Elmo, YPM 1921 collection (Permian of the United States)

Also known as Insect Hill

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.7° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• The Elmo limestone itself about five feet thick; only the very bottom, more massive layer of this stratum contains insects. Overlies underlying terrestrial (swamp) shale with a sharp and irregular contact.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified, gray, argillaceous limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.
• Soft, argillaceous limestone of very light grey color and almost chalky texture, disposed in layers 0.5 to 4 inches thick.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by C.O. Dunbar, R.J. Tillyard in 1921; reposited in the YPM

Collection methods: quarrying,

Primary reference: C. O. Dunbar and R. J. Tillyard. 1924. Kansas Permian insects: Part 1. The geologic occurrence and the environment of the insects, with a description of a new paleodictyopterid. American Journal of Science, Series 5 7(39):171-209 [C. Labandeira/C. Labandeira/R. Day]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111219: authorized by Matthew Clapham, entered by Matthew Clapham on 21.06.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Dicondylia -
Paraneoptera indet.8 winged insect
YPM 5419
 Miomoptera - Palaeomanteidae
"Delopterum minutum" = Delopterum minutum17, "Delopterum dunbari n. sp." = Delopterum minutum17, "Delopteridae indet." = Palaeomanteidae17, Delopsocus fasciatus n. sp.17, Delopsocus elongatus17, Delopsocus latus17
"Delopterum minutum" = Delopterum minutum17 Sellards 1909 winged insect
YPM 5385, ?5393, ?5394
"Delopterum dunbari n. sp." = Delopterum minutum17 Sellards 1909 winged insect
YPM ?5387-8,?5391,5392,?5397,?5399,?5400,5402
"Delopteridae indet." = Palaeomanteidae17 Handlirsch 1906 winged insect
YPM 5395,5396,5398
Delopsocus fasciatus n. sp.17 Tillyard 1928 winged insect
YPM 5376,5377,5404
Delopsocus elongatus17 Sellards 1909 winged insect
YPM 5386,5389
Delopsocus latus17 Sellards 1909 winged insect
YPM 5378-5384,5401
 Hemiptera - Archescytinidae
Archescytina permiana n. gen. n. sp.14, "Permoscytina muiri n. sp." = Archescytina muiri14, "Permoscytina kansasensis n. gen. n. sp." = Archescytina kansasensis14, Permopsylla americana n. gen. n. sp.14
Archescytina permiana n. gen. n. sp.14 Tillyard 1926 true bug
YPM 5111
"Permoscytina muiri n. sp." = Archescytina muiri14 Tillyard 1926 true bug
YPM 5113
"Permoscytina kansasensis n. gen. n. sp." = Archescytina kansasensis14 Tillyard 1926 true bug
YPM 5112
Permopsylla americana n. gen. n. sp.14 Tillyard 1926 true bug
YPM 5114
 Permopsocida - Psocidiidae
"Psocidium minimum n. sp." = Dichentomum minimum13, "Dichentomum tinctum n. gen. n. sp." = Dichentomum tinctum13, "Permentomum tenuiforme n. gen. n. sp." = Dichentomum tinctum13, "Chaetopsocidium sellardsi n. gen. n. sp." = Dichentomum tinctum13, "Psocidium permianum n. gen. n. sp." = Dichentomum tinctum13, "Pentapsocidium indistinctum n. gen. n. sp." = Dichentomum tinctum13, "Psocidium kansasense n. sp." = Dichentomum tinctum13, "Metapsocidium loxoneurum n. gen. n. sp." = Dichentomum tinctum13, "Psocidium robustum n. sp." = Dichentomum tinctum13, "Psocidium abnormale n. sp." = Dichentomum tinctum13, "Psocidium sp." = Dichentomum13
