Madygen, PIN collection 2555, Dzhailoucho area (Triassic of Kyrgyzstan)

Where: Osh, Kyrgyzstan (40.1° N, 70.2° E: paleocoordinates 36.5° N, 60.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Madygen Formation, Ladinian (242.0 - 237.0 Ma)

• Aristov (2020): The age of the Madygen locality is usually reported as the Ladinian or Carnian of the Middle or Upper Triassic (Shcherbakov, 2008). This concept emerged, because I.A. Dobruskina (1995) did not discriminate between Ladinian and Carnian floras. Our data show that only three of the six families found at Madygen are known from younger sediments (Fig. 1), and therefore we can confirm the assumption of the Ladinian age of the Madygen locality (A.G. Ponomarenko, personal communication, 2018).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - small; lithified siliciclastic sediments

• Oxbow lake; Madygen basin; tectonically-induced intermontane basin. Complex facies pattern consistent with active tectonics in upland region.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106088: authorized by Matthew Clapham, entered by Matthew Clapham on 02.03.2011, edited by Lindy Edwards, Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Eoblattida - Megakhosaridae
Megakhosarodes paulivenosus n. sp.36 Storozhenko 1993 insect
PIN 2555/699
Megablattogryllus magister33 Storozhenko 1990 insect
PIN 2555/691
Megablattogryllus austerus33 Storozhenko 1990 insect
PIN 2555/727
Metakhosara sharovi36 Storozhenko 1993 insect
PIN 2555/757, 779
"Microblattogryllus variabilis n. gen. n. sp." = Protoblattogryllus variabilis33 Storozhenko 1990 insect
PIN 2555/744, 674, 770, 777, 830, 833
Protoblattogryllus asiaticus n. sp.33 Storozhenko 1990 insect
PIN 2555/786, 848
Mesoblattogryllus intermedius n. gen. n. sp.33 Storozhenko 1990 insect
PIN 2555/984
 Eoblattida - Blattogryllidae
Baharellinus pectinatus n. sp.35 Storozhenko 1992 insect
PIN 2555/967
Baharellinus dimidiatus n. gen. n. sp.35 Storozhenko 1992 insect
PIN 2555/759, 807, 935
Dorniella primitiva n. sp.35 Storozhenko 1992 insect
PIN 2555/709(931)
Costatoviblatta aenigmatosa35 Storozhenko 1992 insect
PIN 2555/672, 680, 735, 739, 758, 762, 767, 771, 809, 844, 856, 858, 875, 881, 890, 901, 910, 915, 922, 933, 998, 999
Costatoviblatta similis n. sp.1 Aristov 2011 insect
Baharellus madygensis n. sp.35 Storozhenko 1992 insect
PIN 2555/722
 Eoblattida - Mesorthopteridae
Belmophenopterum rasnitsyni n. sp.40 Storozhenko and Aristov 2021 insect
PIN 2555/820
Parastenaropodites longiuscula39 Storozhenko 1996 insect
Parastenaropodites fluxa n. gen. n. sp.39 Storozhenko 1996 insect
PIN 2555/985,789,842,906,911
Sharovites alexanderi3 Aristov and Storozhenko 2013 insect
PIN 2555/729, 2781
Locustoblattina segmentata n. sp.2 Aristov 2017 insect
PIN 2555/2001
Austroidelia asiatica39 Storozhenko 1996 insect
"Parastenaropodites nervosa" = Austroidelia nervosa39 Storozhenko 1996 insect
Mesoidelia ignorata n. gen. n. sp.39 Storozhenko 1996 insect
PIN 2555/676,991
Mesoidelia faceta39 Storozhenko 1996 insect
PIN 2555/686
 Diptera - Psychotipidae
Psychotipa depicta n. sp.31 Shcherbakov 1995 fly
PIN 2555/661
 Diptera - Nadipteridae
Nadiptera pulchella n. sp.31 Lukashevich 1995 fly
PIN 2555/660
 Hymenoptera - Xyelidae
Ferganoxyela destructa n. sp.24, Ferganoxyela sogdiana n. gen. n. sp.24, Oryctoxyela anomala n. gen. n. sp.24, Madygenius primitivus n. sp.24, Triassoxyela orycta n. sp.24
Ferganoxyela destructa n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1753
