Nanligezhuang, Tuanwang Town, Laiyang City (Shandong Prov. Mus.) (Cretaceous of China)

Where: Shandong, China (36.8° N, 120.7° E: paleocoordinates 40.2° N, 122.1° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: 3 Member (Laiyang Formation), Aptian (125.0 - 113.0 Ma)

Environment/lithology: lacustrine - large; lithified, black, green shale

• Large (several hundred km2) lake in the Laiyang basin. In addition to the abundant entomofauna, plant fossil fragments, fish and a single bird feather also have been found. Water plants were apparently rare, and at the centre of the lake I have found no plant or animal fossils. Extensional regime in response to the progressive attenuation of the thickened lithosphere and orogenic collapse.
• The Third Member of the Laiyang Formation consists of 250-705 m of calcareous sandstone, siltstone and fine-granted sandstone intercalated with thin, paper-like, insect-rich shales.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collection methods: Repository: Shandong Provincial Museum

Primary reference: J. Zhang. 1999. Aeschnidiid nymphs from the Jehol biota (latest Jurassic–Early Cretaceous), China, with a discussion of the family Aeschnidiidae (Insecta, Odonata). Cretaceous Research 20:813-827 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 106313: authorized by Matthew Clapham, entered by Jered Karr on 08.03.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera - Sinaphididae
Sinaphidum epichare n. sp.23 Zhang et al. 1989 aphid
L84123
 Hemiptera -
Penaphis nanligezhuangensis n. sp.1 Lin 1995 aphid
L84127 (Penaphis circa in Zhang, 1989)
 Hemiptera - Palaeoaphididae
Caudaphis minulissima n. sp.23 Zhang et al. 1989 aphid
L84118, L84119
 Hemiptera - Archijassidae
? Archijassus plurinervis n. sp.2 Zhang 1985 true bug
L85109
 Hemiptera - Cicadellidae
Sinojassus brevispinatus n. gen. n. sp.2
Sinojassus brevispinatus n. gen. n. sp.2 Zhang 1985 leafhopper
L85108
 Hemiptera - Ignotingidae
Ignotingis mirifica n. gen. n. sp.18
Ignotingis mirifica n. gen. n. sp.18 Zhang et al. 2005 true bug
L86812, 86811, 86813-86818
 Hemiptera - Corixidae
Karataviella shandongensis n. sp.2 Zhang 1985 water boatman
L84999
 Hemiptera - Archegocimicidae
Schizopteryx lacustris n. sp.8 Zhang 1991 shore bug
L88255, 88256, 88258, 88258, 88260
 Hemiptera - Saldidae
Mesolygaeus laiyangensis10 Ping 1928 shore bug
L88221-88227, 88231, 88236, 88237, 88238, 88241, 86317
 Blattodea - Mesoblattinidae
Laiyangia delicatula n. sp.2 Zhang 1985 cockroach
L84993(4)
 Dermaptera -
Archaeosoma serratum n. gen. n. sp.13
Archaeosoma serratum n. gen. n. sp.13 Zhang 1994 earwig
L89203
 Dermaptera - Protodiplatyidae
Longicerciata mesozoica n. gen. n. sp.13 Zhang 1994 earwig
L89201
Longicerciata rumpens n. sp.13 Zhang 1994 earwig
L89202
 Hymenoptera - Scoliidae
Cretoscolia rasnitsyni n. sp.14 Zhang 2004 wasp
