Elmo, MCZ 1932 collection Upper Limestone (Permian of the United States)

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.6° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• Stratigraphy is essentially the same as other Elmo collections except for new upper layer of non extensive limestone. The floor of the quarry where fossils taken is the "Stump layer", resting upon this floor is the white, blocky limestone from which the other collection of insects were taken. In the new quarry the blocky layer is surmounted by about two inches of soft calcareous shale, which in turn is overlain by another blocky layer of limestone. New layer can not be more than 60 feet in diameter. Above this new layer is the shaly limestone found elsewhere in the field.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified, lenticular limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.
• Much harder than lower elmo limestone, sulphur odor when struck by hammer, heavier as well with specific gravity being 12 % higher than lower bed. The new layer contains spots of limonite & numerous dendrites of combined limonite & manganese.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by F.M. Carpenter in 1932; reposited in the MCZ

Collection methods: quarrying,

Primary reference: F. M. Carpenter. 1933. The Lower Permian Insects of Kansas. Part 6: Delopteridae, Protelytroptera, Plectoptera and a New Collection of Protodonata, Odonata, Megasecoptera, Homoptera and Psocoptera. Proceedings of the American Academy of Arts and Sciences 68(11):411-505 [C. Labandeira/C. Labandeira/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111319: authorized by Matthew Clapham, entered by Jered Karr on 23.06.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