"Psocidium minimum n. sp." = Dichentomum minimum13 Tillyard 1926 winged insect
YPM 5110
"Dichentomum tinctum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5071
"Permentomum tenuiforme n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5101
"Chaetopsocidium sellardsi n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5098
"Psocidium permianum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5072
"Pentapsocidium indistinctum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5100
"Psocidium kansasense n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5075-5078, ?5079-82, ?5084-89
"Metapsocidium loxoneurum n. gen. n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5099
"Psocidium robustum n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5074, 5102
"Psocidium abnormale n. sp." = Dichentomum tinctum13 Tillyard 1926 winged insect
YPM 5073
"Psocidium sp." = Dichentomum13 Tillyard 1926 winged insect
YPM 5083, 5090-5097, 5103
 Permopsocida - Permopsocidae
"Progonopsocus permianus n. gen. n. sp." = Progonopsocus permianus13, "Progonopsocus pusillus n. sp." = Progonopsocus permianus13, "Permopsocus latipennis n. gen. n. sp." = Permopsocus latipennis13, "Permopsocus congener n. sp." = Permopsocus latipennis13, "Ancylopsocus insolitus n. gen. n. sp." = Permopsocus latipennis13, "Permopsocus enderleini n. sp." = Permopsocus latipennis13
"Progonopsocus permianus n. gen. n. sp." = Progonopsocus permianus13 Tillyard 1926 winged insect
YPM 5104
"Progonopsocus pusillus n. sp." = Progonopsocus permianus13 Tillyard 1926 winged insect
YPM 5105
"Permopsocus latipennis n. gen. n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5106
"Permopsocus congener n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5107
"Ancylopsocus insolitus n. gen. n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5109
"Permopsocus enderleini n. sp." = Permopsocus latipennis13 Tillyard 1926 winged insect
YPM 5108
 Cnemidolestodea - Nugonioneuridae
Nugonioneura problematica n. gen. n. sp.25
Nugonioneura problematica n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15561, 27536
 Cnemidolestodea - Protembiidae
"Protembia permiana n. gen. n. sp." = Protembia permiana24, "Telactinopteryx striatipennis n. gen. n. sp." = Protembia permiana23
"Protembia permiana n. gen. n. sp." = Protembia permiana24 Tillyard 1937 winged insect
YPM 15507
"Telactinopteryx striatipennis n. gen. n. sp." = Protembia permiana23 Tillyard 1937 winged insect
YPM 15558-9,15602
 Caloneurodea -
Paleuthygramma acuta2 Carpenter 1943 winged insect
YPM 15561,15655-7,15571,15572
 Caloneurodea - Caloneuridae
Permobiella perspicua n. gen. n. sp.25
Permobiella perspicua n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15593
 Blattodea - Spiloblattinidae
? Spiloblattinidae indet.26 Handlirsch 1906 cockroach
protnotum YPM 15698
Permoblattina permiana22 Sellards 1908 cockroach
YPM 15677-83,15699,15702,15724-32
Permoblattina curta n. gen.22 Sellards 1908 cockroach
YPM 15663-76,15678,15700,15703-23
Pycnoblattina sp.26 Sellards 1908 cockroach
hindwings YPM 15701,15746-9, pronotum YPM 15697
 Dicondylia - Phyloblattidae
"Pycnoblattina curvata" = Phyloblatta curvata26, "Pycnoblattina compacta" = Phyloblatta compacta26
"Pycnoblattina curvata" = Phyloblatta curvata26 Sellards 1908 winged insect
YPM 15692-3,15740-5
