Madygen, PIN collection 2069, Dzhailoucho area (Triassic of Kyrgyzstan)

Where: Osh, Kyrgyzstan (40.1° N, 70.2° E: paleocoordinates 36.5° N, 60.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Madygen Formation, Ladinian (242.0 - 237.0 Ma)

• Aristov (2020): The age of the Madygen locality is usually reported as the Ladinian or Carnian of the Middle or Upper Triassic (Shcherbakov, 2008). This concept emerged, because I.A. Dobruskina (1995) did not discriminate between Ladinian and Carnian floras. Our data show that only three of the six families found at Madygen are known from younger sediments (Fig. 1), and therefore we can confirm the assumption of the Ladinian age of the Madygen locality (A.G. Ponomarenko, personal communication, 2018).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - small; lithified siliciclastic sediments

• Oxbow lake; Madygen basin; tectonically-induced intermontane basin. Complex facies pattern consistent with active tectonics in upland region.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106090: authorized by Matthew Clapham, entered by Matthew Clapham on 02.03.2011, edited by Jered Karr, Lindy Edwards and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Eoblattida - Mesorthopteridae
Parastenaropodites fluxa n. gen.29 Storozhenko 1996 insect
Parastenaropodites longiuscula29 Storozhenko 1996 insect
Mesoidelia faceta29 Storozhenko 1996 insect
PIN 2069/1485
Austroidelia asiatica29 Storozhenko 1996 insect
"Parastenaropodites nervosa" = Austroidelia nervosa29 Storozhenko 1996 insect
 Eoblattida - Blattogryllidae
Costatoviblatta aenigmatosa26 Storozhenko 1992 insect
PIN 2069/1619, 1627, 1633, 1699, 1736, 1741, 1797, 1799, 1855
 Eoblattida - Megakhosaridae
"Microblattogryllus variabilis" = Protoblattogryllus variabilis24 Storozhenko 1990 insect
PIN 2069/1714
Madygenocephalus micropteron n. gen. n. sp.1 Aristov 2011 insect
PIN 2069/1518
Megablattogryllus pinguis24 Storozhenko 1990 insect
PIN 2069/1403
Megakhosarodes paulivenosus27 Storozhenko 1993 insect
PIN 2069/1667
 Reculida - Geinitziidae
Shurabia minuta n. sp.19 Rasnitsyn 1982 winged insect
PIN 2069/1742, 1678
Shurabia minutissima n. sp.2 Aristov 2022 winged insect
PIN 2069/1495
 Blattaria - Caloblattinidae
Thuringoblatta sogdianensis n. sp.30 Vishnyakova 1998 cockroach
PIN 2069/41, 42, 88, 139, 187, 963
 Dicondylia - Permosialidae
Permosialis triassica3 Novokshonov and Zhuzhgova 2004 winged insect
PIN 2069/3592
 Hemiptera - Pincombeidae
Madygenopsyllidium djailautshoense n. gen. n. sp.5 Becker-Migdisova 1985 true bug
PIN 2069/3204
 Hemiptera - Paraknightiidae
Triknightia mira n. gen. n. sp.21
Triknightia mira n. gen. n. sp.21 Shcherbakov 2021 true bug
PIN 2069/3278
 Hemiptera - Maguviopseidae
Maguviopsis kotchnevi20 Becker-Migdisova 1953 true bug
PIN 2069/1724
 Hemiptera - Mesojabloniidae
Mesojablonia kukalovae n. gen. n. sp.26
Mesojablonia kukalovae n. gen. n. sp.26 Storozhenko 1992 true bug
PIN 2069/1677
 Grylloblattodea - Madygenophlebiidae
Madygenophlebia nana n. sp.25 Storozhenko 1992 ice crawler
PIN 2069/1640,1609
 Grylloblattodea - Gorochoviidae
Gorochoviella conjuncta n. gen.28 Storozhenko 1994 ice crawler
PIN 2069/1846?
