Karatau-Mikhailovka (PIN collection 2554) (Jurassic to of Kazakhstan)

Where: Chimkent, Kazakhstan (42.9° N, 70.0° E: paleocoordinates 41.6° N, 73.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karabastau Formation, Callovian to Callovian (166.1 - 157.3 Ma)

Environment/lithology: lacustrine - large; lithified siltstone

• "dark shales" Grimaldi & Engel, 2005

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Reposited in the PIN

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 105970: authorized by Matthew Clapham, entered by Matthew Clapham on 28.02.2011, edited by Andrew Spencer-Lee, Jered Karr and Lindy Edwards

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Malmopsyllidae
Malmopsylla karatavica n. gen. n. sp.6
Malmopsylla karatavica n. gen. n. sp.6 Becker-Migdisova 1985 jumping plant louse
PIN 2554/369
 Hemiptera - Protopsyllidiidae
Poljanka "sp. 1"50, "Cicadellopsis kukalovae n. sp." = Poljanka kukalovae5, "Cicadellopsis sharovi n. sp." = Poljanka sharovi6, Carpenterella grombtchevskayae n. sp.6, Aphidulum heckeri6, Karatavopsyllidium auliense n. sp.6
Poljanka "sp. 1"50 Klimaszewski 1995 true bug
PIN 2554/371
"Cicadellopsis kukalovae n. sp." = Poljanka kukalovae5 Becker-Migdisova 1968 true bug
PIN 2554/384
"Cicadellopsis sharovi n. sp." = Poljanka sharovi6 Becker-Migdisova 1985 true bug
PIN 2554/1182
Carpenterella grombtchevskayae n. sp.6 Becker-Migdisova 1985 true bug
PIN 2554/1174
Aphidulum heckeri6 Becker-Migdisova 1985 true bug
PIN 2554/1233
Karatavopsyllidium auliense n. sp.6 Becker-Migdisova 1985 true bug
PIN 2554/389
 Hemiptera - Scaphocoridae
Scaphocoris notatus n. gen. n. sp.35
Scaphocoris notatus n. gen. n. sp.35 Popov 1968 true bug
PIN 2554/406
 Thysanoptera - Liassothripidae
Liassothrips crassipes43 Martynov 1927 thrips
PIN 2554/1077, 1082
 Psocodea - Electrentomidae
Paramesopsocus adibi3 Azar et al. 2008 book louse
PIN 2554/388
 Blattaria - Skokidae
Skok svaba n. gen. n. sp.47
Skok svaba n. gen. n. sp.47 Vršanský 2007 jumping cockroach
PIN 2554/44, 77
 Blattaria - Caloblattinidae
Decomposita triocella n. gen. n. sp.48, Paleovia praecarnia n. gen. n. sp.48
Decomposita triocella n. gen. n. sp.48 Vršanský 2008 cockroach
PIN 2554/36, 30
Paleovia praecarnia n. gen. n. sp.48 Vršanský 2008 cockroach
PIN 2554/33
 Dermaptera - Semenoviolidae
Semenovioloides capitatus46 Vishnyakova 1980 earwig
PIN 2554/335
 Dermaptera - Turanodermatidae
Turanoderma sepultum46 Vishnyakova 1980 earwig
PIN 2554/648
 Dermaptera - Protodiplatyidae
Asiodiplatys speciosus46 Vishnyakova 1980 earwig
PIN 2554/657
 Phasmatodea - Susumaniidae
Phasmomimoides lineatus n. gen. n. sp.
