Soyana (PIN collection 2455) (Permian of Russian Federation)

Also known as Iva-gora

Where: Arkangelsk, Russian Federation (65.6° N, 43.7° E: paleocoordinates 33.2° N, 33.1° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified, fine-grained sandstone

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Gundersen in 1930; reposited in the PIN

Primary reference: A. V. Martynov. 1932. New Permian Palaeoptera with the discussion of some problems of their evolution. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 1:1-44 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111187: authorized by Matthew Clapham, entered by Jered Karr on 20.06.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Megasecoptera - Protohymenidae
Ivahymen constrictus n. gen. n. sp.
Ivahymen constrictus n. gen. n. sp. Martynov 1932 winged insect
PIN 2455/5
 Palaeodictyoptera - Eukulojidae
"Kuloja cubitalis n. sp." = Eukuloja cubitalis
"Kuloja cubitalis n. sp." = Eukuloja cubitalis Martynov 1932 winged insect
PIN 2455/3, 4; 2455/291 in Sinitshenkova 1981
 Hypoperlida - Hypoperlidae
Boreopsocus danksae11 Shcherbakov 1994 winged insect
PIN 2455/258
 Hemiptera - Archescytinidae
"Ivascytina difficilis n. gen. n. sp." = Sojanoscytina difficilis7, "Permoscytina tetraclada n. sp." = Archescytina tetraclada7
"Ivascytina difficilis n. gen. n. sp." = Sojanoscytina difficilis7 Martynov 1933 true bug
PIN 2455/44
"Permoscytina tetraclada n. sp." = Archescytina tetraclada7 Martynov 1933 true bug
PIN 2455/45
 Hemiptera - Scytinopteridae
Scytinoptera sp.8 Handlirsch 1904 true bug
PIN 2455/59
Scytinoptera reducta3 Martynov 1928 true bug
PIN 2455/176A
 Hemiptera - Coleoscytidae
Coleoscytodes venosa n. gen. n. sp.8 Martynov 1935 planthopper
PIN 2455/299 (310 is Coleoscyta rotundata)
Coleoscyta rotundata4 Martynov 1935 planthopper
PIN 2455/310 (C. venosa in Martynov, 1935)
 Hemiptera - Prosbolopseidae
"Prosbolopsis simplex" = Prosbolopsis ovalis8, Scytoneura elliptica n. gen. n. sp.8, Ivaia indistincta n. gen. n. sp.4
"Prosbolopsis simplex" = Prosbolopsis ovalis8 Martynov 1935 true bug
PIN 2455/304
Scytoneura elliptica n. gen. n. sp.8 Martynov 1935 true bug
PIN 2455/267
Ivaia indistincta n. gen. n. sp.4 Becker-Migdisova 1960 true bug
PIN 2455/346, x
 Hemiptera - Prosbolidae
Prosbole "sp 3"8 Handlirsch 1904 true bug
PIN 2455/56, 53
Prosbole "sp 4"8 Handlirsch 1904 true bug
PIN 2455/52
Prosbole dilatata n. sp.8 Martynov 1935 true bug
PIN 2455/292
"Permocicada angustata n. sp." = Permocicadopsis angustata8 Martynov 1935 true bug
PIN 2455/302
Permocicada "sp 2"8 Martynov 1928 true bug
PIN 2455/55
Permocicada subtruncata n. sp.8 Martynov 1935 true bug
PIN 2455/189
Permocicada borealis n. sp.8 Martynov 1935 true bug
PIN 2455/60
Permocicada nigronervosa2 Martynov 1928 true bug
PIN 2455/224
Sojanoneura signata n. sp.8 Martynov 1935 true bug
PIN 2455/57, 58
Sojanoneura dubiosa n. sp.8 Martynov 1935 true bug
PIN 2455/62
Sojanoneura triangularis n. sp.8 Martynov 1935 true bug
PIN 2455/65
Sojanoneura furcatula n. sp.8 Martynov 1935 true bug
PIN 2455/229
Sojanoneura bimaculata8 Martynov 1935 true bug
PIN 2455/170, 213
 Miomoptera - Palaeomanteidae
Archisialis praecox n. gen. n. sp.6
Archisialis praecox n. gen. n. sp.6 Martynov 1932 winged insect
PIN 2455/31
 Protelytroptera -
Arctocoleus ivensis6 Martynov 1932 winged insect