Pseudoliomopterites lucidus n. sp.28 Storozhenko 1994 ice crawler
PIN 2069/1482
Pseudoliomopterites obscurus n. gen. n. sp.28 Storozhenko 1994 ice crawler
PIN 2069/1805
Gorochovia bifurca28 Storozhenko 1994 ice crawler
PIN 2069/1846?
Gorochovia individua n. gen.28 Storozhenko 1994 ice crawler
PIN 2069/1404,1590,1772,1783,1884
Gorochovia minuta n. sp.28 Storozhenko 1994 ice crawler
 Orthoptera - Permelcanidae
Meselcana madygenica n. gen. n. sp.
Meselcana madygenica n. gen. n. sp. Sharov 1968 grasshopper
PIN 2069/2409
 Orthoptera - Haglidae
Proisfaroptera martynovi n. sp. Sharov 1968 hump-winged cricket
PIN 2069/2440, 2360
Protshorkuphlebia triassica Sharov 1968 hump-winged cricket
PIN 2069/2352
Maragella reducta n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2069/2374
Cantohagla gracilis n. gen. n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2069/2442
Tinnihagla handlirschi n. sp.6 Gorochov 1986 hump-winged cricket
PIN 2069/2418
 Orthoptera - Xenopteridae
Pseudoferganopsis zini n. gen. n. sp.9 Gorochov 1989 grasshopper
PIN 2069/2178
Ferganotriassia lata n. gen. n. sp.9 Gorochov 1989 grasshopper
PIN 2069/2266
Ferganopterus clarus Sharov 1968 grasshopper
text says 121 imprints doesn't specify collection
Triassoferganella angusta n. gen. n. sp.9 Gorochov 1989 grasshopper
PIN 2069/2239
 Orthoptera - Proparagryllacrididae
"Brevibatkenella abscisa n. gen. n. sp." = Gryllacrimima perfecta7, Parafergania sharovi n. gen. n. sp.7
"Brevibatkenella abscisa n. gen. n. sp." = Gryllacrimima perfecta7 Sharov 1968 grasshopper
PIN 2069/2240
Parafergania sharovi n. gen. n. sp.7 Gorochov 1987 grasshopper
PIN 2069/2341
 Orthoptera - Mesoedischiidae
Mesoedischia kirgizica n. sp.8 Gorochov 1987 grasshopper
PIN 2069/2318
Mesoedischia obliqua8 Gorochov 1987 grasshopper
PIN 2069/2319
 Orthoptera - Bintoniellidae
"Oshiella oblonga n. sp." = Probintoniella oblonga8, "Oshiella crassa n. gen. n. sp." = Probintoniella crassa8, Probintoniella triassica n. gen. n. sp., Stenoshiella angusta n. gen. n. sp.8, Paroshiella alia n. gen. n. sp.8, Eubintoniella ferganica n. gen. n. sp.8
"Oshiella oblonga n. sp." = Probintoniella oblonga8 Gorochov 1987 grasshopper
PIN 2069/2326
"Oshiella crassa n. gen. n. sp." = Probintoniella crassa8 Gorochov 1987 grasshopper
PIN 2069/2321
Probintoniella triassica n. gen. n. sp. Sharov 1968 grasshopper
PIN 2069/2320
Stenoshiella angusta n. gen. n. sp.8 Gorochov 1987 grasshopper
PIN 2069/2325
Paroshiella alia n. gen. n. sp.8 Gorochov 1987 grasshopper
PIN 2069/2324
Eubintoniella ferganica n. gen. n. sp.8 Gorochov 1987 grasshopper
PIN 2069/2327
 Orthoptera - Gryllavidae
Gryllavus madygenicus Sharov 1968 grasshopper
PIN 2069/2429
 Orthoptera - Locustavidae
Brevilocustavus microscopicus n. sp.10 Gorochov 2005 grasshopper
PIN 2069/2329
 Plecoptera - Perlariopseidae
Tritaniella mera23 Sinitshenkova 1987 stonefly
PIN 2069/1907, 1915
Tritaniella synneura23 Sinitshenkova 1987 stonefly
PIN 2069/1902, 1906, 1913
Fritaniopsis remota23 Sinitshenkova 1987 stonefly