Phasmomimoides lineatus n. gen. n. sp. Sharov 1968 stick insect
PIN 2554/318
 Phasmatodea - Aerophasmidae
Jurophasma mistshenkoi n. gen. n. sp.11
Jurophasma mistshenkoi n. gen. n. sp.11 Gorochov 1993 stick insect
PIN 2554/326
 Orthoptera - Protogryllidae
Karataogryllus gryllotalpiformis n. gen. n. sp. Sharov 1968 grasshopper
PIN 2554/283
 Orthoptera - Prophalangopsidae
Aboilus aulietus n. gen. n. sp. Sharov 1968 hump-winged cricket
PIN 2554/236
Aboilus transversalis n. sp.12 Gorochov 1996 hump-winged cricket
PIN 2554/270
Aboilus tigris12 Gorochov 1996 hump-winged cricket
PIN 2554/267
 Orthoptera - Tuphellidae
Paracyrtophyllites undulatus Sharov 1968 grasshopper
PIN 2554/287
 Orthoptera - Elcanidae
"Elcana karatavica n. sp." = Probaisselcana karatavica, "Elcana minuta" = Panorpidium minuta
"Elcana karatavica n. sp." = Probaisselcana karatavica Sharov 1968 grasshopper
PIN 2554/280
"Elcana minuta" = Panorpidium minuta Sharov 1968 grasshopper
PIN 2554/295
 Hymenoptera - Ephialtitidae
Symphytopterus nigridorsatus n. sp.39 Rasnitsyn 1975 wasp
PIN 2554/1296
Symphytopterus nigricornis39 Rasnitsyn 1975 wasp
PIN 2554/1272
Trigonalopterus brachycerus39 Rasnitsyn 1975 wasp
PIN 2554/1297, 1283
 Hymenoptera - Mesoserphidae
Karataoserphinus minor n. gen. n. sp.41, Mesoserphus karatavicus n. gen. n. sp.18
Karataoserphinus minor n. gen. n. sp.41 Rasnitsyn 1994 wasp
PIN 2554/1201
Mesoserphus karatavicus n. gen. n. sp.18 Kozlov 1968 wasp
PIN 2554/1281
 Hymenoptera - Praeaulacidae
"Aulacogaster ater n. gen. n. sp." = Aulacogastrinus ater38, Praeaulacus ventricosus n. sp.38, Praeaulacus cubocephalus n. sp.38
"Aulacogaster ater n. gen. n. sp." = Aulacogastrinus ater38 Rasnitsyn 1972 wasp
PIN 2554/1287
Praeaulacus ventricosus n. sp.38 Rasnitsyn 1972 wasp
PIN 2554/1271
Praeaulacus cubocephalus n. sp.38 Rasnitsyn 1972 wasp
PIN 2554/1284
 Hymenoptera - Megalyridae
Cleistogaster oculata n. sp.39 Rasnitsyn 1975 wasp
PIN 2554/1278
 Hymenoptera - Bethylonymidae
Bethylonymellus oligocerus n. sp.39 Rasnitsyn 1975 wasp
PIN 2554/1270
 Hymenoptera - Brachycleistogastridae
Microcleistogaster dolichura n. sp.39 Rasnitsyn 1975 wasp
PIN 2554/1297
Brachycleistogaster nana n. sp.39 Rasnitsyn 1975 wasp
PIN 2554/1279
Brachycleistogaster tredecatoma n. sp.39 Rasnitsyn 1975 wasp
PIN 2554/1292
 Hymenoptera - Anaxyelidae
? Syntexyela nana n. sp.37, "Syntexyela magna n. sp." = Urosyntexis magna37
? Syntexyela nana n. sp.37 Rasnitsyn 1968 winged insect
PIN 2554/1300
"Syntexyela magna n. sp." = Urosyntexis magna37 Rasnitsyn 1968 winged insect
PIN 2554/1301
 Hymenoptera - Siricidae
Pararchexyela macroptera n. gen. n. sp.37, Megura magnifica n. gen. n. sp.37
Pararchexyela macroptera n. gen. n. sp.37 Rasnitsyn 1968 horntail
PIN 2554/1299
Megura magnifica n. gen. n. sp.37 Rasnitsyn 1968 horntail