PIN 2455/33
 Mecoptera - Permochoristidae
Petromantis sp.5, "? Permochorista stigmata n. sp." = Petromantis maculipennis5, "Permochorista nigromaculata n. sp." = Petromantis maculipennis5, "Permochorista venosa n. sp." = Petromantis maculipennis5, "Permochorista venosa" = Petromantis maculipennis9, "Petromantis rossica" = Petromantis rossica9, "Petromantis punctata n. sp." = Petromantis rossica5, "Petromantis borealis" = Petromantis rossica5, Petrochorista distincta n. sp.5, "Permochorista sp." = Mesochorista5, "Agetochorista ornata" = Agetopanorpa ornata5, "Agetochorista umbrata n. sp." = Agetopanorpa ornata5, "Agetochorista splendida n. sp." = Agetopanorpa ornata5, "Agetochorista reducta n. sp." = Agetopanorpa dubia5, "Agetochorista sp." = Agetopanorpa5, "Agetochorista fasciata n. sp." = Agetopanorpa fasciata5, Oochorista gunderseni n. gen. n. sp.5
Petromantis sp.5 Handlirsch 1904 scorpionfly
PIN 2455/330
"? Permochorista stigmata n. sp." = Petromantis maculipennis5 Martynov 1932 scorpionfly
PIN 2455/185
"Permochorista nigromaculata n. sp." = Petromantis maculipennis5 Martynov 1932 scorpionfly
PIN 2455/332
"Permochorista venosa n. sp." = Petromantis maculipennis5 Martynov 1932 scorpionfly
PIN 2455/329
"Permochorista venosa" = Petromantis maculipennis9 Martynov 1932 scorpionfly
PIN 2455/157
"Petromantis rossica" = Petromantis rossica9 Handlirsch 1904 scorpionfly
PIN 2455/172, 244, 245, 246, 3032
"Petromantis punctata n. sp." = Petromantis rossica5 Handlirsch 1904 scorpionfly
PIN 2455/29
"Petromantis borealis" = Petromantis rossica5 Handlirsch 1904 scorpionfly
PIN 2445/174, 120, 348, 167; 2445/86, 175 in Martynova, 1948
Petrochorista distincta n. sp.5 Martynov 1932 scorpionfly
PIN 2455/331
"Permochorista sp." = Mesochorista5 Tillyard 1916 scorpionfly
PIN 2455/268
"Agetochorista ornata" = Agetopanorpa ornata5 Martynov 1932 scorpionfly
PIN 2455/147
"Agetochorista umbrata n. sp." = Agetopanorpa ornata5 Martynov 1932 scorpionfly
PIN 2455/43
"Agetochorista splendida n. sp." = Agetopanorpa ornata5 Martynov 1932 scorpionfly
PIN 2445/343
"Agetochorista reducta n. sp." = Agetopanorpa dubia5 Martynov 1932 scorpionfly
PIN 2445/10
"Agetochorista sp." = Agetopanorpa5 Carpenter 1930 scorpionfly
"Agetochorista fasciata n. sp." = Agetopanorpa fasciata5 Martynov 1932 scorpionfly
PIN 2455/27
Oochorista gunderseni n. gen. n. sp.5 Martynov 1932 scorpionfly
PIN 2445/347
 Mecoptera - Permocentropidae
Permocentropus philopotamoides n. gen. n. sp.5 Martynov 1932 scorpionfly
PIN 2455/146
 Permotrichoptera - Microptysmatidae
Kamopanorpa rotundipennis n. sp.5 Martynov 1932 winged insect
PIN 2455/87
 Dicondylia - Permithonidae
Permithonopsis sp.6 Martynov 1932 winged insect
PIN 2455/117
Permithonopsis ivensis n. gen. n. sp.6 Martynov 1932 winged insect
PIN 2455/37
Eopsychops sojanensis n. gen. n. sp.6 Martynov 1932 winged insect
PIN 2455/184
Permosisyra punctata n. sp.6 Martynov 1932 winged insect
PIN 2455/9
 Paoliida - Blattinopsidae
Glaphyrophlebia iva n. sp.1 Aristov and Rasnitsyn 2021 winged insect
PIN 2455/128
 Paoliida - Anthracoptilidae
Strephoneura tenebrosa n. sp.10 Novokshonov and Aristov 2004 winged insect
PIN 2455/339, 340
 Dicondylia - Permagrionidae
Permolestes gracilis n. gen. n. sp.
Permolestes gracilis n. gen. n. sp. Martynov 1932 winged insect
PIN 2455/1,2 One specimen