L97929
Cretoscolia formosa n. sp.14 Zhang 2004 wasp
L91550
Cretoscolia laiyangica n. sp.14 Zhang 2004 wasp
L91548
Cretaproscolia asiatica n. sp.16 Zhang 2006 wasp
L91311
 Hymenoptera - Mutillidae
Mesomutilla aptera n. gen. n. sp.2
Mesomutilla aptera n. gen. n. sp.2 Zhang 1985 velvet ant
L85119
 Hymenoptera - Angarosphecidae
Eubaissodes completus n. gen. n. sp.11, "Calobaissodes strigosus n. gen. n. sp." = Angarosphex strigosus11
Eubaissodes completus n. gen. n. sp.11 Zhang 1992 apoid wasp
L88701
"Calobaissodes strigosus n. gen. n. sp." = Angarosphex strigosus11 Zhang 1992 apoid wasp
L88702
 Hymenoptera - Ichneumonidae
"Tanychora sinensis n. sp." = Khasurtella sinensis5, Polychorella magica n. gen. n. sp.2
"Tanychora sinensis n. sp." = Khasurtella sinensis5 Zhang 1991 ichneumon wasp
L84984
Polychorella magica n. gen. n. sp.2 Zhang 1985 ichneumon wasp
L85117
 Hymenoptera - Pelecinidae
Allopelecinus terpnus n. gen. n. sp.20 Zhang and Rasnitsyn 2006 wasp
L91446 (stated as recently collected by JZ)
Sinopelecinus daspletis n. sp.20 Zhang and Rasnitsyn 2006 wasp
L91448 (stated as recently collected by JZ)
Sinopelecinus hierus n. sp.20 Zhang and Rasnitsyn 2006 wasp
L91444 (stated as recently collected by JZ)
Eopelecinus hodoiporus n. sp.15 Zhang 2005 wasp
L91439
Eopelecinus giganteus n. sp.15 Zhang 2005 wasp
L91436
Eopelecinus leptaleus n. sp.15 Zhang 2005 wasp
L91442
Eopelecinus pusillus n. sp.15 Zhang 2005 wasp
L91435
Eopelecinus mesomicrus n. sp.15 Zhang 2005 wasp
L91443
Eopelecinus mecometasomatus n. sp.15 Zhang 2005 wasp
L91438
Eopelecinus eucallus n. sp.15 Zhang 2005 wasp
L91437
Eopelecinus laiyangicus n. sp.15 Zhang 2005 wasp
L91440
 Hymenoptera - Mesoserphidae
Otlia ectemnia n. gen. n. sp.12
Otlia ectemnia n. gen. n. sp.12 Zhang 1992 wasp
L88714
 Hymenoptera - Heloridae
Laiyanghelorus erymnus n. gen. n. sp.12
Laiyanghelorus erymnus n. gen. n. sp.12 Zhang 1992 wasp
L88713
 Hymenoptera - Baissidae
Humiryssus specialis n. sp.19 Zhang and Rasnitsyn 2004 wasp
L91457
Humiryssus cancellatus n. sp.19 Zhang and Rasnitsyn 2004 wasp
L91454
Humiryssus vulgatus n. sp.19 Zhang and Rasnitsyn 2004 wasp
L91555
Mesepipolaea nanligezhuangica n. gen. n. sp.19 Zhang and Rasnitsyn 2004 wasp
L91466, 91470, 91468
 Coleoptera - Staphylinidae
"Mesostaphylinus fraternus n. sp." = Thayeralinus fraternus21, Mesostaphylinus laiyangensis n. gen. n. sp.4, Laostaphylinus nigritellus n. gen. n. sp.4, Laostaphylinus fuscus n. sp.4, Hesterniasca obesa n. gen. n. sp.21
"Mesostaphylinus fraternus n. sp." = Thayeralinus fraternus21 Zhang et al. 1992 rove beetle
L84139
Mesostaphylinus laiyangensis n. gen. n. sp.4 Zhang 1988 rove beetle
L84136
Laostaphylinus nigritellus n. gen. n. sp.4 Zhang 1988 rove beetle
L84137
Laostaphylinus fuscus n. sp.4 Zhang 1988 rove beetle
L84138
Hesterniasca obesa n. gen. n. sp.21 Zhang et al. 1992 rove beetle