PIN 2554/1298
 Hymenoptera - Sepulcidae
Parapamphilius confusus n. gen. n. sp.37
Parapamphilius confusus n. gen. n. sp.37 Rasnitsyn 1968 stem sawfly
PIN 2554/1303
 Hymenoptera - Paroryssidae
Microryssus crassipes n. sp.37 Rasnitsyn 1968 wood wasp
PIN 2554/1305
Microryssus antennatus n. sp.37 Rasnitsyn 1968 wood wasp
PIN 2554/1304
 Coleoptera - Buprestidae
Metabuprestium ovale n. sp.1 Alexeev 1995 jewel beetle
PIN 2554/448
 Coleoptera - Cerophytidae
Necromera aff. admiranda7, "Idiomerus inflatus n. gen. n. sp." = Necromera inflata10
Necromera aff. admiranda7 Chang and Kirejtshuk 2011 beetle
PIN 2554/460
"Idiomerus inflatus n. gen. n. sp." = Necromera inflata10 Dolin 1980 beetle
PIN 2554/694
 Coleoptera - Elateridae
Cryptocardius mirabilis n. gen. n. sp.10 Dolin 1980 click beetle
PIN 2554/649
Litholacon derumpens n. gen. n. sp.10 Dolin 1980 click beetle
PIN 2554/699
Pseudocardiophorites fragilis n. gen. n. sp.9 Dolin 1976 click beetle
PIN 2554/688
Codemus synaptoides10 Dolin 1980 click beetle
PIN 2554/680, 682
Codemus quadricolis10 Dolin 1980 click beetle
PIN 2554/687, 704
Codemus alatus10 Dolin 1980 click beetle
PIN 2554/654
Codemus teres10 Dolin 1980 click beetle
PIN 2554/695
Hypnomorphus imperspicuus n. sp.8 Dolin 1975 click beetle
PIN 2554/678
Hypnomorphus dubius n. sp.8 Dolin 1975 click beetle
PIN 2554/651
Hypnomorphus distinctus n. sp.8 Dolin 1975 click beetle
PIN 2554/653, 652
Hypnomorphus inventus8 Dolin 1975 click beetle
PIN 2554/697
Hypnomorphus minutus8 Dolin 1975 click beetle
PIN 2554/700
Hypnomorphus aemulus n. sp.8 Dolin 1975 click beetle
PIN 2554/692, 690
Hypnomorphus rohdendorfi8 Dolin 1975 click beetle
PIN 2554/676
Lithoptychus handlirschi n. sp.10 Dolin 1980 click beetle
PIN 2554/696
Parahypnomorphus jurassicus n. gen. n. sp.10 Dolin 1980 click beetle
PIN 2554/703
Negastrioides globicollis n. sp.10 Dolin 1980 click beetle
PIN 2554/705
Platyelater reflexicollis10 Dolin 1980 click beetle
PIN 2554/683
Lithomerus cockerelli10 Dolin 1980 click beetle
PIN 2554/685
Desmatus beckeri8 Dolin 1975 click beetle
PIN 2554/689
Desmatus affinis n. sp.8 Dolin 1975 click beetle
PIN 2554/672
 Coleoptera - Staphylinidae
Tunicopterus sigara n. gen. n. sp.44 Tikhomirova 1968 rove beetle
PIN 2554/669(665)
Abolescus glabratus44 Tikhomirova 1968 rove beetle
PIN 2554/656
Mesoxytelus mandibularis n. sp.44 Tikhomirova 1968 rove beetle
PIN 2554/664
Globoides microps n. sp.44 Tikhomirova 1968 rove beetle
PIN 2554/661
Archodromus brachypterus n. sp.44 Tikhomirova 1968 rove beetle
PIN 2554/650
Archodromus comptus44 Tikhomirova 1968 rove beetle
PIN 2554/668
Mesotachinus major44 Tikhomirova 1968 rove beetle
PIN 2554/675,680
 Coleoptera - Trogossitidae
Juralithinus gracilidorsum n. gen. n. sp.17
