PBDB Collection

Vishnevaya Balka creek, Karagan, Stavropol (PIN collection 254) (Miocene of Russian Federation)

Where: Russian Federation (45.1° N, 42.3° E: paleocoordinates 45.5° N, 41.3° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karagan Formation, Middle Miocene (16.0 - 11.6 Ma)

Environment/lithology: lacustrine; lithified, carbonaceous shale

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by B.F. Kaspiev in 1938; reposited in the PIN

Primary reference: Y. P. Nekrutenko. 1965. Tertiary nymphalid butterflies and some phylogenetic aspects of systematic lepidopterology. Journal of Research on the Lepidoptera 4(3):149-158 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 105571: authorized by Matthew Clapham, entered by Jered Karr on 22.02.2011, edited by Matthew Clapham and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Odonata - Sieblosiidae	Miostenolestes zherikhini n. gen. n. sp.⁹, Stenolestes fasciata n. sp.⁹, Paraoligolestes stavropolensis n. sp.⁹ Miostenolestes zherikhini n. gen. n. sp.⁹ Nel et al. 2005 damselfly PIN 254/2169 Stenolestes fasciata n. sp.⁹ Nel et al. 2005 damselfly PIN 254/3192 Paraoligolestes stavropolensis n. sp.⁹ Nel et al. 2005 damselfly PIN 254/2171, 2202
Odonata - Aeshnidae	Aeshna stavropolensis n. sp.⁸ Aeshna stavropolensis n. sp.⁸ Nel et al. 2005 hawker dragonfly PIN 254/2148
Odonata - Libellulidae	Pantalinae indet.⁸, Sympetrum sp.⁸ Pantalinae indet.⁸ common skimmer PIN 254/2163, 2198 Sympetrum sp.⁸ Newman 1833 common skimmer PIN 254/2141
Trichoptera - Limnephilidae	Limnephilus kaspievi¹² Limnephilus kaspievi¹² Martynova 1939 caddisfly PIN 254/256, 651, 813, 958, 1010, 1212, 1262, 1264, 1268, 1269, 1271, 1272, 1273, 1275, 1276, 1277, 1278, 1281, 1313, 1359, 1955, 1351, 3041, 3186; presumably includes specimens from Temnolesskaya
Lepidoptera - Nymphalidae	"Pyrameis fossilis n. sp." = Vanessa fossilis, "Vanessa karaganica n. sp." = Aglais karaganica "Pyrameis fossilis n. sp." = Vanessa fossilis Nekrutenko 1965 brush-footed butterfly PIN 254/2753 "Vanessa karaganica n. sp." = Aglais karaganica Nekrutenko 1965 brush-footed butterfly PIN 254/2936a
Lepidoptera -	"Xyleutites miocenicus n. gen. n. sp." = Noctuites miocenicus⁴ "Xyleutites miocenicus n. gen. n. sp." = Noctuites miocenicus⁴ Kozhanchikov 1957 moth PIN 254/182
Coleoptera - Dytiscidae	Copelatus spp.¹⁰, Copelatus fossilis n. sp.¹⁰, Copelatus stavropolitanus n. sp.¹⁰, Copelatus ponomarenkoi n. sp.¹⁰, Hydroporus lapideus n. sp.¹⁰, Methles caucasicus n. sp.¹⁰, ? Graphoderus mirabilis n. sp.¹⁰, Cybister fractus n. sp.¹⁰, Colymbetes sp.¹⁰, Colymbetes miocaenicus n. sp.¹⁰ Copelatus spp.¹⁰ Erichson 1832 predaceous diving beetle PIN 254/1654, 1711 Copelatus fossilis n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1623 Copelatus stavropolitanus n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1743 Copelatus ponomarenkoi n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1734, 1733 Hydroporus lapideus n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1715 Methles caucasicus n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1676 ? Graphoderus mirabilis n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1607 Cybister fractus n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/1626 Colymbetes sp.¹⁰ Clairville 1806 predaceous diving beetle PIN 254/1758 Colymbetes miocaenicus n. sp.¹⁰ Říha 1974 predaceous diving beetle PIN 254/2673
Coleoptera - Nitidulidae	"Lasiodactylus caucasicus n. sp." = Phenolia (Lasiodites) caucasicus² "Lasiodactylus caucasicus n. sp." = Phenolia (Lasiodites) caucasicus² Kirejtshuk 1990 sap beetle PIN 254/1794
Hemiptera - Derbidae	Mysidioides migdisovae n. sp.¹ Mysidioides migdisovae n. sp.¹ Emeljanov 2002 planthopper PIN 254/239
Hemiptera - Aphididae	Eutrichosiphum europaeum¹³ Eutrichosiphum europaeum¹³ Wegierek and Peñalver 2002 aphid PIN 254/2940
Orthoptera - Oecanthidae	Paroecanthus caucasicus n. gen. n. sp.¹¹ Paroecanthus caucasicus n. gen. n. sp.¹¹ Sharov 1968 tree cricket PIN 254/2577
Diptera - Tephritidae	"? Pseudacidia clotho n. sp." = Myoleja clotho³ "? Pseudacidia clotho n. sp." = Myoleja clotho³ Korneyev 1982 fruit fly PIN 254/724 (PIN 252/724 is error?)
Diptera - Limoniidae	Toxorhina (Ceratocheilus) caucasiensis n. sp.⁷, Helius stavropolensis n. sp.⁶, Helius verticillis n. sp.⁶, Helius miocenicus n. sp.⁶ Toxorhina (Ceratocheilus) caucasiensis n. sp.⁷ Krzemiński and Freiwald 1991 crane fly PIN 254/671, 275 Helius stavropolensis n. sp.⁶ Krzemiński 2002 crane fly PIN 254/231, 278, 764, 288, 650 Helius verticillis n. sp.⁶ Krzemiński 2002 crane fly PIN 254/2507, 816, 932 Helius miocenicus n. sp.⁶ Krzemiński 2002 crane fly PIN 254/833, 170a, 170b, 135, 322, 337, 533, 590, 598, 612, 821, 1079, 1442, 1580, 2301
Diptera - Trichoceridae	Trichocera sp.⁵, Paracladura caucasiana n. sp.⁵ Trichocera sp.⁵ Meigen 1803 winter crane fly PIN 254/264 Paracladura caucasiana n. sp.⁵ Krzemińska et al. 2009 winter crane fly PIN 254/697

¹Emeljanov 2002; ²Kirejtshuk and Ponomarenko 1990; ³Korneyev 1982; ⁴Kozhanchikov 1957; ⁵Krzemińska et al. 2009; ⁶Krzemiński 2002; ⁷Krzemiński and Freiwald 1991; ⁸Nel et al. 2005; ⁹Nel et al. 2005; ¹⁰Říha 1974; ¹¹Sharov 1968; ¹²Sukatsheva et al. 2018; ¹³Wegierek and Peñalver 2002