"Pycnoblattina compacta" = Phyloblatta compacta26 Sellards 1908 winged insect
YPM 15684-90, 15694-5, 15735-9
 Protorthoptera -
? Lecopterum delicosum23 Sellards 1909 winged insect
YPM 15574
 Orthoptera - Pruvostitidae
Paroedischia recta4 Carpenter 1966 grasshopper
YPM 15757,15638
 Orthoptera - Proparagryllacrididae
Crinoedischia buttsi n. gen. n. sp.1
Crinoedischia buttsi n. gen. n. sp.1 Béthoux and Beckemeyer 2007 grasshopper
YPM 204902
 Orthoptera - Permelcanidae
Promartynovia venicosta n. gen. n. sp.25
Promartynovia venicosta n. gen. n. sp.25 Tillyard 1937 grasshopper
YPM 15594
 Orthoptera - Permoraphidiidae
Permoraphidia americana n. gen. n. sp.19
Permoraphidia americana n. gen. n. sp.19 Tillyard 1932 grasshopper
YPM 5422
 Paoliida - Blattinopsidae
Blattinopsis kukalovae4 Carpenter 1966 winged insect
YPM 15582,15633
 Paoliida - Anthracoptilidae
"Homocladus ornatus n. sp." = Homocladus grandis4
"Homocladus ornatus n. sp." = Homocladus grandis4 Carpenter 1966 winged insect
YPM 15584
 Grylloblattodea - Chelopteridae
Chelopterum peregrinum3 Carpenter 1950 ice crawler
 Grylloblattodea - Camptoneuritidae
Demopterum gracile3 Carpenter 1950 ice crawler
YPM 15793,15562,15646
 Grylloblattodea - Probnidae
Probnis speciosa23 Sellards 1909 ice crawler
YPM 15508-15557,15560,15565-67,15599-601,15603-5,15575,15751-87
 Dicondylia - Permembiidae
Permembia delicatula n. gen. n. sp.17
Permembia delicatula n. gen. n. sp.17 Tillyard 1928 winged insect
YPM 5403
 Dicondylia - Choristosialidae
Choristosialis enigmatica n. gen. n. sp.19
Choristosialis enigmatica n. gen. n. sp.19 Tillyard 1932 winged insect
YPM 5425
 Protelytroptera - Protelytridae
? Protelytron sp.18 Tillyard 1931 winged insect
YPM 5421
Protelytron dunbari n. sp.18 Tillyard 1931 winged insect
YPM 5405,5417-8
Protelytron permianum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 1019
 Protelytroptera - Megelytridae
Megelytron robustum n. gen. n. sp.18
Megelytron robustum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5406
 Protelytroptera - Permelytridae
"? Parablattelytron sp." = Parablattelytron18, "Acosmelytron delicatum n. sp." = Parablattelytron delicatum18, "Acosmelytron elongatum n. gen. n. sp." = Parablattelytron elongatum18, "Acosmelytron acutum n. sp." = Parablattelytron elongatum18, Parablattelytron rectum n. sp.18, "? Acosmelytron sp." = Parablattelytron18, Parablattelytron subincisum n. gen. n. sp.18, Blattelytron permianum n. gen. n. sp.18, Permelytron schucherti n. gen. n. sp.18
"? Parablattelytron sp." = Parablattelytron18 Tillyard 1931 winged insect
YPM 5412
"Acosmelytron delicatum n. sp." = Parablattelytron delicatum18 Tillyard 1931 winged insect
YPM 5416
"Acosmelytron elongatum n. gen. n. sp." = Parablattelytron elongatum18 Tillyard 1931 winged insect
YPM 5415
"Acosmelytron acutum n. sp." = Parablattelytron elongatum18 Tillyard 1931 winged insect
YPM 5414
Parablattelytron rectum n. sp.18 Tillyard 1931 winged insect
YPM 5413
"? Acosmelytron sp." = Parablattelytron18 Tillyard 1931 winged insect
YPM 5407,5410
Parablattelytron subincisum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5408
Blattelytron permianum n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5409
Permelytron schucherti n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5411
 Protelytroptera -
