosa n. gen. n. sp.¹ Aristov 2004 winged insect PIN 1366/152
Arachnida
Araneae - Arthrolycosidae	Arthrolycosa sp.⁶ Arthrolycosa sp.⁶ Harger 1874 spider PIN 1366/490

¹Aristov 2004; ²Aristov 2014; ³Aristov 2015; ⁴Aristov 2015; ⁵Aristov and Rasnitsyn 2022; ⁶Eskov and Selden 2005; ⁷Felker 2020; ⁸Felker 2023; ⁹Gorochov 1987; ¹⁰Novokshonov 1995; ¹¹Novokshonov and Zhuzhgova 2004; ¹²Ponomarenko 1963; ¹³Ponomarenko 2004; ¹⁴Sharov 1962; ¹⁵Storozhenko 1992; ¹⁶Storozhenko 1993; ¹⁷Vishnyakova 1993