Juralithinus gracilidorsum n. gen. n. sp.17 Kirejtshuk and Ponomarenko 1990 bark gnawing beetle
PIN 2554/624
 Coleoptera - Anthribidae
Protoscelis nitidicornis22 Medvedev 1968 fungus weevil
PIN 2554/735
Protoscelis jurassica22 Medvedev 1968 fungus weevil
PIN 2554/743
Protoscelis longicornis22 Medvedev 1968 fungus weevil
PIN 2554/736, 737, 738
 Coleoptera - Nemonychidae
Brenthorrhinus mirabilis n. gen. n. sp.2, "Distenorrhinus (Astenorrhinus) elongatus" = Astenorrhinus elongatus14, "Procurculio fortipes" = Procurculio fortipes14, "Eccoptothorax latipennis n. gen. n. sp." = Procurculio fortipes2, "Distenorrhinus (Distenorrhinus) rotundicollis" = Distenorrhinus (Distenorrhinus) pallidirostris14, "Distenorrhinus antennatus n. sp." = Distenorrhinus (Distenorrhinus) antennatus2, "Distenorrhinus (Distenorrhinus) arnoldii n. sp." = Distenorrhinus (Distenorrhinus) angulatus14, Ampliceps dubius13, "Scelocamptus tenuirostris n. gen. n. sp." = Ampliceps tenuirostris2, "Ampliceps tenuirostris" = Ampliceps tenuirostris13, Ampliceps dentitibia13, Ampliceps furcitibia13, "Oxycorynoides similis n. gen. n. sp." = Oxycorynoides (Oxycorynoides) similis2, Oxycorynoides (Pseudoxycorynoides) brevipes13, Archaeorrhynchus acutirostris n. gen. n. sp.2, "Probelus curvispinus n. gen. n. sp." = Probelus (Probelus) curvispinus2, Belonotaris (Belonotaris) punctatissimus13, Belonotaris (Arnoldibelus) rasnitsyni13
Brenthorrhinus mirabilis n. gen. n. sp.2 Arnoldi 1977 pine flower snout beetle
PIN 2554/708
"Distenorrhinus (Astenorrhinus) elongatus" = Astenorrhinus elongatus14 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2554/733
"Procurculio fortipes" = Procurculio fortipes14 Arnoldi 1977 pine flower snout beetle
PIN 2554/731
"Eccoptothorax latipennis n. gen. n. sp." = Procurculio fortipes2 Arnoldi 1977 pine flower snout beetle
PIN 2554/720
"Distenorrhinus (Distenorrhinus) rotundicollis" = Distenorrhinus (Distenorrhinus) pallidirostris14 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2554/1210
"Distenorrhinus antennatus n. sp." = Distenorrhinus (Distenorrhinus) antennatus2 Arnoldi 1977 pine flower snout beetle
PIN 2554/721
"Distenorrhinus (Distenorrhinus) arnoldii n. sp." = Distenorrhinus (Distenorrhinus) angulatus14 Arnoldi 1977 pine flower snout beetle
PIN 2554/701(725), 565
Ampliceps dubius13 Arnoldi 1977 pine flower snout beetle
PIN 2554/725
"Scelocamptus tenuirostris n. gen. n. sp." = Ampliceps tenuirostris2 Arnoldi 1977 pine flower snout beetle
PIN 2554/732
"Ampliceps tenuirostris" = Ampliceps tenuirostris13 Arnoldi 1977 pine flower snout beetle
PIN 2554/733
Ampliceps dentitibia13 Arnoldi 1977 pine flower snout beetle
PIN 2554/565
Ampliceps furcitibia13 Arnoldi 1977 pine flower snout beetle
PIN 2554/710, 711, 719, 722, 734
"Oxycorynoides similis n. gen. n. sp." = Oxycorynoides (Oxycorynoides) similis2 Arnoldi 1977 pine flower snout beetle