Protelytropsis grandis n. gen. n. sp.18
Protelytropsis grandis n. gen. n. sp.18 Tillyard 1931 winged insect
YPM 5420
 Mecoptera - Anormochoristidae
Anormochorista oligoclada n. gen. n. sp.12
Anormochorista oligoclada n. gen. n. sp.12 Tillyard 1926 scorpionfly
YPM 5068
 Mecoptera - Permochoristidae
"Protochorista tetraclada n. gen. n. sp." = Protochorista tetraclada12, "Protochorista pentaclada n. sp." = Protochorista tetraclada12, Protopanorpa permiana n. gen. n. sp.12
"Protochorista tetraclada n. gen. n. sp." = Protochorista tetraclada12 Tillyard 1926 scorpionfly
YPM 5055
"Protochorista pentaclada n. sp." = Protochorista tetraclada12 Tillyard 1926 scorpionfly
YPM 5056
Protopanorpa permiana n. gen. n. sp.12 Tillyard 1926 scorpionfly
YPM 5064, 5065
 Mecoptera - Permopanorpidae
"Protopanorpa pusilla n. sp." = Lithopanorpa pusilla12, "Permopanorpa inaequalis n. sp." = Permopanorpa inaequalis12, "Permopanorpa dunbari n. sp." = Permopanorpa inaequalis12, "Permopanorpa gracilis n. sp." = Permopanorpa inaequalis12, "Permopanorpa tenuis n. sp." = Permopanorpa inaequalis12, "Permopanorpa sellardsi n. sp." = Permopanorpa inaequalis12, Permopanorpa formosa n. sp.12, Permopanorpa schucherti n. sp.12
"Protopanorpa pusilla n. sp." = Lithopanorpa pusilla12 Tillyard 1926 scorpionfly
YPM 5066
"Permopanorpa inaequalis n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5058
"Permopanorpa dunbari n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5062
"Permopanorpa gracilis n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5060
"Permopanorpa tenuis n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5059
"Permopanorpa sellardsi n. sp." = Permopanorpa inaequalis12 Tillyard 1926 scorpionfly
YPM 5063
Permopanorpa formosa n. sp.12 Tillyard 1926 scorpionfly
YPM 5057
Permopanorpa schucherti n. sp.12 Tillyard 1926 scorpionfly
YPM 5061
 Mecoptera - Protomeropidae
"Platychorista venosa n. gen. n. sp." = Platychorista venosa12, "Protomerope permiana n. gen. n. sp." = Platychorista venosa12
"Platychorista venosa n. gen. n. sp." = Platychorista venosa12 Tillyard 1926 scorpionfly
YPM 5067
"Protomerope permiana n. gen. n. sp." = Platychorista venosa12 Tillyard 1926 scorpionfly
YPM 5069, 5070
 Dicondylia - Permoberothidae
"Permoberotha villosa n. gen. n. sp." = Permoberotha villosa19, "Permoberotha convergens n. gen. n. sp." = Permoberotha villosa25, "Dictyobiella nervosa n. gen. n. sp." = Permoberotha villosa25
"Permoberotha villosa n. gen. n. sp." = Permoberotha villosa19 Tillyard 1932 winged insect
YPM 5426
"Permoberotha convergens n. gen. n. sp." = Permoberotha villosa25 Tillyard 1932 winged insect
YPM 15595
"Dictyobiella nervosa n. gen. n. sp." = Permoberotha villosa25 Tillyard 1932 winged insect
YPM 15592
 Megasecoptera - Protohymenidae
Permohymen schucherti n. gen. n. sp.9, Protohymen permianus n. gen. n. sp.9
Permohymen schucherti n. gen. n. sp.9 Tillyard 1924 winged insect
YPM 5003, 5043
Protohymen permianus n. gen. n. sp.9 Tillyard 1924 winged insect
YPM 5001/5002 (actually counterpart), Cawthron Institute 1702, YPM 5035-43, 5053
 Diaphanopterodea - Martynoviidae
"Martynoviella protohymenoides n. gen. n. sp." = Martynovia protohymenoides19, Martynovia insignis n. gen. n. sp.19
"Martynoviella protohymenoides n. gen. n. sp." = Martynovia protohymenoides19 Tillyard 1932 winged insect