Ferganoxyela sogdiana n. gen. n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1752
Oryctoxyela anomala n. gen. n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1754
Madygenius primitivus n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1755
Triassoxyela orycta n. sp.24 Rasnitsyn 1969 sawfly
PIN 2555/1757
 Mecoptera - Mesopsychidae
Prochoristella longa15, "Mesopsyche shcherbakovi n. sp." = Turbidapsyche shcherbakovi16, "Mesopsyche gentica n. sp." = Turbidapsyche gentica19, "Mesopsyche justa" = Turbidapsyche justa19
Prochoristella longa15 Novokshonov 1997 scorpionfly
PIN 2555/1956
"Mesopsyche shcherbakovi n. sp." = Turbidapsyche shcherbakovi16 Novokshonov 1997 scorpionfly
PIN 2555/2037
"Mesopsyche gentica n. sp." = Turbidapsyche gentica19 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2555/2013
"Mesopsyche justa" = Turbidapsyche justa19 Novokshonov and Sukacheva 2001 scorpionfly
PIN 2555/1950
 Mecoptera - Permochoristidae
Agetopanorpa consueta n. sp.15 Novokshonov 1997 scorpionfly
PIN 2555/1812
Mesageta ignava n. sp.15 Novokshonov 1997 scorpionfly
PIN 2555/1890, 1867
Mesageta pertrita n. sp.15 Novokshonov 1997 scorpionfly
PIN 2555/1927
Liassochorista ? utilis15 Novokshonov 1997 scorpionfly
PIN 2555/1964
 Mecoptera - Parachoristidae
Triassochorista kirgizica n. sp.17 Novokshonov 1997 scorpionfly
PIN 2555/1835, 2045, 2052
Parachorista asiatica n. sp.17 Novokshonov 1997 scorpionfly
PIN 2555/2017
Parachorista sana n. sp.18 Novokshonov 2001 scorpionfly
PIN 2555/836
Parachorista arguta18 Novokshonov 2001 scorpionfly
PIN 2555/1989, 2009
Parachorista multivena n. sp.17 Novokshonov 1997 scorpionfly
PIN 2555/1941
 Trichoptera - Cladochoristidae
Cladochorista multivenosa n. sp.42 Sukatsheva 1973 caddisfly
PIN 2555/174
 Trichoptera - Philopotamidae
Prophilopotamus asiaticus n. gen. n. sp.42
Prophilopotamus asiaticus n. gen. n. sp.42 Sukatsheva 1973 finger-net caddisfly
PIN 2555/1780
 Coleoptera - Triaplidae
Triaplus laticoxa4 Ponomarenko 1977 beetle
PIN 2555/1720
 Coleoptera - Ommatidae
Lithocupes gigas n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1668
 Coleoptera - Phoroschizidae
Hadeocoleus catachtonius22 Ponomarenko 1969 beetle
PIN 2555/1704
Hadeocoleus pelopius22 Ponomarenko 1969 beetle
PIN 2555/1701, 1719, 1699
Hadeocoleus gigas22 Ponomarenko 1969 beetle
PIN 2555/1698, 1700
Triassocoleus tortulosus22 Ponomarenko 1969 beetle
PIN 2555/1708
Schizophoroides rugosus22 Ponomarenko 1969 beetle
PIN 2555/1710
Catabrycus hoplites n. gen. n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1702
Salebroferus asper n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1707
Salebroferus confragosus n. gen. n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1692
Thnesidius xyphophorus22 Ponomarenko 1969 beetle
PIN 2555/1709
 Coleoptera -
"Notocupes tenuis" = Rhabdocupes tenuis22, Rhabdocupes baculatus n. sp.22, "Notocupes rostratus n. sp." = Rhabdocupes rostratus22, "Notocupes laticella n. sp." = Rhabdocupes laticella22
"Notocupes tenuis" = Rhabdocupes tenuis22 Ponomarenko 1969 beetle
PIN 2555/1683,1685
Rhabdocupes baculatus n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1666
"Notocupes rostratus n. sp." = Rhabdocupes rostratus22 Ponomarenko 1969 beetle
PIN 2555/1664
"Notocupes laticella n. sp." = Rhabdocupes laticella22 Ponomarenko 1969 beetle
PIN 2555/1663, 1662, 1686
 Coleoptera - Triadocupedidae
Cupesia sepulta n. sp.22
Cupesia sepulta n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1682, 1658
