PBDB Collection

Grassel, Braunschweig (Jurassic of Germany)

Where: Niedersachsen, Germany (52.4° N, 10.6° E: paleocoordinates 41.6° N, 17.6° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Posidonia Shale Formation, Early/Lower Toarcian (183.0 - 182.0 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: offshore; lithified mudstone

Size class: macrofossils

Preservation: mold/impression

Collection methods: Specimens deposited in Geol. Landesmuseum, Berlin

Primary reference: A. Bode. 1953. Die Insektenfauna des Ostniedersachsischen Oberen Lias. Palaeontographica Abteilung A 103:1-375 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 107321: authorized by Matthew Clapham, entered by Jered Karr on 04.04.2011, edited by Zachary Newman

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
-	Trimerocephalium incisum Trimerocephalium incisum Bode 1953 insect
Reculida - Geinitziidae	"Gryllacris fasciata n. sp." = Geinitzia fasciata¹ "Gryllacris fasciata n. sp." = Geinitzia fasciata¹ Bode 1905 winged insect
Odonata - Liassogomphidae	"Paragomphus angulatus n. gen. n. sp." = Phthitogomphus angulatus, "Gomphites brodiei" = Heterothemis brodiei "Paragomphus angulatus n. gen. n. sp." = Phthitogomphus angulatus Handlirsch 1939 dragonfly "Gomphites brodiei" = Heterothemis brodiei Brodie 1845 dragonfly
Odonata - Myopophlebiidae	Strongylogomphus grasselianus n. gen. n. sp., Strongylogomphus grasselianus, "Liadothemis wunnenbergi" = Paraheterophlebia wunnenbergi, "Myopophlebia libera" = Myopophlebia libera, "Myopophlebia tecta n. sp." = Myopophlebia libera Strongylogomphus grasselianus n. gen. n. sp. Bode 1953 dragonfly Strongylogomphus grasselianus Bode 1953 dragonfly "Liadothemis wunnenbergi" = Paraheterophlebia wunnenbergi Bode 1953 dragonfly "Myopophlebia libera" = Myopophlebia libera Bode 1953 dragonfly "Myopophlebia tecta n. sp." = Myopophlebia libera Bode 1953 dragonfly
Odonata - Sphenophlebiidae	Ensphingophlebia undulata n. gen. n. sp. Ensphingophlebia undulata n. gen. n. sp. Bode 1953 dragonfly
Odonata -	Syrrhoe commissa n. gen. n. sp. Syrrhoe commissa n. gen. n. sp. Bode 1953 dragonfly
Dicondylia - Protomyrmeleontidae	"Protomyrmeleon (Tillyardagrion) grasselensis n. sp." = Protomyrmeleon grasselensis "Protomyrmeleon (Tillyardagrion) grasselensis n. sp." = Protomyrmeleon grasselensis Bode 1953 winged insect
Blattaria - Raphidiomimidae	"Ptyctoblattina completa n. sp." = Liadoblattina blakei, Ptyctoblattina simplicior n. sp. "Ptyctoblattina completa n. sp." = Liadoblattina blakei Scudder 1886 cockroach Ptyctoblattina simplicior n. sp. Bode 1953 cockroach
Necrotrichoptera - Necrotauliidae	"Necrotaulius marginatus" = Necrotaulius parvulus "Necrotaulius marginatus" = Necrotaulius parvulus Geinitz 1884 winged insect
Mecoptera - Bittacidae	"Polydicrobittacus lingula" = Parabittacus lingula, Haplobittacus parvus "Polydicrobittacus lingula" = Parabittacus lingula Bode 1953 hangingfly Haplobittacus parvus Bode 1953 hangingfly
Mecoptera - Orthophlebiidae	Orthophlebia latipennisimilis, Parorthophlebia grasselensis Orthophlebia latipennisimilis Bode 1953 scorpionfly Parorthophlebia grasselensis Bode 1953 scorpionfly
Mecoptera - Pseudopolycentropodidae	Pseudopolycentropus obtusus Pseudopolycentropus obtusus Bode 1953 scorpionfly