YPM 5424
Martynovia insignis n. gen. n. sp.19 Tillyard 1932 winged insect
YPM 5423
 Diaphanopterodea - Asthenohymenidae
Asthenohymen sp.11 Tillyard 1924 winged insect
YPM 5052
Asthenohymen affinis n. sp.11 Tillyard 1926 winged insect
YPM 5050, ?5048
Asthenohymen gracilis n. sp.11 Tillyard 1926 winged insect
YPM 5045, ?5047
Asthenohymen pusillus n. sp.11 Tillyard 1926 winged insect
YPM 5051
Asthenohymen stigmatizans n. sp.11 Tillyard 1926 winged insect
YPM 5046
Asthenohymen dunbari n. gen. n. sp.9 Tillyard 1924 winged insect
YPM 5004, Cawthron Institute 1706
Asthenohymen stenobasis n. sp.11 Tillyard 1926 winged insect
YPM 5049
Asthenohymen kansasensis n. sp.11 Tillyard 1926 winged insect
YPM 5044
 Diaphanopterodea - Elmoidae
Elmoa trisecta n. gen. n. sp.25
Elmoa trisecta n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15591
 Palaeodictyoptera - Calvertiellidae
Calvertiella permiana n. gen. n. sp.10
Calvertiella permiana n. gen. n. sp.10 Tillyard 1925 winged insect
YPM 5007
 Palaeodictyoptera - Spilapteridae
Dunbaria fasciipennis n. gen. n. sp.
Dunbaria fasciipennis n. gen. n. sp. Tillyard 1924 winged insect
YPM 1001,1002,1050, 5020-2
 Palaeodictyoptera -
Kansasia pulchra n. gen. n. sp.25
Kansasia pulchra n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15589
 Reculida - Liomopteridae
Liomopterum ornatum3 Sellards 1909 winged insect
Liomopterum elongatum3 Sellards 1909 winged insect
YPM 15612, 15749, 15573
Liomopterum sellardsi3 Carpenter 1950 winged insect
YPM 16180
Semopterum venosum3 Carpenter 1950 winged insect
 Reculida - Sylvaphlebiidae
Phenopterum elongatum3 Sellards 1909 winged insect
YPM 15637,15583,15617,15632
 Reculida - Lemmatophoridae
"Artinska clara" = Artinska clara16, "Artinska simplex n. sp." = Artinska clara7, "Estadia banneria n. sp." = Artinska clara7, "Estadia reducta n. sp." = Artinska clara7, "Estadia aucta n. sp." = Artinska clara7, "Estadia altera n. sp." = Artinska clara7, "Estadia confusa n. sp." = Artinska clara7, "Artinska nova n. sp." = Artinska clara7, "Artinska pentamera n. sp." = Artinska clara7, "Orta quadrifida n. sp." = Artinska clara7, "Artinska inversa n. sp." = Artinska clara7, "Artinska tillyardi n. sp." = Artinska clara7, "Estadia kansasia n. sp." = Artinska clara7, "Estadia maculipennis n. sp." = Artinska clara7, "Artinska ovata" = Artinska ovata16, "Orta reducta n. sp." = Artinska ovata7, Artinska sellardsi n. sp.16, "Lemmatophora typica" = Lemmatophora typa15, "Paraprisca fragilis" = Paraprisca fragilis16, "Lemmatophora longipennis n. sp." = Paraprisca fragilis7, "Paraprisca stigmatica n. sp." = Paraprisca fragilis25, Paraprisca grandis25, "Lecorium elongatum" = Lecorium elongatum16, "Metalecorium confusum n. sp." = Lecorium elongatum7, "Sellardsia lecorioides n. sp." = Lecorium elongatum16, "Metalecorium anomalum n. sp." = Lecorium elongatum7, "Sellardsia separanda n. sp." = Lecorium elongatum7, "Sellardsia abnormis n. sp." = Lecorium elongatum7, "Sellardsia kansasensis n. gen. n. sp." = Lecorium elongatum16, "Lemmatophora minuta" = Lisca minuta15, "Lemmatophora cubitalis n. sp." = Lisca minuta7, "Lemmatophora reducta n. sp." = Lisca minuta15
"Artinska clara" = Artinska clara16 Sellards 1909 winged insect
YPM 1003,5263-5265,5285-5290,5292-5312,5332-5363 (several specimens made types of Handlirsch spp.)