PIN 2069/1914
Dicronemoura dira n. sp.23 Sinitshenkova 1987 stonefly
PIN 2069/1909, 1906
Dicronemoura acaulis23 Sinitshenkova 1987 stonefly
PIN 2069/1905, 1908, 1910
Ramonemoura constricta n. gen. n. sp.23 Sinitshenkova 1987 stonefly
PIN 2069/1899
 Coleoptera - Triadocupedidae
Platycupes pusillus n. sp.14, Platycupes reticulatus n. sp.14, Platycupes dolichocerus n. gen. n. sp.14, Triadocupes ferghanensis n. gen. n. sp.14, Triadocupes ellipticus n. sp.14, Triadocupes latus n. sp.14, Cupesia monilicornia n. gen. n. sp.14
Platycupes pusillus n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1099
Platycupes reticulatus n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1063
Platycupes dolichocerus n. gen. n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1107
Triadocupes ferghanensis n. gen. n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1098
Triadocupes ellipticus n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1111
Triadocupes latus n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1101
Cupesia monilicornia n. gen. n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1115
 Coleoptera - Triaplidae
Triassocatinius brachynotus n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1291 (1069 is error)
Avocatinus elongatus15 Ponomarenko 1969 beetle
PIN 2069/1300
Catinoides rotundatus n. gen. n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1358
Triaplus laticoxa4 Ponomarenko 1977 beetle
PIN 2069/1228, 1294
 Coleoptera - Obrieniidae
Obrienia illaetabilis31 Zherikhin and Gratshev 1993 weevil
PIN 2069/3550
 Coleoptera - Ademosynidae
Ademosyne kirghizica n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1369
Cephalosyne capitata15 Ponomarenko 1969 beetle
PIN 2069/1342
Dolichosyne confragosa n. gen. n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1336, 1340
 Coleoptera - Phoroschizidae
Lethocoleus sternalis n. gen. n. sp.15, Schizophoroides rugosus n. gen. n. sp.15
Lethocoleus sternalis n. gen. n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1273
Schizophoroides rugosus n. gen. n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1290
 Coleoptera - Tricoleidae
Tricoleodes acutus n. gen. n. sp.15
Tricoleodes acutus n. gen. n. sp.15 Ponomarenko 1969 beetle
PIN 2069/1194, 1196
 Coleoptera - Ommatidae
Notocupoides fasciatus n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1072
Notocupoides capitatus n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1121
 Coleoptera - Cupedidae
Pterocupes antennatus n. gen. n. sp.14, Pterocupes leptocerus n. sp.14, "Mesocupoides proporeius" = Kirghizocupes proporeius15, Kirghizocupes cellulosus n. gen. n. sp.14
Pterocupes antennatus n. gen. n. sp.14 Ponomarenko 1966 reticulated beetle
PIN 2069/1076
Pterocupes leptocerus n. sp.14 Ponomarenko 1966 reticulated beetle
PIN 2069/1126
"Mesocupoides proporeius" = Kirghizocupes proporeius15 Ponomarenko 1969 reticulated beetle
PIN 2069/1053, 1034
Kirghizocupes cellulosus n. gen. n. sp.14 Ponomarenko 1966 reticulated beetle