PIN 2554/713; 723, 726, 728, 730 in Gratshev & Legalov 2014
Oxycorynoides (Pseudoxycorynoides) brevipes13 Arnoldi 1977 pine flower snout beetle
PIN 2554/718, 729
Archaeorrhynchus acutirostris n. gen. n. sp.2 Arnoldi 1977 pine flower snout beetle
PIN 2554/715
"Probelus curvispinus n. gen. n. sp." = Probelus (Probelus) curvispinus2 Arnoldi 1977 pine flower snout beetle
PIN 2554/79
Belonotaris (Belonotaris) punctatissimus13 Arnoldi 1977 pine flower snout beetle
PIN 2554/717
Belonotaris (Arnoldibelus) rasnitsyni13 Legalov 2014 pine flower snout beetle
PIN 2554/714
 Coleoptera - Coptoclavidae
Charonoscapha grossa n. gen. n. sp.2, "Necronectes gigas n. sp." = Timarchopsis gigas2, Exedia plana n. gen. n. sp.2
Charonoscapha grossa n. gen. n. sp.2 Ponomarenko 1977 beetle
PIN 2554/446
"Necronectes gigas n. sp." = Timarchopsis gigas2 Ponomarenko 1977 beetle
PIN 2554/477
Exedia plana n. gen. n. sp.2 Ponomarenko 1977 beetle
PIN 2554/478
 Coleoptera - Trachypachidae
Karadromeus latus2 Ponomarenko 1977 false ground beetle
PIN 2554/724
Psacodromeus gutta n. gen. n. sp.2 Ponomarenko 1977 false ground beetle
PIN 2554/456
 Coleoptera - Carabidae
Ovrabites ovalis2 Ponomarenko 1977 ground beetle
PIN 2554/526
Protorabus magnus n. sp.2 Ponomarenko 1977 ground beetle
PIN 2554/447
 Coleoptera - Schizophoridae
Xyphosternum minor34 Ponomarenko 1968 beetle
PIN 2554/444
Tersus karatavicus34 Ponomarenko 1968 beetle
PIN 2554/441,442
 Coleoptera - Obrieniidae
Kararhynchus occiduus21 Zherikhin and Gratshev 1993 beetle
PIN 2554/502
 Coleoptera - Cupedidae
Notocupes nigrimonticola34 Ponomarenko 1968 reticulated beetle
PIN 2554/430
Notocupes lapidarius34 Ponomarenko 1968 reticulated beetle
PIN 2554/433,437
Omma jurassicum34 Ponomarenko 1968 reticulated beetle
PIN 2554/432,436
 Mecoptera - Bittacidae
Orobittacus antiquus30 Novokshonov 1993 hangingfly
PIN 2554/1094
Prohylobittacus sorosi31 Novokshonov 1993 hangingfly
PIN 2554/924
 Mecoptera - Orthophlebiidae
"Orthophlebia grandis" = Gigaphlebia grandis49
"Orthophlebia grandis" = Gigaphlebia grandis49 Martynov 1927 scorpionfly
PIN 2554/870 (not 810)
 Mecoptera - Aneuretopsychidae
? Aneuretopsyche minima n. sp.42 Rasnitsyn and Kozlov 1990 scorpionfly
PIN 2554/1324
 Diptera - Empididae
Homalocnemimus abjugatus28 Mostovski 1999 dagger fly
PIN 2554/1131
 Diptera - Archisargidae
Mesosolva hennigi n. sp.26 Mostovski 1996 fly
PIN 2554/1159+1160
 Diptera - Rhagionidae
Protorhagio karataviensis29 Mostovski 2008 snipe fly
PIN 2554/941
Protorhagio ruptus n. sp.29 Mostovski 2008 snipe fly
PIN 2554/943
 Diptera - Nemestrinidae
Protonemestrius rasnitsyni n. sp.27 Mostovski 1998 tangle-veined fly
PIN 2554/908
 Diptera - Rhagionemestriidae
Rhagionemestrius rapidus n. gen. n. sp.45