"Artinska simplex n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5286
"Estadia banneria n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5302, 5304
"Estadia reducta n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5296
"Estadia aucta n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5295
"Estadia altera n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5298, 5303
"Estadia confusa n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5306
"Artinska nova n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5290
"Artinska pentamera n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5263
"Orta quadrifida n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5269
"Artinska inversa n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5287
"Artinska tillyardi n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5265
"Estadia kansasia n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5297, 5299
"Estadia maculipennis n. sp." = Artinska clara7 Sellards 1909 winged insect
YPM 5264
"Artinska ovata" = Artinska ovata16 Sellards 1909 winged insect
YPM 5313-5331,5364-5375
"Orta reducta n. sp." = Artinska ovata7 Sellards 1909 winged insect
YPM 5277
Artinska sellardsi n. sp.16 Tillyard 1928 winged insect
YPM 5291
"Lemmatophora typica" = Lemmatophora typa15 Sellards 1909 winged insect
YPM 5115-5192,5194,5195,5199-5205
"Paraprisca fragilis" = Paraprisca fragilis16 Sellards 1909 winged insect
YPM 5195,5250-5258,5262,5284
"Lemmatophora longipennis n. sp." = Paraprisca fragilis7 Sellards 1909 winged insect
YPM 5194 (one of Paraprisca fragilis in Tillyard)
"Paraprisca stigmatica n. sp." = Paraprisca fragilis25 Sellards 1909 winged insect
YPM 15579,15596-7
Paraprisca grandis25 Carpenter 1935 winged insect
YPM 15576
"Lecorium elongatum" = Lecorium elongatum16 Sellards 1909 winged insect
YPM 5267,5270,5272-7,5279,5281 (several specimens made types of Handlirsch spp.)
"Metalecorium confusum n. sp." = Lecorium elongatum7 Sellards 1909 winged insect
YPM 5272
"Sellardsia lecorioides n. sp." = Lecorium elongatum16 Sellards 1909 winged insect
YPM 5271
"Metalecorium anomalum n. sp." = Lecorium elongatum7 Sellards 1909 winged insect
YPM 5267
"Sellardsia separanda n. sp." = Lecorium elongatum7 Sellards 1909 winged insect
YPM 5278
"Sellardsia abnormis n. sp." = Lecorium elongatum7 Sellards 1909 winged insect
YPM 5268
"Sellardsia kansasensis n. gen. n. sp." = Lecorium elongatum16 Sellards 1909 winged insect
YPM 5266,5268-9,5278,5280,5282-3 (several specimens made types of Handlirsch spp.)