L84140
 Coleoptera - Trogossitidae
Sinosoronia longiantenna n. gen. n. sp.9, Palaeoendomychus gymnus n. gen. n. sp.9
Sinosoronia longiantenna n. gen. n. sp.9 Zhang 1992 bark gnawing beetle
L85309
Palaeoendomychus gymnus n. gen. n. sp.9 Zhang 1992 bark gnawing beetle
L85301/2
 Coleoptera - Coptoclavidae
Coptoclava longipoda10 Ping 1928 beetle
dominant species
 Diptera - Chironomidae
Coelochironoma xantha n. gen. n. sp.22, Protobibio orientalis n. sp.22, Sinoryctochlus insolitus n. gen. n. sp.6
Coelochironoma xantha n. gen. n. sp.22 Zhang et al. 1986 non biting midge
L84106
Protobibio orientalis n. sp.22 Zhang et al. 1986 non biting midge
L84111
Sinoryctochlus insolitus n. gen. n. sp.6 Zhang 1991 non biting midge
L85417, 85418
 Diptera - Chaoboridae
Chironomaptera vesca10 Kalugina 1980 phantom midge
dominant species
Chironomaptera gregaria10 Grabau 1923 phantom midge
dominant species
 Diptera - Ironomyiidae
Palaeopetia laiyangensis n. gen. n. sp.3
Palaeopetia laiyangensis n. gen. n. sp.3 Zhang 1987 ironic fly
L84130
 Diptera -
Lithopetia hirsuta n. gen. n. sp.3
Lithopetia hirsuta n. gen. n. sp.3 Zhang 1987 fly
L81432(3)
 Diptera - Zhangsolvidae
"Archisolva cupressa n. gen. n. sp." = Zhangsolva cupressa24
"Archisolva cupressa n. gen. n. sp." = Zhangsolva cupressa24 Zhang et al. 1993 fly
L88811
 Diptera - Rhagionidae
Scelorhagio mecomastigus n. gen. n. sp.24, Pseudopetia grandis n. gen. n. sp.3
Scelorhagio mecomastigus n. gen. n. sp.24 Zhang et al. 1993 snipe fly
L88812
Pseudopetia grandis n. gen. n. sp.3 Zhang 1987 snipe fly
L84134
 Diptera - Athericidae
Sinocretomyia minuscula n. gen. n. sp.17
Sinocretomyia minuscula n. gen. n. sp.17 Zhang 2012 water snipe fly
L91919
 Diptera - Tabanidae
Laiyangitabanus formosus n. gen. n. sp.17
Laiyangitabanus formosus n. gen. n. sp.17 Zhang 2012 horse fly
L91806
 Diptera - Antefungivoridae
"Pleciomimella perbella n. sp." = Lycoriomimodes perbella22, Aortomima shandongensis n. gen. n. sp.22
"Pleciomimella perbella n. sp." = Lycoriomimodes perbella22 Zhang et al. 1986 fly
L84110
Aortomima shandongensis n. gen. n. sp.22 Zhang et al. 1986 fly
L84109
 Diptera - Pediciidae
Ceuthoneura dolichoptera n. gen. n. sp.22
Ceuthoneura dolichoptera n. gen. n. sp.22 Zhang et al. 1986 hairy-eyed cranefly
L84103
 Neuroptera - Mesochrysopidae
Allopterus luianus n. gen. n. sp.7
Allopterus luianus n. gen. n. sp.7 Zhang 1991 brown lacewing
L88501(2)
 Odonata -
Zygoptera indet.10 Selys-Longchamps 1854 damselfly
L88760(1) & L88762
 Odonata - Dysagrionidae
Congqingia rhora n. gen. n. sp.10
Congqingia rhora n. gen. n. sp.10 Zhang 1992 damselfly
L88758(9) (collected by JZ in 1989)
 Odonata - Aeschnidiidae
"Aeschnidium heishankowense" = Sinaeschnidia heishankowensis
"Aeschnidium heishankowense" = Sinaeschnidia heishankowensis Hong 1965 small darner
L90403 basal portion of hindwing