 Coleoptera - Ademosynidae
Dolichosyne rostrata n. sp.22 Ponomarenko 1969 beetle
PIN 2555/1735
Ademosyne kirghizica22 Ponomarenko 1969 beetle
PIN 2555/1696
Cephalosyne capitata22 Ponomarenko 1969 beetle
PIN 2555/1732
 Neuroptera - Archeosmylidae
Osmylotriasia superba n. gen. n. sp.13
Osmylotriasia superba n. gen. n. sp.13 Khramov 2024 lacewing
PIN 2555/2096
 Orthoptera - Dzhajloutshellidae
Adzhajloutshella plana n. sp.11 Gorochov 2005 grasshopper
PIN 2555/1440
 Orthoptera - Locustavidae
Locustavus intermedius11, Locustavus deformatus n. sp.11, Locustavus minutus11, Locustavus problematicus n. sp.11, "Ferganopsis lanceolatus n. gen. n. sp." = Locustavus lanceolatus
Locustavus intermedius11 Gorochov 2005 grasshopper
PIN 2555/1393
Locustavus deformatus n. sp.11 Gorochov 2005 grasshopper
PIN 2555/1396
Locustavus minutus11 Gorochov 2005 grasshopper
PIN 2555/1399, 1406, 1417
Locustavus problematicus n. sp.11 Gorochov 2005 grasshopper
PIN 2555/1403
"Ferganopsis lanceolatus n. gen. n. sp." = Locustavus lanceolatus Sharov 1968 grasshopper
PIN 2555/1392
 Orthoptera - Xenopteridae
Axenopterum venosum n. gen. n. sp.10, Pteroferganodes rieki n. gen. n. sp.7, Ferganopterodes reductus n. gen. n. sp., Ferganopterus clarus, Proxenopterum primitivum n. gen. n. sp.10
Axenopterum venosum n. gen. n. sp.10 Gorochov 2005 grasshopper
PIN 2555/1316
Pteroferganodes rieki n. gen. n. sp.7 Gorochov 1989 grasshopper
PIN 2555/2240
Ferganopterodes reductus n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1280
Ferganopterus clarus Sharov 1968 grasshopper
text says 121 imprints but doesn't specify collection
Proxenopterum primitivum n. gen. n. sp.10 Gorochov 2005 grasshopper
PIN 2555/1313, 1307, 1312, 1315
 Orthoptera - Bintoniellidae
"Provitimia ramivenosa n. sp." = Proshiella ramivenosa, Provitimia pectinata n. gen. n. sp.
"Provitimia ramivenosa n. sp." = Proshiella ramivenosa Sharov 1968 grasshopper
PIN 2555/1436
Provitimia pectinata n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1420
 Orthoptera - Mesoedischiidae
Mesoedischia madygenica Sharov 1968 grasshopper
PIN 2555/1250,1256,1249,1251,1252,1253,1254,1255,1257,1326
 Orthoptera - Proparagryllacrididae
Madygenia orientalis Sharov 1968 grasshopper
PIN 2555/1340
Madygenia ovalis6 Gorochov 1987 grasshopper
PIN 2555/1342
"Fergania reducta n. gen. n. sp." = Kashgarlimahmutia reducta Sharov 1968 grasshopper
PIN 2555/1445
Gryllacrimima perfecta Sharov 1968 grasshopper
 Orthoptera - Tuphellidae
Tuphella rohdendorfi n. sp.5 Gorochov 1986 grasshopper
PIN 2555/1081
Tuphella sharovi n. sp.5 Gorochov 1986 grasshopper
PIN 2555/1159
 Orthoptera - Haglidae
Dinohagla corrugata n. gen. n. sp.5, Dulcihagla beybienkoi n. gen. n. sp.5, Eumaraga madygenica n. gen. n. sp.5, "Maraga tshorkuphlebioides n. sp." = Triassaga tshorkuphlebioides5, Voliopinae indet.5, Zavoliopus densus n. gen. n. sp.5, Turkestania deviata n. gen. n. sp., Platyvoliopus maximus n. gen. n. sp.5, Stenovoliopus elongatus n. gen. n. sp.5, Proisfaroptera martynovi, Hagloptera intermedia n. gen. n. sp.5, Protshorkuphlebia similis n. sp.5, Protshorkuphlebia triassica n. gen. n. sp., Microhagla minuta n. gen. n. sp.5, Sonohagla saussurei n. gen. n. sp.5, Sonohagla chopardi n. sp.5, Lyrohagla decipiens n. sp.5, Lyrohagla pravdini n. sp.5, Archihagla tenuis n. sp.5
Dinohagla corrugata n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1137
Dulcihagla beybienkoi n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1150
