Huangbanjigou, Chaomidian Village, CNU 2009 collection (Cretaceous of China)

Where: Liaoning, China (41.6° N, 120.8° E: paleocoordinates 44.1° N, 122.9° E)

• coordinate estimated from map

• outcrop-level geographic resolution

When: Yixian Formation (Jehol Group), Late/Upper Barremian (130.0 - 125.5 Ma)

• The Yixian Formation can be subdivided into five units, in ascending order, Lujiatun unit, Lower Lava, Jianshangou unit, Upper Lava, Dawangzhangzi unit, Jingangshan unit, and Huanghuashan unit. Most of the Jehol fossils from the Yixian Formation are preserved in four beds, that is, the Lujiatun bed (recalculated age is 124.0 ± 1.0/1.3 Ma, He et al., 2006), the Jianshangou bed (125.43 ± 0.50/0.88 Ma, Swisher et al., 1999), the Dawangzhangzi bed, and Jinggangshan bed (~122 Ma, Smith et al., 1995).U-Pb dating suggests that the Huangbanjigou section ranges from 124.9 ± 1.7 Ma to 122.8 ± 1.6 Ma (Yang et al., 2007). Chang et al. (2017) obtained Ar/Ar ages of 125.8 ± 1.0 Ma and 126.0 ± 0.8 Ma from the Lujiatun Unit, the lowest part of the Yixian Formation, below the lacustrine shales.

•He et al. (2004) reported a 40Ar/39Ar age of 120.3 ± 0.7 Ma for a tuff layer in the lower part of the Jiufotang Formation, and Chang et al. (2009) obtained a 40Ar/39Ar age of 122.1 ± 0.3 Ma for a tuff layer from the lowermost part of the Jiufotang Formation. Yu et al. (2021) dated an ash bed just above the Yixian/Jiufotang formation boundary to 122.0 ± 0.9 Ma and three tuff beds from the Xiaotaizi section to 118.9 ± 0.8 Ma, 118.8 ± 0.6 Ma, and 118.6 ± 1.1 Ma. These ages suggest that the Jiufotang Formation in Jianchang Basin was deposited at ca. 122.0–118.9 Ma.

•Olierrok et al. (2019) used a selection of U-Pb and 40Ar-39Ar geochronology to place the Barremian–Aptian boundary (the base of the Chron M0r) between 123.8 and 121.8 Ma. The GTS 2020 uses an age of 121.4 Ma for the base of the Aptian, and multiproxy orbital tuning gives an age of 120.2 Ma for the Barremian-Aptian boundary (Leandro et al., 2022). Therefore, the Yixian Formation should be entirely Late Barremian, with the Jiufotang Formation deposited in the Early Aptian (or latest Barremian at its base).

