Karatau-Mikhailovka (PIN collection 2384) (Jurassic to of Kazakhstan)

Where: Chimkent, Kazakhstan (42.9° N, 70.0° E: paleocoordinates 41.6° N, 73.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Karabastau Formation, Callovian to Callovian (166.1 - 157.3 Ma)

Environment/lithology: lacustrine - large; lithified siltstone

• "dark shales" Grimaldi & Engel, 2005

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Reposited in the PIN

Primary reference: A. G. Sharov. 1968. Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 118:1-216 [C. Beard/C. Beard/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106102: authorized by Matthew Clapham, entered by Matthew Clapham on 02.03.2011, edited by Andrew Spencer-Lee and Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Liadopsyllidae
Liadopsylla grandis4 Becker-Migdisova 1985 jumping plant louse
PIN 2384/324, 328
Liadopsylla tenuicornis4 Martynov 1926 jumping plant louse
PIN 2384/318
 Hemiptera - Protopsyllidiidae
Karatavopsyllidium martynovi n. sp.4 Becker-Migdisova 1985 true bug
PIN 2384/293
 Hemiptera - Naucoridae
Aidium pleurale n. gen. n. sp.34
Aidium pleurale n. gen. n. sp.34 Popov 1968 creeping water bug
PIN 2384/368
 Hemiptera - Corixidae
Archaecorixa lata n. gen. n. sp.34
Archaecorixa lata n. gen. n. sp.34 Popov 1968 water boatman
PIN 2384/377
 Permopsocida - Archipsyllidae
Archipsylla lata n. sp.48 Vishnyakova 1976 winged insect
PIN 2384/335
 Psocodea - Electrentomidae
Paramesopsocus adibi3 Azar et al. 2008 book louse
PIN 2384/286, 291
 Blattaria - Caloblattinidae
Asioblatta punctata47 Vishnyakova 1968 cockroach
PIN 2384/35,37
 Plecoptera - Perlariopseidae
Karanemoura abrupta n. gen. n. sp.45
Karanemoura abrupta n. gen. n. sp.45 Sinitshenkova 1987 stonefly
PIN 2384/2367
 Dicondylia -
Blattodea indet.1 Latreille 1810 cockroach
PIN 2384/8 described as a paratype of Blattogryllus karatavicus
 Blattodea - Mesoblattinidae
Latiblatta lativalvata47 Vishnyakova 1968 cockroach
PIN 2384/192
 Blattodea - Liberiblattinidae
Hydrokhoohydra aquabella50 Vršanský 2019 cockroach
PIN 2384/186, 194, 1274
 Dermaptera - Protodiplatyidae
Microdiplatys oculatus n. sp.49 Vishnyakova 1980 earwig
PIN 2384/196
 Dicondylia - Brachyphyllophagidae
Brachyphyllophagus phantasus n. sp.43 Rasnitsyn 2000 winged insect
PIN 2384/260
 Phasmatodea - Susumaniidae
Phasmomimoides minutus n. sp.8 Gorochov 2000 stick insect
PIN 2384/261
 Orthoptera - Protogryllidae
"Protogryllus karatavicus n. sp." = Falsispeculum karatavicus
"Protogryllus karatavicus n. sp." = Falsispeculum karatavicus Sharov 1968 grasshopper
PIN 2384/255
 Orthoptera - Prophalangopsidae
Aboilus aulietus Sharov 1968 hump-winged cricket
PIN 2384/248
Aboilus femineus7 Gorochov 1996 hump-winged cricket
PIN 2384/217
 Hymenoptera - Ephialtitidae
? Asiephialtites ventricosus n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1332
Sessiliventer symphytus n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1328
Stephanogaster pyriformis n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1320
Symphytopterus nigricornis40 Rasnitsyn 1975 wasp
PIN 2384/1323
 Hymenoptera - Mesoserphidae
Scoliuroserphus pallidulus42 Rasnitsyn 1986 wasp