Neuroptera - Prohemerobiidae	Prohemerobius mediolatus², "Parhemerobius dilatatus" = Parhemerobius dilatatus, "Parhemerobius aequalirotundatus" = Parhemerobius dilatatus, "Parhemerobius praeauctus" = Parhemerobius dilatatus, "Prohemerobius triangulus" = Parhemerobius dilatatus Prohemerobius mediolatus² Bode 1953 lacewing "Parhemerobius dilatatus" = Parhemerobius dilatatus Bode 1953 lacewing "Parhemerobius aequalirotundatus" = Parhemerobius dilatatus Bode 1953 lacewing "Parhemerobius praeauctus" = Parhemerobius dilatatus Bode 1953 lacewing "Prohemerobius triangulus" = Parhemerobius dilatatus Bode 1953 lacewing
Neuroptera - Kalligrammatidae	"Liassopsychops curvata" = Liassopsychops curvatus "Liassopsychops curvata" = Liassopsychops curvatus Bode 1953 lacewing
Coleoptera - Trachypachidae	"Liassocarabites praefrictus" = Prosynactus scissus, "Euenarthrus mandibulatus" = Prosynactus scissus, "Megelytrites mutilatus" = Prosynactus scissus, "Mesalocistus constrictus" = Prosynactus gracilis "Liassocarabites praefrictus" = Prosynactus scissus Bode 1953 false ground beetle "Euenarthrus mandibulatus" = Prosynactus scissus Bode 1953 false ground beetle "Megelytrites mutilatus" = Prosynactus scissus Bode 1953 false ground beetle "Mesalocistus constrictus" = Prosynactus gracilis Bode 1953 false ground beetle
Coleoptera -	Grasselites pusillus, "Aptilotus capitecarens" = Aptilotitus capitecarens, "Perosoma praecisum" = Peridosoma praecisum, Gastroratus dispertitus, Sideriosemion punctolineatum, Scalopoides inscissus, Diplocelides minutus, Metanastes denudatus, Tripsalis praecisa, Brachylaimon striatus, "Loxoncus striatulus" = Mesoncus striatulus, Omogongylus ovatus, Leptosolenophorus brevicollis, Trochiscites capitapertus, Pholipheron articulatus, Pleuralocista insculpta, Diatrypamene angulocollis, Rhomaleus ornatus Grasselites pusillus Bode 1953 beetle "Aptilotus capitecarens" = Aptilotitus capitecarens Bode 1953 beetle "Perosoma praecisum" = Peridosoma praecisum Bode 1953 beetle Gastroratus dispertitus Bode 1953 beetle Sideriosemion punctolineatum Bode 1953 beetle Scalopoides inscissus Bode 1953 beetle Diplocelides minutus Bode 1953 beetle Metanastes denudatus Bode 1953 beetle Tripsalis praecisa Bode 1953 beetle Brachylaimon striatus Bode 1953 beetle "Loxoncus striatulus" = Mesoncus striatulus Bode 1953 beetle Omogongylus ovatus Bode 1953 beetle Leptosolenophorus brevicollis Bode 1953 beetle Trochiscites capitapertus Bode 1953 beetle Pholipheron articulatus Bode 1953 beetle Pleuralocista insculpta Bode 1953 beetle Diatrypamene angulocollis Bode 1953 beetle Rhomaleus ornatus Bode 1953 beetle
Hemiptera - Pachymeridiidae	Stiphroschema longealatum, Adelocoris ambiguus Stiphroschema longealatum Bode 1953 true bug Adelocoris ambiguus Bode 1953 true bug
Hemiptera - Archegocimicidae	Megalocoris laticlavus, Corynecoris occultatus, Ensphingocoris parvulus Megalocoris laticlavus Bode 1953 shore bug Corynecoris occultatus Bode 1953 shore bug Ensphingocoris parvulus Bode 1953 shore bug
Hemiptera - Fulgoridiidae	Fulgoridium sp., Elasmoscelidium rectemarginatum, Elasmoscelidium promotum Fulgoridium sp. Handlirsch 1906 planthopper Elasmoscelidium rectemarginatum Bode 1953 planthopper Elasmoscelidium promotum Bode 1953 planthopper
Hemiptera -	"Tetragonidium parallelogramma" = Tetrafulgoria parallelogramma, "Fulgoridium (Procerofulgoridium) verticillatum" = Procerofulgoridium verticillatum, Margaroptilon formosum, "Fulgoridium (Productofulgoridium) filiferum" = Productofulgoridium filiferum, "Fulgoridium (Compactofulgoridium) obesum" = Compactofulgoridium obesum, "Fulgoridium (Compactofulgoridium) fronterotundum" = Compactofulgoridium fronterotundum "Tetragonidium parallelogramma" = Tetrafulgoria parallelogramma Bode 1953 planthopper "Fulgoridium (Procerofulgoridium) verticillatum" = Procerofulgoridium verticillatum Bode 1953 planthopper Margaroptilon formosum Bode 1953 planthopper "Fulgoridium (Productofulgoridium) filiferum" = Productofulgoridium filiferum Bode 1953 planthopper "Fulgoridium (Compactofulgoridium) obesum" = Compactofulgoridium obesum Bode 1953 planthopper "Fulgoridium (Compactofulgoridium) fronterotundum" = Compactofulgoridium fronterotundum Bode 1953 planthopper