Eumaraga madygenica n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1085
"Maraga tshorkuphlebioides n. sp." = Triassaga tshorkuphlebioides5 Gorochov 1986 hump-winged cricket
PIN 2555/1099
Voliopinae indet.5 Gorochov 1986 hump-winged cricket
PIN 2555/1079
Zavoliopus densus n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1148
Turkestania deviata n. gen. n. sp. Sharov 1968 hump-winged cricket
PIN 2555/1142
Platyvoliopus maximus n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1168
Stenovoliopus elongatus n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1119
Proisfaroptera martynovi Sharov 1968 hump-winged cricket
PIN 2555/1114
Hagloptera intermedia n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1141
Protshorkuphlebia similis n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1092
Protshorkuphlebia triassica n. gen. n. sp. Sharov 1968 hump-winged cricket
PIN 2555/1088,1097
Microhagla minuta n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1134
Sonohagla saussurei n. gen. n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1102
Sonohagla chopardi n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1118
Lyrohagla decipiens n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1125
Lyrohagla pravdini n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1165
Archihagla tenuis n. sp.5 Gorochov 1986 hump-winged cricket
PIN 2555/1128
 Orthoptera - Paratitanidae
Paratitan modestus n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1479
Paratitan latispeculum n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1478
Paratitan intermedius n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1520
Paratitan venosus n. sp.9 Gorochov 2003 grasshopper
PIN 2555/1456
Paratitan libelluloides n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1530
 Orthoptera - Mesotitanidae
Mesotitanodes tillyardi n. gen. n. sp., "Prototitan similis n. sp." = Mesotitanodes similis, "Nanotitan extentus" = Gigatitan extensus, "Nanotitan magnificus n. gen. n. sp." = Gigatitan magnificus, Gigatitan ovatus n. sp., Gigatitan similis n. sp., "Prototitan primitivus" = Mesotitan primitivus
Mesotitanodes tillyardi n. gen. n. sp. Sharov 1968 grasshopper
PIN 2555/1524,1455
"Prototitan similis n. sp." = Mesotitanodes similis Sharov 1968 grasshopper
PIN 2555/1481,1521
"Nanotitan extentus" = Gigatitan extensus Sharov 1968 grasshopper
PIN 2555/1513
"Nanotitan magnificus n. gen. n. sp." = Gigatitan magnificus Sharov 1968 grasshopper
PIN 2555/1509
Gigatitan ovatus n. sp. Sharov 1968 grasshopper
PIN 2555/1504,1461
Gigatitan similis n. sp. Sharov 1968 grasshopper
PIN 2555/1462
"Prototitan primitivus" = Mesotitan primitivus Sharov 1968 grasshopper
PIN 2555/1473
 Plecoptera - Perlariopseidae
Fritaniopsis remota n. sp.32 Sinitshenkova 1987 stonefly
PIN 2555/1027, 1019
Fritaniopsis brevicaulis n. gen. n. sp.32 Sinitshenkova 1987 stonefly
PIN 2555/1013, 1017
Tritaniella mera n. gen. n. sp.32 Sinitshenkova 1987 stonefly
PIN 2555/1036
Tritaniella synneura32 Sinitshenkova 1987 stonefly
PIN 2555/1024, 1029
Dicronemoura acaulis32 Sinitshenkova 1987 stonefly
PIN 2555/1014, 1016, 1020, 1022, 1028, 1031
 Embioptera - Alexarasniidae
Nestorembia sp.12 Shcherbakov 2015 webspinners
PIN 2555/1873
 Dicondylia - Aeroplanidae
"Paraplana affinis" = Sharovoplana affinis
"Paraplana affinis" = Sharovoplana affinis Sharov 1968 winged insect
PIN 2555/1563
 Dicondylia - Xiphopteridae
Xiphopterum curvatum n. gen. n. sp.
