PBDB Collection

Elmo, MCZ 1927 collection (Permian of the United States)

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.6° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• The Elmo limestone itself is a chalky, soft, almost white deposit, about five feet thick; only the very bottom, more massive layer of this stratum contains insects.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by F.M. Carpenter, J.W. Wilson, W.S. Creighton in 1927; reposited in the MCZ

Collection methods: quarrying,

Primary reference: F. M. Carpenter. 1930. The Lower Permian Insects of Kansas: Part 1. Introduction and the Order Mecoptera. Bulletin of the Museum of Comparative Zoology 70(2):69-101 [C. Labandeira/C. Labandeira/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111217: authorized by Matthew Clapham, entered by Matthew Clapham on 21.06.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Eoblattida -	Elmonympha carpenteri n. gen. n. sp.¹² Elmonympha carpenteri n. gen. n. sp.¹² Aristov 2019 insect MCZ 3622 (year not stated)
Meganisoptera - Meganeuridae	Oligotypus sp.², Oligotypus tillyardi n. gen. n. sp.², "Typus permianus" = Tupus permianus², Megatypus schucherti² Oligotypus sp.² Carpenter 1931 griffinfly MCZ 3041 Oligotypus tillyardi n. gen. n. sp.² Carpenter 1931 griffinfly MCZ 3049 "Typus permianus" = Tupus permianus² Sellards 1906 griffinfly MCZ 3050-5 Megatypus schucherti² Tillyard 1925 griffinfly MCZ 3048
Dicondylia - Ditaxineuridae	Ditaxineura anomalostigma² Ditaxineura anomalostigma² Tillyard 1926 winged insect MCZ 3043-6
Dicondylia - Progoneuridae	Progoneura minuta n. gen. n. sp.² Progoneura minuta n. gen. n. sp.² Carpenter 1931 winged insect MCZ 3042
Protelytroptera - Elytroneuridae	Elytroneura permiana⁵ Elytroneura permiana⁵ Carpenter 1933 winged insect
Dicondylia - Apheloneuridae	Apheloneura amplia n. sp.¹⁰, Apheloneura minutissima n. gen. n. sp.¹⁰ Apheloneura amplia n. sp.¹⁰ Carpenter 1976 winged insect MCZ 8604, 7525, 7522 Apheloneura minutissima n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 7527, 7534
Dicondylia - Permembiidae	Permembia delicatula¹⁰ Permembia delicatula¹⁰ Tillyard 1928 winged insect MCZ 7526,7539,7547,7596,7474
Dicondylia - Gelasopteridae	Gelasopteron gracile n. gen. n. sp.¹⁰ Gelasopteron gracile n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 7511
Dicondylia - Trachopterygidae	Trachopteryx martynovi n. gen. n. sp.¹⁰ Trachopteryx martynovi n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 7497
Permopsocida - Psocidiidae	Dichentomum parvulum n. sp.⁴, Dichentomum minimum⁴, Dichentomum tinctum⁴, Dichentomum latum n. sp.⁴ Dichentomum parvulum n. sp.⁴ Carpenter 1932 winged insect MCZ 3173 Dichentomum minimum⁴ Tillyard 1926 winged insect MCZ 3167 and 3171 are best Dichentomum tinctum⁴ Tillyard 1926 winged insect Over 100 specimens including MCZ 3138,3141-7,3162-5 Dichentomum latum n. sp.⁴ Carpenter 1932 winged insect MCZ 3160
