Elmo, MCZ 1927 collection (Permian of the United States)

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.6° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• The Elmo limestone itself is a chalky, soft, almost white deposit, about five feet thick; only the very bottom, more massive layer of this stratum contains insects.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by F.M. Carpenter, J.W. Wilson, W.S. Creighton in 1927; reposited in the MCZ

Collection methods: quarrying,

Primary reference: F. M. Carpenter. 1930. The Lower Permian Insects of Kansas: Part 1. Introduction and the Order Mecoptera. Bulletin of the Museum of Comparative Zoology 70(2):69-101 [C. Labandeira/C. Labandeira/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111217: authorized by Matthew Clapham, entered by Matthew Clapham on 21.06.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Eoblattida -
Elmonympha carpenteri n. gen. n. sp.12
Elmonympha carpenteri n. gen. n. sp.12 Aristov 2019 insect
MCZ 3622 (year not stated)
 Diaphanopterodea - Asthenohymenidae
Asthenohymen dunbari5, "Doter pusillus" = Asthenohymen pusillus1, "Doter affinis" = Asthenohymen affinis1
Asthenohymen dunbari5 Tillyard 1924 winged insect
MCZ 3059,3074-94,3111-3119
"Doter pusillus" = Asthenohymen pusillus1 Tillyard 1926 winged insect
MCZ 3097
"Doter affinis" = Asthenohymen affinis1 Tillyard 1926 winged insect
MCZ 3095
 Megasecoptera - Protohymenidae
Permohymen schucherti1 Tillyard 1924 winged insect
MCZ 3071-4
Protohymen permianus1 Tillyard 1924 winged insect
MCZ 3060-8,3070
Protohymen elongatus n. sp.1 Carpenter 1930 winged insect
MCZ 3069
 Palaeodictyoptera - Spilapteridae
Dunbaria fasciipennis2 Tillyard 1924 winged insect
MCZ 3056-8
 Palaeodictyoptera - Elmoboriidae
Elmoboria piperi n. gen. n. sp.10
Elmoboria piperi n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5839
 Dicondylia - Gelasopteridae
Gelasopteron gracile n. gen. n. sp.10
Gelasopteron gracile n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 7511
 Dicondylia - Apheloneuridae
Apheloneura amplia n. sp.10, Apheloneura minutissima n. gen. n. sp.10
Apheloneura amplia n. sp.10 Carpenter 1976 winged insect
MCZ 8604, 7525, 7522
Apheloneura minutissima n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 7527, 7534
 Protelytroptera - Elytroneuridae
Elytroneura permiana5 Carpenter 1933 winged insect
 Orthoptera - Pruvostitidae
Paroedischia recta n. gen. n. sp.9
Paroedischia recta n. gen. n. sp.9 Carpenter 1966 grasshopper
MCZ 5897 (type from here; 5 specimens in total)
 Paoliida - Anthracoptilidae
Homocladus grandis n. gen. n. sp.9
Homocladus grandis n. gen. n. sp.9 Carpenter 1966 winged insect
 Paoliida - Blattinopsidae
Glaphyrophlebia ovata9 Sellards 1909 winged insect
MCZ 4965
Glaphyrophlebia speciosa9 Sellards 1909 winged insect
MCZ 6303
 Grylloblattodea - Liomopteridae
Liomopterum ornatum8 Sellards 1909 ice crawler
36 from lower limestone, at least 1 from 1927
 Grylloblattodea - Lemmatophoridae
Paraprisca grandis n. sp.6 Carpenter 1935 ice crawler
MCZ 3584,5
Paraprisca fragilis6 Sellards 1909 ice crawler
36 total from 1927 and 1932
Lecorium elongatum6 Sellards 1909 ice crawler
46 total, at least 1 from 1927
Lemmatophora typa6 Sellards 1909 ice crawler
More than 200 total, but years not given
Artinska ovata6 Sellards 1909 ice crawler
76 total, minus 21 collected in 1932
Artinska clara6 Sellards 1909 ice crawler
160 total, minus 64 collected in 1932
Lisca minuta6 Sellards 1909 ice crawler
104 total, minus 42 collected in 1932
 Dicondylia - Nugonioneuridae
Nugonioneura problematica10 Tillyard 1937 winged insect
 Cnemidolestodea - Psoropteridae
Psoroptera cubitalia n. gen. n. sp.10
Psoroptera cubitalia n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5840
 Cnemidolestodea - Tillyardembiidae
Heteroptilon costale n. gen. n. sp.10
Heteroptilon costale n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5878
 Cnemidolestodea - Tococladidae