Xiphopterum curvatum n. gen. n. sp. Sharov 1968 winged insect
PIN 2555/1554
 Dicondylia - Prochresmodidae
Prochresmoda minuta n. sp.8 Gorochov 1994 winged insect
PIN 2555/1611
Triassophasma minutissimum n. sp.8 Gorochov 1994 winged insect
PIN 2555/1614
Triassophasma intermedium8 Gorochov 1994 winged insect
PIN 2555/1557, 1616, 1596
"Prochresmoda brevipoda n. sp." = Triassophasma brevipoda Sharov 1968 winged insect
PIN 2555/1594
 Dicondylia - Subioblattidae
Subioblatta madygenica21 Papier and Nel 2001 winged insect
PIN 2555/76; 38, 39, 43, 46, 48, 51, 53, 59, 62, 66, 80, 81, 82, 86, 87, 89, 98, 100, 104 in Hinkelman 2022
 Grylloblattodea -
Madygenidelia conjuncta n. gen. n. sp.38, Pseudoshurabia pallidula n. gen. n. sp.38
Madygenidelia conjuncta n. gen. n. sp.38 Storozhenko 1996 ice crawler
PIN 2555/791
Pseudoshurabia pallidula n. gen. n. sp.38 Storozhenko 1996 ice crawler
PIN 2555/716
 Grylloblattodea - Gorochoviidae
Gorochovia individua n. gen. n. sp.37
Gorochovia individua n. gen. n. sp.37 Storozhenko 1994 ice crawler
PIN 2555/731,763,873,857
 Grylloblattodea - Madygenophlebiidae
Micromadygenophlebia obscura n. gen. n. sp.34 Storozhenko 1992 ice crawler
PIN 2555/826
Madygenophlebia primitiva n. sp.34 Storozhenko 1992 ice crawler
PIN 2555/718
Madygenophlebia bella n. gen. n. sp.34 Storozhenko 1992 ice crawler
PIN 2555/819,748,824,828
 Cnemidolestodea - Necrophasmidae
Ferganamadygenia plicata n. gen. n. sp.41
Ferganamadygenia plicata n. gen. n. sp.41 Storozhenko and Vršanský 1995 winged insect
PIN 2555/717
 Paoliida - Paoliidae
Ideliopsina ornata n. sp.38 Storozhenko 1996 winged insect
PIN 2555/895, 766
Ideliopsina nana38 Storozhenko 1996 winged insect
PIN 2555/687, 738, 846, 992
 Blattaria - Caloblattinidae
Sogdoblatta nana n. sp.43 Vishnyakova 1998 cockroach
PIN 2555/191, 213, 506
Thuringoblatta sogdianensis43 Vishnyakova 1998 cockroach
PIN 2555/135, 140, 142, 147-152, 154, 170
 Hemiptera - Saaloscytinidae
Tingiopsis reticulata28 Becker-Migdisova 1953 true bug
PIN 2555/2213
 Hemiptera - Maguviopseidae
Asiocula lima n. gen.28, Falcarta bella n. gen.28, Phyllotexta latens n. gen.28, Nevicia imitans n. gen.28
Asiocula lima n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2245
Falcarta bella n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2508
Phyllotexta latens n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2751
Nevicia imitans n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2382
 Hemiptera - Palaeontinidae
Papiliontina spectans n. sp.30 Shcherbakov 2022 true bug
PIN 2555/2137
Papiliontina machaon n. sp.30 Shcherbakov 2022 true bug
PIN 2555/2314
Papiliontina dracomima n. gen. n. sp.30 Shcherbakov 2022 true bug
PIN 2555/2136
 Hemiptera - Mesojabloniidae
Fulgobole evansi n. gen. n. sp.28, Scytachile cf. emeljanovi n. gen.28
Fulgobole evansi n. gen. n. sp.28 Shcherbakov 2011 true bug
PIN 2555/2228,2762
Scytachile cf. emeljanovi n. gen.28 Shcherbakov 2011 true bug
PIN 2555/2499
 Hemiptera - Curvicubitidae
Beaconiella pulchra29 Shcherbakov 2021 true bug