PIN 2384/1367
 Hymenoptera - Praeaulacidae
Praeaulacites medius39 Rasnitsyn 1972 wasp
PIN 2384/1317, 1318, 1329
Praeaulacites subniger n. sp.39 Rasnitsyn 1972 wasp
PIN 2384/1945
 Hymenoptera - Anomopterellidae
"Anomopterella stenocera n. sp." = Choristopterella stenocera40
"Anomopterella stenocera n. sp." = Choristopterella stenocera40 Rasnitsyn 1975 wasp
PIN 2384/1078
 Hymenoptera - Megalyridae
Mesaulacinus stenocerus n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1325
Leptocleistogaster alboscuta n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1319
 Hymenoptera - Bethylonymidae
Bethylonymus nigricornis n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1946
Bethylonymus pedalis n. sp.40 Rasnitsyn 1975 wasp
PIN 2384/1333
 Hymenoptera - Brachycleistogastridae
Brachycleistogaster nigritibialis40 Rasnitsyn 1975 wasp
PIN 2384/1344
 Hymenoptera - Anaxyelidae
? Anaxyelidae indet.16 Martynov 1925 winged insect
PIN 2384/1313
Anaxyela destructa n. sp.38 Rasnitsyn 1969 winged insect
PIN 2384/1314
 Hymenoptera -
Prolyda karatavica n. gen. n. sp.37
Prolyda karatavica n. gen. n. sp.37 Rasnitsyn 1968 sawfly
PIN 2384/1311
 Hymenoptera - Xyelidae
Lydoxyela excellens n. gen. n. sp.36
Lydoxyela excellens n. gen. n. sp.36 Rasnitsyn 1966 sawfly
PIN 2384/1316
 Hymenoptera - Xyelotomidae
Xyelocerus admirandus n. gen. n. sp.37
Xyelocerus admirandus n. gen. n. sp.37 Rasnitsyn 1968 wood wasp
PIN 2384/1315
 Coleoptera - Cerophytidae
"Idiomerus musculus" = Necromera muscula6, "Idiomerus intermedius n. sp." = Necromera intermedia6
"Idiomerus musculus" = Necromera muscula6 Dolin 1980 beetle
PIN 2384/475
"Idiomerus intermedius n. sp." = Necromera intermedia6 Dolin 1980 beetle
PIN 2384/497
 Coleoptera - Elateridae
Hypnomorphoides latus6 Dolin 1980 click beetle
PIN 2384/449
Adiagnostus minutulus n. sp.6 Dolin 1980 click beetle
PIN 2384/455
Codemus synaptoides6 Dolin 1980 click beetle
PIN 2384/481(483)
Codemus quadricolis6 Dolin 1980 click beetle
PIN 2384/500, 502
Codemus carinatus n. sp.6 Dolin 1980 click beetle
PIN 2384/464, 483
Codemus sharovi n. sp.6 Dolin 1980 click beetle
PIN 2384/474, 465
Hypnomorphus curtus6 Dolin 1980 click beetle
PIN 2384/472
Hypnomorphus inventus n. sp.5 Dolin 1975 click beetle
PIN 2384/491, 450, 479
Hypnomorphus rohdendorfi n. gen. n. sp.5 Dolin 1975 click beetle
PIN 2384/457
Platyelater reflexicollis6 Dolin 1980 click beetle
PIN 2384/485
Lithomerus cockerelli n. gen. n. sp.6 Dolin 1980 click beetle
PIN 2384/454, 460, 492
Acheonus abbreviatus n. gen. n. sp.6 Dolin 1980 click beetle
PIN 2384/462 (paratype also given as specimen 462 in text
Lithocoelus detrusus n. gen. n. sp.5 Dolin 1975 click beetle
PIN 2384/484
Desmatus beckeri5 Dolin 1975 click beetle
PIN 2384/489
 Coleoptera - Staphylinidae
Abolescus glabratus46 Tikhomirova 1968 rove beetle
PIN 2384/423,427
Mesoxytelus mandibularis46 Tikhomirova 1968 rove beetle
PIN 2384/444
Archodromus comptus n. gen. n. sp.46 Tikhomirova 1968 rove beetle
PIN 2384/418
Mesotachinus major n. sp.46 Tikhomirova 1968 rove beetle
PIN 2384/417,419
 Coleoptera - Parandrexidae
Parandrexis parvula15 Martynov 1926 beetle
PIN 2384/701
 Coleoptera - Anthribidae