Permopsocida - Permopsocidae	Orthopsocus singularis n. gen. n. sp.⁴, Lithopsocidium permianum n. gen. n. sp.⁴, Permopsocus latipennis⁴, Progonopsocus permianus⁴ Orthopsocus singularis n. gen. n. sp.⁴ Carpenter 1932 winged insect MCZ 3200 Lithopsocidium permianum n. gen. n. sp.⁴ Carpenter 1932 winged insect MCZ 3168 Permopsocus latipennis⁴ Tillyard 1926 winged insect MCZ 3155,3181,2,4,5,8 Progonopsocus permianus⁴ Tillyard 1926 winged insect MCZ 3175-7
Lophioneurida - Lophioneuridae	"Cyphoneura reducta n. sp." = Cyphoneurodes reducta⁴, Cyphoneura permiana n. gen. n. sp.⁴ "Cyphoneura reducta n. sp." = Cyphoneurodes reducta⁴ Carpenter 1932 winged insect MCZ 3198 Cyphoneura permiana n. gen. n. sp.⁴ Carpenter 1932 winged insect MCZ 3197
Hemiptera - Archescytinidae	Paleoscytina brevistigma n. gen. n. sp.³, "Permoscytina muiri" = Archescytina muiri³, Archescytina permiana³, "Permoscytina kansasensis" = Archescytina kansasensis³, Permopsylla americana³, Permopsylla permiana n. sp.³, Permopsylla minuta n. sp.³ Paleoscytina brevistigma n. gen. n. sp.³ Carpenter 1931 true bug MCZ 3133 "Permoscytina muiri" = Archescytina muiri³ Tillyard 1926 true bug MCZ 3122 Archescytina permiana³ Tillyard 1926 true bug MCZ 3120,3136-7 "Permoscytina kansasensis" = Archescytina kansasensis³ Tillyard 1926 true bug MCZ 3121 Permopsylla americana³ Tillyard 1926 true bug MCZ 3123-3125,?3126,?3128,?3134 Permopsylla permiana n. sp.³ Carpenter 1931 true bug MCZ 3129-31 Permopsylla minuta n. sp.³ Carpenter 1931 true bug MCZ 3132
Grylloblattodea - Liomopteridae	Liomopterum ornatum⁸ Liomopterum ornatum⁸ Sellards 1909 ice crawler 36 from lower limestone, at least 1 from 1927
Grylloblattodea - Lemmatophoridae	Lemmatophora typa⁶, Artinska clara⁶, Artinska ovata⁶, Lisca minuta⁶, Lecorium elongatum⁶, Paraprisca fragilis⁶, Paraprisca grandis n. sp.⁶ Lemmatophora typa⁶ Sellards 1909 ice crawler More than 200 total, but years not given Artinska clara⁶ Sellards 1909 ice crawler 160 total, minus 64 collected in 1932 Artinska ovata⁶ Sellards 1909 ice crawler 76 total, minus 21 collected in 1932 Lisca minuta⁶ Sellards 1909 ice crawler 104 total, minus 42 collected in 1932 Lecorium elongatum⁶ Sellards 1909 ice crawler 46 total, at least 1 from 1927 Paraprisca fragilis⁶ Sellards 1909 ice crawler 36 total from 1927 and 1932 Paraprisca grandis n. sp.⁶ Carpenter 1935 ice crawler MCZ 3584,5
Orthoptera - Pruvostitidae	Paroedischia recta n. gen. n. sp.⁹ Paroedischia recta n. gen. n. sp.⁹ Carpenter 1966 grasshopper MCZ 5897 (type from here; 5 specimens in total)
Dicondylia - Nugonioneuridae	Nugonioneura problematica¹⁰ Nugonioneura problematica¹⁰ Tillyard 1937 winged insect
Cnemidolestodea - Tococladidae	"Opisthocladus strictus n. sp." = Elmocladus strictus¹⁰ "Opisthocladus strictus n. sp." = Elmocladus strictus¹⁰ Carpenter 1976 winged insect MCZ 5881
Cnemidolestodea - Parmapteridae	Opisthocladus arcuatus n. gen. n. sp.¹⁰ Opisthocladus arcuatus n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 5882
