Soyana (PIN collection 2455) (Permian of Russian Federation)

Also known as Iva-gora

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified, fine-grained sandstone

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Gundersen in 1930; reposited in the PIN

Primary reference: A. V. Martynov. 1932. New Permian Palaeoptera with the discussion of some problems of their evolution. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 1:1-44 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111187: authorized by Matthew Clapham, entered by Jered Karr on 20.06.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Protelytroptera -
Arctocoleus ivensis5 Martynov 1932 winged insect
PIN 2455/33
 Miomoptera - Palaeomanteidae
Archisialis praecox n. gen. n. sp.5
Archisialis praecox n. gen. n. sp.5 Martynov 1932 winged insect
PIN 2455/31
 Hypoperlida - Hypoperlidae
Boreopsocus danksae10 Shcherbakov 1994 winged insect
PIN 2455/258
 Hemiptera - Coleoscytidae
Coleoscyta rotundata3 Martynov 1935 planthopper
PIN 2455/310 (C. venosa in Martynov, 1935)
Coleoscytodes venosa n. gen. n. sp.7 Martynov 1935 planthopper
PIN 2455/299 (310 is Coleoscyta rotundata)
 Hemiptera - Scytinopteridae
Scytinoptera sp.7 Handlirsch 1904 true bug
PIN 2455/59
Scytinoptera reducta2 Martynov 1928 true bug
PIN 2455/176A
 Hemiptera - Archescytinidae
"Permoscytina tetraclada n. sp." = Archescytina tetraclada6, "Ivascytina difficilis n. gen. n. sp." = Sojanoscytina difficilis6
"Permoscytina tetraclada n. sp." = Archescytina tetraclada6 Martynov 1933 true bug
PIN 2455/45
"Ivascytina difficilis n. gen. n. sp." = Sojanoscytina difficilis6 Martynov 1933 true bug
PIN 2455/44
 Hemiptera - Prosbolopseidae
Scytoneura elliptica n. gen. n. sp.7, Ivaia indistincta n. gen. n. sp.3, "Prosbolopsis simplex" = Prosbolopsis ovalis7
Scytoneura elliptica n. gen. n. sp.7 Martynov 1935 true bug
PIN 2455/267
Ivaia indistincta n. gen. n. sp.3 Becker-Migdisova 1960 true bug
PIN 2455/346, x
"Prosbolopsis simplex" = Prosbolopsis ovalis7 Martynov 1935 true bug
PIN 2455/304
 Hemiptera - Prosbolidae
Permocicada "sp 2"7 Martynov 1928 true bug
PIN 2455/55
Permocicada subtruncata n. sp.7 Martynov 1935 true bug
PIN 2455/189
Permocicada nigronervosa1 Martynov 1928 true bug
PIN 2455/224
Permocicada borealis n. sp.7 Martynov 1935 true bug
PIN 2455/60
"Permocicada angustata n. sp." = Permocicadopsis angustata7 Martynov 1935 true bug
PIN 2455/302
Sojanoneura bimaculata7 Martynov 1935 true bug
PIN 2455/170, 213
Sojanoneura dubiosa n. sp.7 Martynov 1935 true bug
PIN 2455/62
Sojanoneura furcatula n. sp.7 Martynov 1935 true bug
PIN 2455/229
Sojanoneura triangularis n. sp.7 Martynov 1935 true bug
PIN 2455/65
Sojanoneura signata n. sp.7 Martynov 1935 true bug
PIN 2455/57, 58
Prosbole "sp 3"7 Handlirsch 1904 true bug
PIN 2455/56, 53
Prosbole "sp 4"7 Handlirsch 1904 true bug
PIN 2455/52
Prosbole dilatata n. sp.7 Martynov 1935 true bug
PIN 2455/292
 Paoliida - Anthracoptilidae
Strephoneura tenebrosa n. sp.9 Novokshonov and Aristov 2004 winged insect
PIN 2455/339, 340
 Mecoptera - Permocentropidae
Permocentropus philopotamoides n. gen. n. sp.4 Martynov 1932 scorpionfly
PIN 2455/146
 Mecoptera - Permochoristidae