Protoscelis parvula20 Medvedev 1968 fungus weevil
PIN 2384/509, 639
Protoscelis jurassica24 Medvedev 1968 fungus weevil
PIN 2384/770; 655,725,726,733,751,752,781,789 (in Legalov, 2013)
"Pseudomegamerus grandis" = Protoscelis longicornis24 Medvedev 1968 fungus weevil
PIN 2384/415
Protoscelis medvedevi20 Legalov 2013 fungus weevil
PIN 2384/569
 Coleoptera - Nemonychidae
"Brenthorrhinus brevirostris n. sp." = Abrenthorrhinus brevirostris11 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2384/510
"Distenorrhinus (Astenorrhinus) major n. sp." = Astenorrhinus major12 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2384/526
Procurculio fortipes12 Arnoldi 1977 pine flower snout beetle
PIN 2384/517
"Pseudobrenthorrhinus crassicornis n. gen. n. sp." = Eccoptarthrus crassipes11 Arnoldi 1977 pine flower snout beetle
PIN 2384/537
Distenorrhinus (Distenorrhinus) pallidirostris12 Gratshev and Zherikhin 1995 pine flower snout beetle
PIN 2384/536
Ampliceps dubius10 Arnoldi 1977 pine flower snout beetle
PIN 2384/511, 613
"Scelocamptus curvipes n. sp." = Ampliceps tenuirostris2 Arnoldi 1977 pine flower snout beetle
PIN 2384/512
Ampliceps dentitibia10 Arnoldi 1977 pine flower snout beetle
PIN 2384/530
Ampliceps furcitibia10 Arnoldi 1977 pine flower snout beetle
PIN 2384/507, 653
Oxycorynoides (Oxycorynoides) similis10 Arnoldi 1977 pine flower snout beetle
PIN 2384/513, 520, 532, 533, 538
Oxycorynoides (Pseudoxycorynoides) sukatshevae10 Legalov 2014 pine flower snout beetle
PIN 2384/514, 521, 534
Eobelus longipes2 Arnoldi 1977 pine flower snout beetle
PIN 2384/516
Belonotaris (Belonotaris) punctatissimus10 Arnoldi 1977 pine flower snout beetle
PIN 2384/516
Belonotaris (Arnoldibelus) rasnitsyni10 Legalov 2014 pine flower snout beetle
PIN 2384/531
 Coleoptera - Brentidae
Karacar contractus n. gen. n. sp.9
Karacar contractus n. gen. n. sp.9 Gratshev and Legalov 2011 weevil
 Coleoptera - Hybosoridae
Protohybosorus mesoasiaticus n. sp.29 Nikolajev 2010 scavenger scarab beetle
PIN 2384/760
 Coleoptera - Scarabaeidae
Antemnacrassa albosulcata n. sp.28 Nikolajev 2004 scarab beetle
PIN 2384/575
 Coleoptera - Coptoclavidae
Charonoscaphidia elongata n. gen. n. sp.2
Charonoscaphidia elongata n. gen. n. sp.2 Ponomarenko 1977 beetle
PIN 2384/717, 707, 718
 Coleoptera - Trachypachidae
Karadromeus latus n. sp.2 Ponomarenko 1977 false ground beetle
PIN 2384/567
Psacodromeus ovalis2 Ponomarenko 1977 false ground beetle
PIN 2384/757
 Coleoptera - Carabidae
Ovrabites ovalis2 Ponomarenko 1977 ground beetle
PIN 2384/554
 Coleoptera - Schizophoridae
Tersus leptocerus33 Ponomarenko 1968 beetle
PIN 2384/715
 Coleoptera - Triaplidae
Catinius pelta n. gen. n. sp.33
Catinius pelta n. gen. n. sp.33 Ponomarenko 1968 beetle
PIN 2384/634
 Coleoptera - Obrieniidae
Pseudobrienia rasnitsyni n. gen. n. sp.19
Pseudobrienia rasnitsyni n. gen. n. sp.19 Legalov 2012 beetle
PIN 2384/720
 Coleoptera - Cupedidae
Cupidium abavum n. gen. n. sp.33 Ponomarenko 1968 reticulated beetle
PIN 2384/398
Notocupes nigrimonticola n. sp.33 Ponomarenko 1968 reticulated beetle
PIN 2384/397
Notocupes lapidarius n. sp.33 Ponomarenko 1968 reticulated beetle
PIN 2384/404
Omma aberratum n. sp.33 Ponomarenko 1968 reticulated beetle