PIN 2069/1071
 Coleoptera -
Rhabdocupes minor n. sp.14, Rhabdocupes longus n. gen. n. sp.14
Rhabdocupes minor n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1039
Rhabdocupes longus n. gen. n. sp.14 Ponomarenko 1966 beetle
PIN 2069/1041
 Mecoptera - Parachoristidae
Parachorista sana13 Novokshonov 2001 scorpionfly
PIN 2069/3442
 Mecoptera - Mesopsychidae
Prochoristella ignara12 Novokshonov 1997 scorpionfly
PIN 2069/3394
 Mecoptera - Orthophlebiidae
Choristopanorpa opinata13 Novokshonov 2001 scorpionfly
PIN 2069/3460
 Mecoptera - Permochoristidae
Mesageta rieki12 Novokshonov 1997 scorpionfly
PIN 2069/3525
Mecolusor confusicius n. gen. n. sp.13 Novokshonov 2001 scorpionfly
PIN 2069/3391
 Hymenoptera - Xyelidae
Triassoxyela kirgizica n. sp.17, Triassoxyela orycta18, "Euryxyela lata n. sp." = Leioxyela lata17, "Asioxyela antiqua n. sp." = Leioxyela antiqua17, "Triassoxyela striata n. sp." = Xiphoxyela striata17, Xyelinus angustiradius n. gen. n. sp.17, Euryxyela euryptera n. gen. n. sp.17, Asioxyela smilodon n. gen. n. sp.17, Asioxyela parvula n. sp.17, "Triassoxyela grandipennis n. sp." = Asioxyela grandipennis17
Triassoxyela kirgizica n. sp.17 Rasnitsyn 1964 sawfly
PIN 2069/1
Triassoxyela orycta18 Rasnitsyn 1969 sawfly
PIN 2069/4 (paratype of Triassoxyela kirghizica Rasnitsyn, 1964)
"Euryxyela lata n. sp." = Leioxyela lata17 Rasnitsyn 1964 sawfly
PIN 2069/9
"Asioxyela antiqua n. sp." = Leioxyela antiqua17 Rasnitsyn 1964 sawfly
PIN 2069/7
"Triassoxyela striata n. sp." = Xiphoxyela striata17 Rasnitsyn 1964 sawfly
PIN 2069/2
Xyelinus angustiradius n. gen. n. sp.17 Rasnitsyn 1964 sawfly
PIN 2069/10, 11
Euryxyela euryptera n. gen. n. sp.17 Rasnitsyn 1964 sawfly
PIN 2069/8
Asioxyela smilodon n. gen. n. sp.17 Rasnitsyn 1964 sawfly
PIN 2069/5
Asioxyela parvula n. sp.17 Rasnitsyn 1964 sawfly
PIN 2069/6
"Triassoxyela grandipennis n. sp." = Asioxyela grandipennis17 Rasnitsyn 1964 sawfly
PIN 2069/3
 Dicondylia - Paurophlebiidae
Cladophlebia parvula16 Pritykina 1981 winged insect
PIN 2069/29, 38
Paurophlebia angusta16 Pritykina 1981 winged insect
PIN 2069/31
 Dicondylia - Batkeniidae
"Triassoneura primitiva n. sp." = Paratriassoneura primitiva16
"Triassoneura primitiva n. sp." = Paratriassoneura primitiva16 Pritykina 1981 winged insect
PIN 2069/18
 Dicondylia - Kennedyidae
Kennedya carpenteri16 Pritykina 1981 winged insect
PIN 2069/37
 Dicondylia - Protomyrmeleontidae
Ferganagrion kirghiziensis n. gen. n. sp.11
Ferganagrion kirghiziensis n. gen. n. sp.11 Nel et al. 2005 winged insect
PIN 2069/3266
 Diptera - Hennigmatidae
Anemeca liya n. gen. n. sp.22
Anemeca liya n. gen. n. sp.22 Shcherbakov 1995 fly
PIN 2068/3354
 Diptera - Gnomuscidae
Gnomusca renyxa n. sp.22
Gnomusca renyxa n. sp.22 Shcherbakov 1995 fly
PIN 2069/3756
 Diptera - Protorhyphidae
Vymrhyphus triassicus n. sp.22 Blagoderov 1995 fly
PIN 2069/3757
 Diptera - Limoniidae
Mabelysia charlesi n. gen. n. sp.22
Mabelysia charlesi n. gen. n. sp.22 Shcherbakov 1995 crane fly
PIN 2069/3541