"Opisthocladus strictus n. sp." = Elmocladus strictus10
"Opisthocladus strictus n. sp." = Elmocladus strictus10 Carpenter 1976 winged insect
MCZ 5881
 Cnemidolestodea - Parmapteridae
Opisthocladus arcuatus n. gen. n. sp.10
Opisthocladus arcuatus n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5882
 Caloneurodea - Caloneuridae
"Plesiogramma reducta n. sp." = Pleisiogramma medialis7
"Plesiogramma reducta n. sp." = Pleisiogramma medialis7 Carpenter 1943 winged insect
 Dicondylia - Trachopterygidae
Trachopteryx martynovi n. gen. n. sp.10
Trachopteryx martynovi n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 7497
 Permopsocida - Permopsocidae
Lithopsocidium permianum n. gen. n. sp.4 Carpenter 1932 winged insect
MCZ 3168
Permopsocus latipennis4 Tillyard 1926 winged insect
MCZ 3155,3181,2,4,5,8
Orthopsocus singularis n. gen. n. sp.4 Carpenter 1932 winged insect
MCZ 3200
Progonopsocus permianus4 Tillyard 1926 winged insect
MCZ 3175-7
 Permopsocida - Psocidiidae
Dichentomum parvulum n. sp.4 Carpenter 1932 winged insect
MCZ 3173
Dichentomum tinctum4 Tillyard 1926 winged insect
Over 100 specimens including MCZ 3138,3141-7,3162-5
Dichentomum minimum4 Tillyard 1926 winged insect
MCZ 3167 and 3171 are best
Dichentomum latum n. sp.4 Carpenter 1932 winged insect
MCZ 3160
 Lophioneurida - Lophioneuridae
"Cyphoneura reducta n. sp." = Cyphoneurodes reducta4, Cyphoneura permiana n. gen. n. sp.4
"Cyphoneura reducta n. sp." = Cyphoneurodes reducta4 Carpenter 1932 winged insect
MCZ 3198
Cyphoneura permiana n. gen. n. sp.4 Carpenter 1932 winged insect
MCZ 3197
 Hemiptera - Archescytinidae
"Permoscytina kansasensis" = Archescytina kansasensis3, Archescytina permiana3, "Permoscytina muiri" = Archescytina muiri3, Paleoscytina brevistigma n. gen. n. sp.3, Permopsylla permiana n. sp.3, Permopsylla americana3, Permopsylla minuta n. sp.3
"Permoscytina kansasensis" = Archescytina kansasensis3 Tillyard 1926 true bug
MCZ 3121
Archescytina permiana3 Tillyard 1926 true bug
MCZ 3120,3136-7
"Permoscytina muiri" = Archescytina muiri3 Tillyard 1926 true bug
MCZ 3122
Paleoscytina brevistigma n. gen. n. sp.3 Carpenter 1931 true bug
MCZ 3133
Permopsylla permiana n. sp.3 Carpenter 1931 true bug
MCZ 3129-31
Permopsylla americana3 Tillyard 1926 true bug
MCZ 3123-3125,?3126,?3128,?3134
Permopsylla minuta n. sp.3 Carpenter 1931 true bug
MCZ 3132
 Dicondylia - Permembiidae
Permembia delicatula10 Tillyard 1928 winged insect
MCZ 7526,7539,7547,7596,7474
 Mecoptera - Permopanorpidae
Permopanorpa inaequalis Tillyard 1926 scorpionfly
MCZ 3009-3033
 Mecoptera - Permochoristidae
Protopanorpa permiana Tillyard 1926 scorpionfly
MCZ 3034-5
Agetopanorpa maculata n. gen. n. sp. Carpenter 1930 scorpionfly
MCZ 3037
Protochorista tetraclada Tillyard 1926 scorpionfly
MCZ 3036
 Protomeroptera - Protomeropidae
Platychorista venosa Tillyard 1926 winged insect
MCZ 3001-8
 Dicondylia - Permithonidae
Elmothone martynovae n. gen. n. sp.10
Elmothone martynovae n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5585
 Meganisoptera - Meganeuridae
Oligotypus sp.2, Oligotypus tillyardi n. gen. n. sp.2, "Typus permianus" = Tupus permianus2, Megatypus schucherti2
Oligotypus sp.2 Carpenter 1931 griffinfly
MCZ 3041
Oligotypus tillyardi n. gen. n. sp.2 Carpenter 1931 griffinfly
MCZ 3049
"Typus permianus" = Tupus permianus2 Sellards 1906 griffinfly
MCZ 3050-5
Megatypus schucherti2 Tillyard 1925 griffinfly
MCZ 3048
 Dicondylia - Ditaxineuridae
Ditaxineura anomalostigma2 Tillyard 1926 winged insect
MCZ 3043-6
 Dicondylia - Progoneuridae
Progoneura minuta n. gen. n. sp.2
Progoneura minuta n. gen. n. sp.2 Carpenter 1931 winged insect
MCZ 3042
 Dicondylia - Permoneuridae
Permoneura lameerei n. gen. n. sp.3
Permoneura lameerei n. gen. n. sp.3 Carpenter 1931 winged insect
MCZ 3135
Xiphosura
 Xiphosurida - Paleolimulidae
"Paleolimulus avitus" = Paleolimulus signatus11
"Paleolimulus avitus" = Paleolimulus signatus11 Beecher 1904