PIN 2384/396
Tetraphalerus maximus33 Ponomarenko 1968 reticulated beetle
PIN 2384/401
 Mecoptera - Bittacidae
Sibirobittacus kazakhstanensis n. sp.30 Novokshonov 1993 hangingfly
PIN 2384/942
 Diptera -
Apystomimus zaitzevi26 Mostovski 1999 fly
PIN 2384/1258
 Diptera - Rhagionidae
Protorhagio karataviensis27 Mostovski 2008 snipe fly
PIN 2384/1146
Protorhagio ruptus27 Mostovski 2008 snipe fly
PIN 2384/1224 (also listed under Protorhagio capitatus)
 Diptera - Nemestrinidae
"Eohirmoneura carpenteri n. gen. n. sp." = Hirmoneura (Eohirmoneura) carpenteri44
"Eohirmoneura carpenteri n. gen. n. sp." = Hirmoneura (Eohirmoneura) carpenteri44 Rohdendorf 1968 tangle-veined fly
PIN 2384/1236
 Diptera - Protorhyphidae
Archirhyphus asiatcus22 Rohdendorf 1964 fly
PIN 2384/1021
 Diptera - Procramptonomyiidae
Procramptonomyia kovalevi22 Lukashevich 2012 fly
PIN 2384/1149,1173, 1281
 Diptera - Elliidae
Polyanka minuta17 Krzemińska et al. 1993 fly
PIN 2384/988, 87
 Diptera - Trichoceridae
Tanyochoreta (Trichokara) minuta18 Krzemińska et al. 2009 winter crane fly
PIN 2384/1201
 Diptera - Tanyderidae
Praemacrochile ansorgei23 Lukashevich and Krzemiński 2009 primitive crane fly
PIN 2384/1067, 1266
 Diptera - Ptychopteridae
Eoptychopterina karatavica21 Lukashevich 1993 phantom crane fly
PIN 2384/1238; 2323 in Lukashevich, 2004
 Neuroptera - Coniopterygidae
Juraconiopteryx zherichini n. gen. n. sp.25
Juraconiopteryx zherichini n. gen. n. sp.25 Meinander 1975 dustywing
PIN 2384/1300
 Neuroptera - Polystoechotidae
Osmyloides distinctus n. gen. n. sp.32, "Kasachstania fasciata n. gen. n. sp." = Panfilovdvia fasciata32
Osmyloides distinctus n. gen. n. sp.32 Panfilov 1980 giant lacewing
PIN 2384/863
"Kasachstania fasciata n. gen. n. sp." = Panfilovdvia fasciata32 Panfilov 1980 giant lacewing
PIN 2384/814
 Neuroptera -
Chrysoleonites plexus n. sp.32 Panfilov 1980 lacewing
PIN 2384/842
 Neuroptera - Kalligrammatidae
Lithogramma oculatum n. gen. n. sp.31, Kalligramma flexuosum n. sp.31
Lithogramma oculatum n. gen. n. sp.31 Panfilov 1968 lacewing
PIN 2384/856
Kalligramma flexuosum n. sp.31 Panfilov 1968 lacewing
PIN 2384/857
 Neuroptera - Chrysopidae
Baisochrysa pumila14 Khramov et al. 2015 green lacewing
PIN 2384/838
 Neuroptera - Osmylidae
Jurosmylus parvulus13 Khramov 2014 lance lacewing
PIN 2384/849
Scapoptera recta n. gen. n. sp.32 Panfilov 1980 lance lacewing
PIN 2384/862
 Odonata - Stenophlebiidae
Stenophlebia karatavica35 Pritykina 1968 dragonfly
PIN 2384/3
 Odonata -
"Protolindenia aktassica" = Kazakhophlebiella aktassica35
"Protolindenia aktassica" = Kazakhophlebiella aktassica35 Pritykina 1968 true dragonfly
PIN 2384/2
 Odonata - Aktassiidae
Aktassia magna n. gen. n. sp.35
Aktassia magna n. gen. n. sp.35 Pritykina 1968 true dragonfly
PIN 2384/4
 Odonata - Isophlebiidae
Isophlebiodes obscurus n. gen. n. sp.35
Isophlebiodes obscurus n. gen. n. sp.35 Pritykina 1968 dragonfly
PIN 2384/18
 Odonata - Selenothemistidae
Turanothemis nodalis n. gen. n. sp.35
Turanothemis nodalis n. gen. n. sp.35 Pritykina 1968 dragonfly
PIN 2384/9
 Eoblattida - Blattogryllidae
Blattogryllus karatavicus41 Rasnitsyn 1976 insect
PIN 2384/8, 205