PIN 2555/2360, 2368(2386), 2429, 2908, 2320
Beaconiella cincta29 Shcherbakov 2021 true bug
PIN 2555/2192(2866), 3514(3642)
 Hemiptera - Naibiidae
Coccavus supercubitus n. gen. n. sp.27
Coccavus supercubitus n. gen. n. sp.27 Shcherbakov 2007 true bug
PIN 2555/2312, 2392, 2936
 Dicondylia - Permosialidae
Permosialis triassica n. sp.20, "Perloblatta ordinata n. gen. n. sp." = Permonka triassica35, Permonka unica n. sp.25
Permosialis triassica n. sp.20 Novokshonov and Zhuzhgova 2004 winged insect
PIN 2555/2066, 2067, 2068, 3582
"Perloblatta ordinata n. gen. n. sp." = Permonka triassica35 Rasnitsyn 1977 winged insect
PIN 2555/723a
Permonka unica n. sp.25 Rasnitsyn 1977 winged insect
PIN 2555/1060
 Reculida - Geinitziidae
Shurabia anomala n. sp.26 Rasnitsyn 1982 winged insect
PIN 2555/788
Shurabia ferganensis26 Rasnitsyn 1982 winged insect
PIN 2555/787
Shurabia minuta26 Rasnitsyn 1982 winged insect
PIN 2555/792
 Dicondylia - Mitophlebiidae
Mitophlebia enormis n. gen. n. sp.23
Mitophlebia enormis n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/641
 Dicondylia - Triadophlebiidae
Triadophlebia honesta n. sp.23 Pritykina 1981 winged insect
PIN 2555/646
Triadophlebia minuta n. sp.23 Pritykina 1981 winged insect
PIN 2555/635
Triadophlebia madygenica n. sp.23 Pritykina 1981 winged insect
PIN 2555/639
Triadophlebia magna n. sp.23 Pritykina 1981 winged insect
PIN 2555/635, 645
Triadophlebia modica n. sp.23 Pritykina 1981 winged insect
PIN 2555/642
 Dicondylia - Paurophlebiidae
Neritophlebia longa n. sp.23, Paurophlebia lepida n. gen. n. sp.23, Cladophlebia parvula n. gen. n. sp.23, Cladophlebia brevis n. sp.23, Nonymophlebia venosa n. gen. n. sp.23
Neritophlebia longa n. sp.23 Pritykina 1981 winged insect
PIN 2555/614
Paurophlebia lepida n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/624
Cladophlebia parvula n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/617, 654
Cladophlebia brevis n. sp.23 Pritykina 1981 winged insect
PIN 2555/634
Nonymophlebia venosa n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/649, 653
 Dicondylia - Xamenophlebiidae
Xamenophlebia ornata14 Pritykina 1981 winged insect
PIN 2555/2074
 Dicondylia - Zygophlebiidae
Zygophlebiella curta n. gen. n. sp.23, Zygophlebia ramosa n. gen. n. sp.23, Mixophlebia mixta n. gen. n. sp.14
Zygophlebiella curta n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/633, 631
Zygophlebia ramosa n. gen. n. sp.23 Pritykina 1981 winged insect
PIN 2555/619, 656
Mixophlebia mixta n. gen. n. sp.14 Pritykina 1981 winged insect
PIN 2555/625
 Odonata - Triassolestidae
Triassolestodes asiaticus n. gen. n. sp.23
Triassolestodes asiaticus n. gen. n. sp.23 Pritykina 1981 dragonfly
PIN 2555/637
 Dicondylia - Kennedyidae
Kennedya carpenteri n. sp.23 Pritykina 1981 winged insect
PIN 2555/616
 Dicondylia -
Terskeja pumilio n. sp.23, Terskeja tenuis n. sp.23
Terskeja pumilio n. sp.23 Pritykina 1981 winged insect
PIN 2555/630
Terskeja tenuis n. sp.23 Pritykina 1981 winged insect
PIN 2555/632