Rhagionemestrius rapidus n. gen. n. sp.45 Ussatchov 1968 fly
PIN 2554/959(1132)
 Diptera - Procramptonomyiidae
Procramptonomyia kovalevi24 Lukashevich 2012 fly
PIN 2554/1029, 1085, 1168, 1017, 1106
 Diptera - Elliidae
Polyanka minuta19 Krzemińska et al. 1993 fly
PIN 2554/1121, 1207
 Diptera - Trichoceridae
Zherikhinina karatavica20 Krzemińska et al. 2009 winter crane fly
PIN 2554/1049
 Diptera - Tanyderidae
Praemacrochile ansorgei n. sp.25 Lukashevich and Krzemiński 2009 primitive crane fly
PIN 2554/934
Praemacrochile kaluginae n. sp.25 Lukashevich and Krzemiński 2009 primitive crane fly
PIN 2554/958(1095)
 Diptera - Ptychopteridae
"Proptychoptera aucta n. sp." = Eoptychoptera aucta23 Lukashevich 1993 phantom crane fly
PIN 2554/1047
Eoptychopterina karatavica23 Lukashevich 1993 phantom crane fly
PIN 2554/942
Proptychopterina gracilis23 Lukashevich 1993 phantom crane fly
PIN 2554/970(972), 1064, 879, 925
 Neuroptera - Grammolingiidae
Grammolingiidae indet.15 Ren 2002 lacewing
PIN 2554/166
 Neuroptera -
Microsmylus foliformis n. gen. n. sp.33
Microsmylus foliformis n. gen. n. sp.33 Panfilov 1980 lacewing
PIN 2554/784
 Neuroptera - Kalligrammatidae
Kalligramma sharovi n. sp.32, Kalligrammina areolata n. gen. n. sp.33
Kalligramma sharovi n. sp.32 Panfilov 1968 lacewing
PIN 2554/788
Kalligrammina areolata n. gen. n. sp.33 Panfilov 1980 lacewing
PIN 2554/777
 Neuroptera - Osmylidae
Arbusella sp.16 Khramov 2014 lance lacewing
PIN 2554/798
Jurakempynus arcanus n. sp.16 Khramov 2014 lance lacewing
PIN 2554/775
Jurosmylus parvulus n. sp.16 Khramov 2014 lance lacewing
PIN 2554/765
 Neuroptera - Berothidae
"Mesithone gracilis n. sp." = Berothone gracilis33
"Mesithone gracilis n. sp." = Berothone gracilis33 Panfilov 1980 beaded lacewing
PIN 2554/807
 Dicondylia - Tarsophlebiidae
Turanophlebia martynovi n. gen. n. sp.36
Turanophlebia martynovi n. gen. n. sp.36 Pritykina 1968 winged insect
PIN 2554/210
 Odonata -
Cymatophlebiella euryptera n. gen. n. sp.4, "Protolindenia aktassica n. sp." = Kazakhophlebiella aktassica36
Cymatophlebiella euryptera n. gen. n. sp.4 Pritykina 1968 true dragonfly
PIN 2554/211
"Protolindenia aktassica n. sp." = Kazakhophlebiella aktassica36 Pritykina 1968 true dragonfly
PIN 2554/218
 Odonata - Erichschmidtiidae
Erichschmidtia nigrimontana n. gen. n. sp.36
Erichschmidtia nigrimontana n. gen. n. sp.36 Pritykina 1968 dragonfly
PIN 2554/214
 Odonata - Euthemistidae
Euthemis cellulata n. sp.36 Pritykina 1968 dragonfly
PIN 2554/219
 Odonata - Asiopteridae
Turanopteron minor n. gen. n. sp.36
Turanopteron minor n. gen. n. sp.36 Pritykina 1968 dragonfly
PIN 2554/215
 Eoblattida - Blattogryllidae
Blattogryllus karatavicus n. gen. n. sp.40
Blattogryllus karatavicus n. gen. n. sp.40 Rasnitsyn 1976 insect
PIN 2554/225, 226, 227