Elmo, MCZ 1927 collection (Permian of the United States)

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.6° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• The Elmo limestone itself is a chalky, soft, almost white deposit, about five feet thick; only the very bottom, more massive layer of this stratum contains insects.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by F.M. Carpenter, J.W. Wilson, W.S. Creighton in 1927; reposited in the MCZ

Collection methods: quarrying,

Primary reference: F. M. Carpenter. 1930. The Lower Permian Insects of Kansas: Part 1. Introduction and the Order Mecoptera. Bulletin of the Museum of Comparative Zoology 70(2):69-101 [C. Labandeira/C. Labandeira/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111217: authorized by Matthew Clapham, entered by Matthew Clapham on 21.06.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Dicondylia - Permoneuridae
Permoneura lameerei n. gen. n. sp.3
Permoneura lameerei n. gen. n. sp.3 Carpenter 1931 winged insect
MCZ 3135
 Protelytroptera - Elytroneuridae
Elytroneura permiana5 Carpenter 1933 winged insect
 Hemiptera - Archescytinidae
Archescytina permiana3, "Permoscytina muiri" = Archescytina muiri3, "Permoscytina kansasensis" = Archescytina kansasensis3, Permopsylla permiana n. sp.3, Permopsylla americana3, Permopsylla minuta n. sp.3, Paleoscytina brevistigma n. gen. n. sp.3
Archescytina permiana3 Tillyard 1926 true bug
MCZ 3120,3136-7
"Permoscytina muiri" = Archescytina muiri3 Tillyard 1926 true bug
MCZ 3122
"Permoscytina kansasensis" = Archescytina kansasensis3 Tillyard 1926 true bug
MCZ 3121
Permopsylla permiana n. sp.3 Carpenter 1931 true bug
MCZ 3129-31
Permopsylla americana3 Tillyard 1926 true bug
MCZ 3123-3125,?3126,?3128,?3134
Permopsylla minuta n. sp.3 Carpenter 1931 true bug
MCZ 3132
Paleoscytina brevistigma n. gen. n. sp.3 Carpenter 1931 true bug
MCZ 3133
 Dicondylia - Lophioneuridae
"Cyphoneura reducta n. sp." = Cyphoneurodes reducta4, Cyphoneura permiana n. gen. n. sp.4
"Cyphoneura reducta n. sp." = Cyphoneurodes reducta4 Carpenter 1932 winged insect
MCZ 3198
Cyphoneura permiana n. gen. n. sp.4 Carpenter 1932 winged insect
MCZ 3197
 Permopsocida - Psocidiidae
Dichentomum minimum4 Tillyard 1926 winged insect
MCZ 3167 and 3171 are best
Dichentomum parvulum n. sp.4 Carpenter 1932 winged insect
MCZ 3173
Dichentomum tinctum4 Tillyard 1926 winged insect
Over 100 specimens including MCZ 3138,3141-7,3162-5
Dichentomum latum n. sp.4 Carpenter 1932 winged insect
MCZ 3160
 Permopsocida - Permopsocidae
Lithopsocidium permianum n. gen. n. sp.4 Carpenter 1932 winged insect
MCZ 3168
Progonopsocus permianus4 Tillyard 1926 winged insect
MCZ 3175-7
Orthopsocus singularis n. gen. n. sp.4 Carpenter 1932 winged insect
MCZ 3200
Permopsocus latipennis4 Tillyard 1926 winged insect
MCZ 3155,3181,2,4,5,8
 Dicondylia - Trachopterygidae
Trachopteryx martynovi n. gen. n. sp.10
Trachopteryx martynovi n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 7497
 Dicondylia - Apheloneuridae
Apheloneura amplia n. sp.10, Apheloneura minutissima n. gen. n. sp.10
Apheloneura amplia n. sp.10 Carpenter 1976 winged insect
MCZ 8604, 7525, 7522
Apheloneura minutissima n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 7527, 7534
 Cnemidolestodea - Psoropteridae
Psoroptera cubitalia n. gen. n. sp.10
Psoroptera cubitalia n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5840
 Cnemidolestodea - Tococladidae
Opisthocladus strictus n. sp.10 Carpenter 1976 winged insect
MCZ 5881
Opisthocladus arcuatus n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5882
 Caloneurodea - Caloneuridae
"Plesiogramma reducta n. sp." = Pleisiogramma medialis7
"Plesiogramma reducta n. sp." = Pleisiogramma medialis7 Carpenter 1943 winged insect
 Protorthoptera - Heteroptilidae