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified, tuffaceous mudstone

Size classes: macrofossils, mesofossils

Preservation: adpression

Collection methods: Repository: Capital Normal University, Beijing

Primary reference: W. T. Zhang, Y. Z. Yao, D. Ren and Y. Y. Zhao. 2011. First description of fossil Naucoridae (Heteroptera: Nepomorpha) from late Mesozoic of China. Acta Geologica Sinica 85:490-500 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111712: authorized by Matthew Clapham, entered by Matthew Clapham on 28.06.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Orthoptera - Prophalangopsidae
Parahagla sibirica3 Sharov 1968 hump-winged cricket
Allaboilus hani n. sp.3 Gu et al. 2010 hump-winged cricket
 Hemiptera - Naucoridae
Exilcrus cameriferus n. gen. Zhang et al. 2011 creeping water bug
 Hemiptera - Ochteridae
Pristinochterus ovatus n. sp.24 Yao et al. 2011 velvety shore bug
 Hemiptera - Kobdocoridae
Clavaticoris zhengi n. gen. n. sp.23
Clavaticoris zhengi n. gen. n. sp.23 Yao et al. 2012 true bug
CNU-HE-LB2009016
 Hemiptera - Primipentatomidae
Quadrocoris radius n. gen. n. sp.22 Yao et al. 2013 true bug
CNU-HE-LB2009017
Primipentatoma peregrina22 Yao et al. 2013 true bug
CNU-HE-LB2009019-027
 Trichoptera -
Cathayamodus fournieri n. gen. n. sp.2
Cathayamodus fournieri n. gen. n. sp.2 Gao et al. 2016 caddisfly
CNU-TRI-LB-2009001
 Raphidioptera - Mesoraphidiidae
Allocotoraphidia myrioneura7 Ren 1997 snakefly
CNU-RAP-LB-2009003
Caloraphidia glossophylla7 Ren 1997 snakefly
CNU-RAP-LB-2009002
 Neuroptera - Mesochrysopidae
Longicellochrysa yixiana n. gen. n. sp.19
Longicellochrysa yixiana n. gen. n. sp.19 Ren et al. 2010 lacewing
CNU-NEU-LJ2009012 locality given as Chaomidian village
 Neuroptera - Kalligrammatidae
Oregramma illecebrosa n. sp.21 Yang et al. 2014 lacewing
CNU-NEU-LB2009031
 Coleoptera - Dytiscidae
Sinoporus lineatus n. gen. n. sp.18, Liadroporus elegans n. gen. n. sp.18, Mesoderus magnus n. gen. n. sp.18, Mesoderus ventralis n. sp.18, Liadytiscus latus n. sp.18, Liadytiscus cretaceus n. gen. n. sp.18, Liadytiscus longitibialis n. sp.18
Sinoporus lineatus n. gen. n. sp.18 Prokin and Ren 2010 predaceous diving beetle
Liadroporus elegans n. gen. n. sp.18 Prokin and Ren 2010 predaceous diving beetle
Mesoderus magnus n. gen. n. sp.18 Prokin and Ren 2010 predaceous diving beetle
Mesoderus ventralis n. sp.18 Prokin and Ren 2010 predaceous diving beetle
Liadytiscus latus n. sp.18 Prokin and Ren 2010 predaceous diving beetle
Liadytiscus cretaceus n. gen. n. sp.18 Prokin and Ren 2010 predaceous diving beetle
Liadytiscus longitibialis n. sp.18 Prokin and Ren 2010 predaceous diving beetle
 Coleoptera - Parandrexidae
Cretoparacucujinae indet.5 Kirejtshuk 2023 beetle
CNU-COL-LB-2009825
 Coleoptera - Chrysomelidae
Mesolpinus angusticollis n. sp.6 Kirejtshuk et al. 2015 leaf beetle
CNU-COL-LB-2009872
Mesolpinus antennatus n. gen. n. sp.6 Kirejtshuk et al. 2015 leaf beetle
CNU-COL-LB-2009840
 Coleoptera - Lasiosynidae
Lasiosyne integera4 Guo et al. 2022 beetle
CNU-COL-LB-2009488
 Coleoptera - Staphylinidae
Cretoquedius infractus n. sp.20 Solodovnikov and Yue 2013 rove beetle
CNU-COL-LB200951
 Coleoptera - Ochodaeidae
Yixianochodaeus horridus n. gen. n. sp.11
Yixianochodaeus horridus n. gen. n. sp.11 Nikolajev 2015 sand-loving scarab beetle
CNU-COL-LB-2009686, 726, 727, 763
 Coleoptera - Pleocomidae
Pleocoma dolichophylla n. sp.13 Nikolajev and Ren 2012 rain beetle
CNU-COL-LB2009725
 Coleoptera - Lucanidae
Litholamprima longimana n. gen. n. sp.16
Litholamprima longimana n. gen. n. sp.16 Nikolajev and Ren 2015 stag beetle
CNU-COL-LB20099817
 Coleoptera - Scarabaeidae
Yixianscarabaeus tenuistriatus n. sp.10 Nikolajev 2015 scarab beetle
CNU-COL-LB2009753, 755, 778
Yixianscarabaeus sulcatus n. gen. n. sp.10 Nikolajev 2015 scarab beetle
CNU-COL-LB2009757, 768
 Coleoptera - Glaphyridae
Cretohypna yixianensis n. sp.17 Nikolajev and Ren 2015 bumble bee scarab beetle
CNU-COL-LB-2009729
Paraglaphyrus subtilis n. sp.9 Nikolajev 2015 bumble bee scarab beetle
CNU-COL-LB2009614(682)
Paraglaphyrus yixianensis n. gen. n. sp.9 Nikolajev 2015 bumble bee scarab beetle
CNU-COL-LB2009664, 608, 605
Paraglaphyrus ovalis n. sp.9 Nikolajev 2015 bumble bee scarab beetle
CNU-COL-LB2009618
Paraglaphyrus robustus n. sp.9 Nikolajev 2015 bumble bee scarab beetle
CNU-COL-LB2009744
Lithohypna tuberculata n. sp.14 Nikolajev and Ren 2012 bumble bee scarab beetle
CNU-COL-LB2009639(655), 632(681)
Lithohypna yuxiana n. sp.14 Nikolajev and Ren 2012 bumble bee scarab beetle
CNU-COL-LB2009634(625)
Lithohypna lepticephala n. sp.14 Nikolajev and Ren 2012 bumble bee scarab beetle
CNU-COL-LB2009648
Lithohypna longula n. sp.14 Nikolajev and Ren 2012 bumble bee scarab beetle
CNU-COL-LB2009649(666)
Lithohypna laoningensis n. sp.8 Nikolajev 2014 bumble bee scarab beetle
CNU-COL-LB2009603(770), 619, 606
Glaphyrus ancestralis n. sp.12 Nikolajev and Ren 2011 bumble bee scarab beetle
CNU-COL-LB2009604, 662
Mesohypna lopatini n. gen. n. sp.15 Nikolajev and Ren 2013 bumble bee scarab beetle
CNU-COL-LB2009710
Mesohypna probata n. sp.8 Nikolajev 2014 bumble bee scarab beetle
CNU-COL-LB2009607
 Coleoptera - Hydrophilidae
Hydroyixia elongata1 Fikáček et al. 2014 water scavenger beetle
CNU-2009075
Hydroyixia latissima n. sp.1 Fikáček et al. 2014 water scavenger beetle
CNU-COL-LB2009074, 107
Alegorius sp.1 Fikáček et al. 2014 water scavenger beetle
CNU-2009081
Alegorius yixianus n. gen. n. sp.1 Fikáček et al. 2014 water scavenger beetle
CNU-2009079, 078, 080, 077