Petromantis sp.4, "Petromantis rossica" = Petromantis rossica8, "Petromantis borealis" = Petromantis rossica4, "Petromantis punctata n. sp." = Petromantis rossica4, "Permochorista venosa n. sp." = Petromantis maculipennis4, "Permochorista venosa" = Petromantis maculipennis8, "Permochorista nigromaculata n. sp." = Petromantis maculipennis4, "? Permochorista stigmata n. sp." = Petromantis maculipennis4, Petrochorista distincta n. sp.4, "Permochorista sp." = Mesochorista4, "Agetochorista ornata" = Agetopanorpa ornata4, "Agetochorista umbrata n. sp." = Agetopanorpa ornata4, "Agetochorista splendida n. sp." = Agetopanorpa ornata4, "Agetochorista reducta n. sp." = Agetopanorpa dubia4, "Agetochorista sp." = Agetopanorpa4, "Agetochorista fasciata n. sp." = Agetopanorpa fasciata4, Oochorista gunderseni n. gen. n. sp.4
Petromantis sp.4 Handlirsch 1904 scorpionfly
PIN 2455/330
"Petromantis rossica" = Petromantis rossica8 Handlirsch 1904 scorpionfly
PIN 2455/172, 244, 245, 246, 3032
"Petromantis borealis" = Petromantis rossica4 Handlirsch 1904 scorpionfly
PIN 2445/174, 120, 348, 167; 2445/86, 175 in Martynova, 1948
"Petromantis punctata n. sp." = Petromantis rossica4 Handlirsch 1904 scorpionfly
PIN 2455/29
"Permochorista venosa n. sp." = Petromantis maculipennis4 Martynov 1932 scorpionfly
PIN 2455/329
"Permochorista venosa" = Petromantis maculipennis8 Martynov 1932 scorpionfly
PIN 2455/157
"Permochorista nigromaculata n. sp." = Petromantis maculipennis4 Martynov 1932 scorpionfly
PIN 2455/332
"? Permochorista stigmata n. sp." = Petromantis maculipennis4 Martynov 1932 scorpionfly
PIN 2455/185
Petrochorista distincta n. sp.4 Martynov 1932 scorpionfly
PIN 2455/331
"Permochorista sp." = Mesochorista4 Tillyard 1916 scorpionfly
PIN 2455/268
"Agetochorista ornata" = Agetopanorpa ornata4 Martynov 1932 scorpionfly
PIN 2455/147
"Agetochorista umbrata n. sp." = Agetopanorpa ornata4 Martynov 1932 scorpionfly
PIN 2455/43
"Agetochorista splendida n. sp." = Agetopanorpa ornata4 Martynov 1932 scorpionfly
PIN 2445/343
"Agetochorista reducta n. sp." = Agetopanorpa dubia4 Martynov 1932 scorpionfly
PIN 2445/10
"Agetochorista sp." = Agetopanorpa4 Carpenter 1930 scorpionfly
"Agetochorista fasciata n. sp." = Agetopanorpa fasciata4 Martynov 1932 scorpionfly
PIN 2455/27
Oochorista gunderseni n. gen. n. sp.4 Martynov 1932 scorpionfly
PIN 2445/347
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis n. sp.4 Martynov 1932 winged insect
PIN 2455/87
 Dicondylia - Permithonidae
Permithonopsis sp.5 Martynov 1932 winged insect
PIN 2455/117
Permithonopsis ivensis n. gen. n. sp.5 Martynov 1932 winged insect
PIN 2455/37
Eopsychops sojanensis n. gen. n. sp.5 Martynov 1932 winged insect
PIN 2455/184
Permosisyra punctata n. sp.5 Martynov 1932 winged insect
PIN 2455/9
 Megasecoptera - Protohymenidae
Ivahymen constrictus n. gen. n. sp.
Ivahymen constrictus n. gen. n. sp. Martynov 1932 winged insect
PIN 2455/5
 Palaeodictyoptera - Eukulojidae
"Kuloja cubitalis n. sp." = Eukuloja cubitalis
"Kuloja cubitalis n. sp." = Eukuloja cubitalis Martynov 1932 winged insect
PIN 2455/3, 4; 2455/291 in Sinitshenkova 1981
 Dicondylia - Permagrionidae
Permolestes gracilis n. gen. n. sp.
Permolestes gracilis n. gen. n. sp. Martynov 1932 winged insect
PIN 2455/1,2 One specimen