Heteroptilon costale n. gen. n. sp.10
Heteroptilon costale n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5878
 Protorthoptera - Nugonioneuridae
Nugonioneura problematica10 Tillyard 1937 winged insect
 Orthoptera - Pruvostitidae
Paroedischia recta n. gen. n. sp.9
Paroedischia recta n. gen. n. sp.9 Carpenter 1966 grasshopper
MCZ 5897 (type from here; 5 specimens in total)
 Paoliida - Blattinopsidae
Glaphyrophlebia ovata9 Sellards 1909 winged insect
MCZ 4965
Glaphyrophlebia speciosa9 Sellards 1909 winged insect
MCZ 6303
 Paoliida - Anthracoptilidae
Homocladus grandis n. gen. n. sp.9
Homocladus grandis n. gen. n. sp.9 Carpenter 1966 winged insect
 Dicondylia - Permembiidae
Permembia delicatula10 Tillyard 1928 winged insect
MCZ 7526,7539,7547,7596,7474
 Mecoptera - Permochoristidae
Agetopanorpa maculata n. gen. n. sp. Carpenter 1930 scorpionfly
MCZ 3037
Protochorista tetraclada Tillyard 1926 scorpionfly
MCZ 3036
Protopanorpa permiana Tillyard 1926 scorpionfly
MCZ 3034-5
 Mecoptera - Permopanorpidae
Permopanorpa inaequalis Tillyard 1926 scorpionfly
MCZ 3009-3033
 Mecoptera - Protomeropidae
Platychorista venosa Tillyard 1926 scorpionfly
MCZ 3001-8
 Dicondylia - Permithonidae
Elmothone martynovae n. gen. n. sp.10
Elmothone martynovae n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5585
 Dicondylia - Gelasopteridae
Gelasopteron gracile n. gen. n. sp.10
Gelasopteron gracile n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 7511
 Megasecoptera - Protohymenidae
Permohymen schucherti1 Tillyard 1924 winged insect
MCZ 3071-4
Protohymen permianus1 Tillyard 1924 winged insect
MCZ 3060-8,3070
Protohymen elongatus n. sp.1 Carpenter 1930 winged insect
MCZ 3069
 Diaphanopterodea - Asthenohymenidae
"Doter affinis" = Asthenohymen affinis1, "Doter pusillus" = Asthenohymen pusillus1, Asthenohymen dunbari5
"Doter affinis" = Asthenohymen affinis1 Tillyard 1926 winged insect
MCZ 3095
"Doter pusillus" = Asthenohymen pusillus1 Tillyard 1926 winged insect
MCZ 3097
Asthenohymen dunbari5 Tillyard 1924 winged insect
MCZ 3059,3074-94,3111-3119
 Palaeodictyoptera - Elmoboriidae
Elmoboria piperi n. gen. n. sp.10
Elmoboria piperi n. gen. n. sp.10 Carpenter 1976 winged insect
MCZ 5839
 Palaeodictyoptera - Spilapteridae
Dunbaria fasciipennis2 Tillyard 1924 winged insect
MCZ 3056-8
 Reculida - Liomopteridae
Liomopterum ornatum8 Sellards 1909 winged insect
36 from lower limestone, at least 1 from 1927
 Reculida - Lemmatophoridae
Artinska clara6 Sellards 1909 winged insect
160 total, minus 64 collected in 1932
Artinska ovata6 Sellards 1909 winged insect
76 total, minus 21 collected in 1932
Lemmatophora typa6 Sellards 1909 winged insect
More than 200 total, but years not given
Paraprisca fragilis6 Sellards 1909 winged insect
36 total from 1927 and 1932
Paraprisca grandis n. sp.6 Carpenter 1935 winged insect
MCZ 3584,5
Lecorium elongatum6 Sellards 1909 winged insect
46 total, at least 1 from 1927
Lisca minuta6 Sellards 1909 winged insect
104 total, minus 42 collected in 1932
 Dicondylia - Ditaxineuridae
Ditaxineura anomalostigma2 Tillyard 1926 winged insect
MCZ 3043-6
 Dicondylia - Kennedyidae
Progoneura minuta n. gen. n. sp.2
Progoneura minuta n. gen. n. sp.2 Carpenter 1931 winged insect
MCZ 3042
 Meganisoptera - Meganeuridae
Megatypus schucherti2, "Typus permianus" = Tupus permianus2, Oligotypus sp.2, Oligotypus tillyardi n. gen. n. sp.2
Megatypus schucherti2 Tillyard 1925 griffinfly
MCZ 3048
"Typus permianus" = Tupus permianus2 Sellards 1906 griffinfly
MCZ 3050-5
Oligotypus sp.2 Carpenter 1931 griffinfly
MCZ 3041
Oligotypus tillyardi n. gen. n. sp.2 Carpenter 1931 griffinfly
MCZ 3049
 Eoblattida -
Elmonympha carpenteri n. gen. n. sp.11
Elmonympha carpenteri n. gen. n. sp.11 Aristov 2019 insect
MCZ 3622 (year not stated)