"Lemmatophora minuta" = Lisca minuta15 Sellards 1909 winged insect
YPM 5193,5196-5198,5206-5223,5225-5246,5259-5261
"Lemmatophora cubitalis n. sp." = Lisca minuta7 Sellards 1909 winged insect
YPM 5208 (one of Lemmatophora minuta in Tillyard)
"Lemmatophora reducta n. sp." = Lisca minuta15 Sellards 1909 winged insect
YPM 5224
 Ephemerida - Protereismatidae
Protereisma sp.20, Protereisma elongatum20, Protereisma sellardsi n. sp.20, "Protereisma permianum" = Protereisma permianum20, "Protereisma minus" = Protereisma permianum20, "Protereisma gracile" = Protereisma permianum20, Protereisma latum20, Protereisma arcuatum20, Protereisma insigne n. sp.20
Protereisma sp.20 Sellards 1907 mayfly
YPM 5466-5469,5473-5506
Protereisma elongatum20 Sellards 1907 mayfly
YPM 1102,1113,5432,5439,5453-4
Protereisma sellardsi n. sp.20 Tillyard 1932 mayfly
YPM 5434
"Protereisma permianum" = Protereisma permianum20 Sellards 1907 mayfly
YPM 1009,1023,1103,1105,1114,1119,1120,5427-30,5433,5436,5437,5449-50
"Protereisma minus" = Protereisma permianum20 Sellards 1907 mayfly
YPM 5451
"Protereisma gracile" = Protereisma permianum20 Sellards 1907 mayfly
YPM 1104,5431,5435,5438,5440-2,5444,5446-8
Protereisma latum20 Sellards 1907 mayfly
YPM 1109,1115-6,5452,5460-4
Protereisma arcuatum20 Sellards 1907 mayfly
YPM 5455-7
Protereisma insigne n. sp.20 Tillyard 1932 mayfly
YPM 1112
 Ephemerida - Misthodotidae
Misthodotes obtusus20, Misthodotes ovalis n. sp.20, "Eudoter delicatulus n. gen. n. sp." = Misthodotes delicatulus21, Misthodotes biguttatus n. sp.20
Misthodotes obtusus20 Sellards 1907 mayfly
YPM 1100,1117,5443,?5459,?5465,5470,5471
Misthodotes ovalis n. sp.20 Tillyard 1932 mayfly
YPM 1118
"Eudoter delicatulus n. gen. n. sp." = Misthodotes delicatulus21 Tillyard 1936 mayfly
YPM 1014
Misthodotes biguttatus n. sp.20 Tillyard 1932 mayfly
YPM 1122
 Dicondylia - Ditaxineuridae
Ditaxineura anomalostigma n. gen. n. sp.11
Ditaxineura anomalostigma n. gen. n. sp.11 Tillyard 1926 winged insect
YPM 5054
 Dicondylia - Kennedyidae
Kennedya mirabilis n. gen. n. sp.10
Kennedya mirabilis n. gen. n. sp.10 Tillyard 1925 winged insect
YPM 5005-6
 Meganisoptera - Meganeuridae
Meganeuridae indet.10, Megatypus ingentissimus n. sp.10, Megatypus schucherti n. gen. n. sp.10, "? Typus sp." = Tupus10, "Typus permianus" = Tupus permianus10
Meganeuridae indet.10 Handlirsch 1906 griffinfly
YPM 5014-9, 5027-34; wing fragments
Megatypus ingentissimus n. sp.10 Tillyard 1925 griffinfly
YPM 5010, ?5025, ?5026
Megatypus schucherti n. gen. n. sp.10 Tillyard 1925 griffinfly
YPM 1021, 5009
"? Typus sp." = Tupus10 Sellards 1906 griffinfly
YPM 5011, 5023; abdominal fragments
"Typus permianus" = Tupus permianus10 Sellards 1906 griffinfly
YPM 5008, ?5024, ?5012, ?5013
 Dicondylia - Camptotaxineuridae
Camptotaxineura ephialtes n. gen. n. sp.25
Camptotaxineura ephialtes n. gen. n. sp.25 Tillyard 1937 winged insect
YPM 15590
 Archaeognatha - Dasyleptidae
"Lepidodasypus sharovi n. gen. n. sp." = Dasyleptus sharovi6
"Lepidodasypus sharovi n. gen. n. sp." = Dasyleptus sharovi6 Durden 1978 jumping bristletails
YPM 28152
 Eoblattida - Euryptilonidae
Stereopterum maculosum n. sp.4 Carpenter 1966 insect
YPM 15653
Stereopterum rotundum3 Carpenter 1950 insect
YPM 15634
Xiphosura
 Xiphosurida - Paleolimulidae
"Paleolimulus avitus n. gen. n. sp." = Paleolimulus signatus5
"Paleolimulus avitus n. gen. n. sp." = Paleolimulus signatus5 Beecher 1904 horseshoe crab
Bivalvia
 Myalinida - Myalinidae
"Myalina meeki n. sp." = Myalinella meeki
"Myalina meeki n. sp." = Myalinella meeki Dunbar 1924 clam