Cnemidolestodea - Psoropteridae	Psoroptera cubitalia n. gen. n. sp.¹⁰ Psoroptera cubitalia n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 5840
Cnemidolestodea - Tillyardembiidae	Heteroptilon costale n. gen. n. sp.¹⁰ Heteroptilon costale n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 5878
Caloneurodea - Caloneuridae	"Plesiogramma reducta n. sp." = Pleisiogramma medialis⁷ "Plesiogramma reducta n. sp." = Pleisiogramma medialis⁷ Carpenter 1943 winged insect
Paoliida - Anthracoptilidae	Homocladus grandis n. gen. n. sp.⁹ Homocladus grandis n. gen. n. sp.⁹ Carpenter 1966 winged insect
Paoliida - Blattinopsidae	Glaphyrophlebia ovata⁹, Glaphyrophlebia speciosa⁹ Glaphyrophlebia ovata⁹ Sellards 1909 winged insect MCZ 4965 Glaphyrophlebia speciosa⁹ Sellards 1909 winged insect MCZ 6303
Protomeroptera - Protomeropidae	Platychorista venosa Platychorista venosa Tillyard 1926 winged insect MCZ 3001-8
Mecoptera - Permopanorpidae	Permopanorpa inaequalis Permopanorpa inaequalis Tillyard 1926 scorpionfly MCZ 3009-3033
Mecoptera - Permochoristidae	Protochorista tetraclada, Agetopanorpa maculata n. gen. n. sp., Protopanorpa permiana Protochorista tetraclada Tillyard 1926 scorpionfly MCZ 3036 Agetopanorpa maculata n. gen. n. sp. Carpenter 1930 scorpionfly MCZ 3037 Protopanorpa permiana Tillyard 1926 scorpionfly MCZ 3034-5
Dicondylia - Permithonidae	Elmothone martynovae n. gen. n. sp.¹⁰ Elmothone martynovae n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 5585
Diaphanopterodea - Asthenohymenidae	"Doter affinis" = Asthenohymen affinis¹, Asthenohymen dunbari⁵, "Doter pusillus" = Asthenohymen pusillus¹ "Doter affinis" = Asthenohymen affinis¹ Tillyard 1926 winged insect MCZ 3095 Asthenohymen dunbari⁵ Tillyard 1924 winged insect MCZ 3059,3074-94,3111-3119 "Doter pusillus" = Asthenohymen pusillus¹ Tillyard 1926 winged insect MCZ 3097
Megasecoptera - Protohymenidae	Protohymen permianus¹, Protohymen elongatus n. sp.¹, Permohymen schucherti¹ Protohymen permianus¹ Tillyard 1924 winged insect MCZ 3060-8,3070 Protohymen elongatus n. sp.¹ Carpenter 1930 winged insect MCZ 3069 Permohymen schucherti¹ Tillyard 1924 winged insect MCZ 3071-4
Palaeodictyoptera - Spilapteridae	Dunbaria fasciipennis² Dunbaria fasciipennis² Tillyard 1924 winged insect MCZ 3056-8
Palaeodictyoptera - Elmoboriidae	Elmoboria piperi n. gen. n. sp.¹⁰ Elmoboria piperi n. gen. n. sp.¹⁰ Carpenter 1976 winged insect MCZ 5839
Dicondylia - Permoneuridae	Permoneura lameerei n. gen. n. sp.³ Permoneura lameerei n. gen. n. sp.³ Carpenter 1931 winged insect MCZ 3135
Xiphosura
Xiphosurida - Paleolimulidae	"Paleolimulus avitus" = Paleolimulus signatus¹¹ "Paleolimulus avitus" = Paleolimulus signatus¹¹ Beecher 1904

¹Carpenter 1930; ²Carpenter 1931; ³Carpenter 1931; ⁴Carpenter 1932; ⁵Carpenter 1933; ⁶Carpenter 1935; ⁷Carpenter 1943; ⁸Carpenter 1950; ⁹Carpenter 1966; ¹⁰Carpenter 1976; ¹¹Raymond 1944; ¹²Vršanský et al. 2019