Hemiptera - Procercopidae	Procercopis lacerata Procercopis lacerata Bode 1953 froghopper
Orthoptera - Protogryllidae	Protogryllus formosus n. sp., Protogryllus formosus Protogryllus formosus n. sp. Bode 1953 grasshopper Protogryllus formosus Bode 1953 grasshopper
Orthoptera - Elcanidae	"Elcana infralaesa n. sp." = Panorpidium media, "Elcana prominens" = Panorpidium media, "Elcana varia n. sp." = Panorpidium media, "Elcana rescripta n. sp." = Panorpidium media, "Elcana planta n. sp." = Panorpidium minima, "Elcana novemmaculata n. sp." = Panorpidium minima, "Elcana incurvata n. sp." = Panorpidium minima, "Elcana hildesiana n. sp." = Panorpidium geinitzi, "Elcana dilatata n. sp." = Panorpidium geinitzi "Elcana infralaesa n. sp." = Panorpidium media Handlirsch 1906 grasshopper "Elcana prominens" = Panorpidium media Handlirsch 1906 grasshopper "Elcana varia n. sp." = Panorpidium media Handlirsch 1906 grasshopper "Elcana rescripta n. sp." = Panorpidium media Handlirsch 1906 grasshopper "Elcana planta n. sp." = Panorpidium minima Handlirsch 1906 grasshopper "Elcana novemmaculata n. sp." = Panorpidium minima Handlirsch 1906 grasshopper "Elcana incurvata n. sp." = Panorpidium minima Handlirsch 1906 grasshopper "Elcana hildesiana n. sp." = Panorpidium geinitzi Heer 1880 grasshopper "Elcana dilatata n. sp." = Panorpidium geinitzi Heer 1880 grasshopper
Dicondylia - Aerophasmidae	"Chresmodella culminecontracta n. sp." = Schesslitziella integra "Chresmodella culminecontracta n. sp." = Schesslitziella integra Bode 1953 winged insect
-	Agmatozoon articulatum Agmatozoon articulatum Bode 1953 insect
Eoblattida - Blattogryllidae	"Dorniella pulchra" = Dorniella pulchra, "Dorniella biramosa" = Dorniella pulchra, "Dorniella multoramosa" = Dorniella pulchra "Dorniella pulchra" = Dorniella pulchra Bode 1953 insect "Dorniella biramosa" = Dorniella pulchra Bode 1953 insect "Dorniella multoramosa" = Dorniella pulchra Bode 1953 insect
Diptera -	Homoeoptychopteris incerta Homoeoptychopteris incerta Bode 1953 fly
Diptera - Heterorhyphidae	Heterorhyphus analivarius, "Heterorhyphus latus" = Heterorhyphus anomalus Heterorhyphus analivarius Bode 1953 fly "Heterorhyphus latus" = Heterorhyphus anomalus Handlirsch 1939 fly
Diptera - Protorhyphidae	Protorhyphus ovisimilis Protorhyphus ovisimilis Bode 1953 fly
Diptera -	Amianta eurycephala Amianta eurycephala Bode 1953 fly
Diptera - Ptychopteridae	"Acritorhyphus ramosus" = Eoptychoptera simplex "Acritorhyphus ramosus" = Eoptychoptera simplex Handlirsch 1906 phantom crane fly
Diptera -	Culiciscolex gibberatus Culiciscolex gibberatus Bode 1953 fly
Diptera - Limoniidae	"Paratipula formosa" = Architipula formosa, Architipula bodeisimilis, Ozotipula tarda "Paratipula formosa" = Architipula formosa Bode 1953 crane fly Architipula bodeisimilis Bode 1953 crane fly Ozotipula tarda Bode 1953 crane fly
Diptera -	Amblylexis gibberata, "Ellipes laesa" = Ellipibodus laesa Amblylexis gibberata Bode 1953 fly "Ellipes laesa" = Ellipibodus laesa Bode 1953 fly

¹Bode 1905; ²Ponomarenko 1995