• Includes specimens in later publications where collecting year not specified
Insecta
 Ephemerida - Protereismatidae
Protereisma arcuatum Sellards 1907 mayfly
MCZ 3413ab
Protereisma permianum Sellards 1907 mayfly
Protereisma insigne Tillyard 1932 mayfly
MCZ 3436
Protereisma latum Sellards 1907 mayfly
MCZ 3391ab & MCZ 3419ab
 Meganisoptera - Meganeuridae
Megatypus vetustus n. sp., "Typus permianus" = Tupus permianus, Oligotypus tillyardi, "Typus readi n. sp." = Shenzhousia readi
Megatypus vetustus n. sp. Carpenter 1933 griffinfly
MCZ 3254; wing fragment
"Typus permianus" = Tupus permianus Sellards 1906 griffinfly
wing fragment
Oligotypus tillyardi Carpenter 1931 griffinfly
wing fragment
"Typus readi n. sp." = Shenzhousia readi Carpenter 1933 griffinfly
MCZ 3253; wing fragment
 Dicondylia - Ditaxineuridae
Ditaxineura cellulosa Carpenter 1933 winged insect
MCZ 3222; wing fragment
 Dicondylia - Kennedyidae
Kennedya mirabilis Tillyard 1925 winged insect
MCZ 3220; wing fragment
 Reculida - Sylvaphlebiidae
Phenopterum elongatum4 Sellards 1909 winged insect
Year not specified; about half from upper layer
 Permopsocida - Permopsocidae
Lithopsocidium permianum Carpenter 1932 winged insect
MCZ 3365ab; single wing
Progonopsocus permianus Tillyard 1926 winged insect
MCZ 3361ab
Permopsocus latipennis Tillyard 1926 winged insect
MCZ 3359ab; 3 wings & parts of body
 Permopsocida - Psocidiidae
Dichentomum grande n. sp. Carpenter 1933 winged insect
MCZ 3358ab
Dichentomum tinctum Tillyard 1926 winged insect
Common; 125 specimens collected in 1932
 Hemiptera - Archescytinidae
Permopsylla sp. Tillyard 1926 true bug
either P. americana or P. permiana but to poorly preserved
Permopsylla grandis n. gen. n. sp. Carpenter 1933 true bug
MCZ 3287; tegmen
Permopsylla americana Tillyard 1926 true bug
MCZ 3282; 2 isolated tegmina
Permopsylla anomala n. sp. Carpenter 1933 true bug
MCZ 3288; Tegmen
Lithoscytina cubitalis n. gen. n. sp. Carpenter 1933 true bug
MCZ 3289; Tegmen
 Miomoptera - Palaeomanteidae
Delopterum minutum, "Delopterum elongatum" = Delopsocus elongatus
Delopterum minutum Sellards 1909 winged insect
MCZ 3295ab; complete Individual, MCZ 3304ab
"Delopterum elongatum" = Delopsocus elongatus Sellards 1909 winged insect
MCZ 3305
 Grylloblattodea - Liomopteridae
Tapopterum celsum4 Carpenter 1950 ice crawler
Year and layer not specified
Liomopterum ornatum4 Sellards 1909 ice crawler
Year not specified; 37 specimens from upper limestone
 Grylloblattodea - Lemmatophoridae
Lisca minuta2 Sellards 1909 ice crawler
Artinska ovata2 Sellards 1909 ice crawler
Artinska clara2 Sellards 1909 ice crawler
Lemmatophora typa2 Sellards 1909 ice crawler
 Grylloblattodea - Chelopteridae
Chelopterum peregrinum4 Carpenter 1950 ice crawler
Year not specified; about equally from both layers
 Grylloblattodea - Probnidae
Probnis speciosa3 Sellards 1909 ice crawler
Year and layer not specified; about 250 specimens
 Cnemidolestodea - Protembiidae
Aibolitus minutus n. sp.1 Béthoux and Beckemeyer 2007 winged insect
MCZ 7467ab
Protembia permiana4 Tillyard 1937 winged insect
Year not specified; 13 specimens, about equally from each layer
 Cnemidolestodea - Sylvabestiidae
Elmopterum rotundum n. gen. n. sp.1
Elmopterum rotundum n. gen. n. sp.1 Béthoux and Beckemeyer 2007 winged insect
MCZ 7468/9
 Caloneurodea - Caloneuridae
Plesiogramma medialis n. gen. n. sp.3
Plesiogramma medialis n. gen. n. sp.3 Carpenter 1943 winged insect
Year not specified
 Caloneurodea -
Paleuthygramma acuta3 Carpenter 1943 winged insect
Year not specified; 20 specimens from unspecified layer
 Caloneurodea - Apsidoneuridae
Apsidoneura flexa n. gen. n. sp.3
Apsidoneura flexa n. gen. n. sp.3 Carpenter 1943 winged insect
 Paoliida - Blattinopsidae
Blattinopsis kukalovae n. sp.5 Carpenter 1966 winged insect
Year not specified
 Orthoptera - Permoraphidiidae
Permoraphidia grandis n. sp.3 Carpenter 1943 grasshopper
Year not specified
Permoraphidia americana3 Tillyard 1932 grasshopper
Year not specified
 Protelytroptera - Archelytridae
Archelytron superbum n. gen. n. sp.
Archelytron superbum n. gen. n. sp. Carpenter 1933 winged insect
MCZ3389
 Protelytroptera - Permelytridae
"Acosmelytron elongatum" = Parablattelytron elongatum Tillyard 1931 winged insect
MCZ 3386ab
Parablattelytron rectum Tillyard 1931 winged insect
MCZ 3384, MCZ 3388 & MCZ 3383; isolated Elytra
Parablattelytron subincisum Tillyard 1931 winged insect
MCZ 3379ab & MCZ 3375
 Protelytroptera - Protelytridae
Permelytropsis cubitalis n. gen. n. sp. Carpenter 1933 winged insect
MCZ 3373ab
Protelytron permianum Tillyard 1931 winged insect
MCZ 3368ab
Protelytron angustum n. sp. Carpenter 1933 winged insect
MCZ 3371ab
 Protoneuroptera - Permoberothidae
Permoberotha villosa3 Tillyard 1932 winged insect
Year not specified
 Megasecoptera - Protohymenidae
Protohymen readi n. sp. Carpenter 1933 winged insect
MCZ 3257ab; Complete Individual
Protohymen permianus Tillyard 1924 winged insect
MCZ 3261ab; wing fragment
Protohymen elongatus Carpenter 1930 winged insect
MCZ 3263; wing fragment, MCZ 3262ab; Four wings and Abdomen
 Diaphanopterodea - Martynoviidae
Martynovia insignis3 Tillyard 1932 winged insect
MCZ 4600,4601,4604; year not specified
 Diaphanopterodea - Asthenohymenidae
Asthenohymen dunbari Tillyard 1924 winged insect
3 isolated wings
Asthenohymen affinis Tillyard 1926 winged insect
Asthenohymen pusillus Tillyard 1926 winged insect
 Diaphanopterodea - Elmoidae
Elmoa trisecta3 Tillyard 1937 winged insect
Year not specified
 Eoblattida - Euryptilonidae
Stereopterum rotundum4 Carpenter 1950 insect
Year not specified