PBDB Collection

Lengwu, upper Shiziling Member, Tonglu (Permian of China)

Where: Zhejiang, China (29.9° N, 119.5° E: paleocoordinates 8.5° N, 102.7° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Shiziling Member (Lengwu Formation), Capitanian (264.3 - 259.5 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: coastal; lithified marl and lithified, muddy limestone

Size class: macrofossils

Preservation: mold/impression

Primary reference: W. P. Liang. 1990. Lengwu Formation of Permian and its Brachiopod Fauna in Zhejiang Province. Geological Memoirs 2(10):1-522 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 111933: authorized by Matthew Clapham, entered by Matthew Clapham on 30.06.2011

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Rhynchonellata
Athyridida - Neoretziidae	Hustedia cf. indica Hustedia cf. indica Waagen 1883
Athyridida - Athyrididae	"Juxathyris abrupta" = Juxathyris abrupta¹, "Araxathyris zhinanensis n. sp." = Juxathyris abrupta, Tongzithyris "sp. 1", Araxathyris tongluensis n. sp. "Juxathyris abrupta" = Juxathyris abrupta¹ Liang 1990 "Araxathyris zhinanensis n. sp." = Juxathyris abrupta Liang 1990 Tongzithyris "sp. 1" Jin et al. 1974 Araxathyris tongluensis n. sp. Liang 1990
Spiriferinida - Reticulariinidae	Altiplecus zhejiangensis n. sp. Altiplecus zhejiangensis n. sp. Liang 1990
Spiriferinida - Spiriferellinidae	"Punctospirifer aff. alpheus" = Crenispirifer alpheus, "Genuspirifer orientalis" = Spiriferellina orientalis, ""Metriolepis" asiatica n. sp." = Spiriferellina orientalis, "Punctospirifer shenzhouensis n. sp." = Spiriferellina orientalis, Spiriferellina discoformis n. sp. "Punctospirifer aff. alpheus" = Crenispirifer alpheus Huang 1933 "Genuspirifer orientalis" = Spiriferellina orientalis Frech 1911 ""Metriolepis" asiatica n. sp." = Spiriferellina orientalis Frech 1911 "Punctospirifer shenzhouensis n. sp." = Spiriferellina orientalis Frech 1911 Spiriferellina discoformis n. sp. Liang 1990
Spiriferida - Ambocoeliidae	Cruricella "sp. 1" Cruricella "sp. 1" Grant 1976
Spiriferida - Martiniidae	Martinia zhexiensis n. sp., Martinia avicula, Martinia anorbicularis Martinia zhexiensis n. sp. Liang 1990 Martinia avicula Liang 1990 Martinia anorbicularis Liang 1990
Spiriferida - Choristitidae	"Semibrachythyrina pyramidiformis" = Alphaneospirifer anshunensis "Semibrachythyrina pyramidiformis" = Alphaneospirifer anshunensis Liao 1980
Spiriferida - Strophopleuridae	Spiriferinaella lengwuensis n. sp. Spiriferinaella lengwuensis n. sp. Liang 1990
Spiriferida - Elythidae	Zhinania eximiusa n. gen. n. sp., Phricodothyris asiatica, "Bullarina indica" = Permophricodothyris indica, "Anomaloria dorsatusa n. sp." = Permophricodothyris indica, "Astegosia qianjiangensis" = Permophricodothyris indica, "Astegosia stropharostrisa" = Permophricodothyris indica, "Bullarina pseudoasiatica n. sp." = Permophricodothyris indica, "Anomaloria puerilis n. sp." = Permophricodothyris indica, "Phricodothyris rotiformis n. sp." = Permophricodothyris rotiformis, Permophricodothyris elegantula¹ Zhinania eximiusa n. gen. n. sp. Liang 1990 Phricodothyris asiatica Chao 1929 "Bullarina indica" = Permophricodothyris indica Waagen 1883 "Anomaloria dorsatusa n. sp." = Permophricodothyris indica Waagen 1883 "Astegosia qianjiangensis" = Permophricodothyris indica Waagen 1883 "Astegosia stropharostrisa" = Permophricodothyris indica Waagen 1883 "Bullarina pseudoasiatica n. sp." = Permophricodothyris indica Waagen 1883 "Anomaloria puerilis n. sp." = Permophricodothyris indica Waagen 1883 "Phricodothyris rotiformis n. sp." = Permophricodothyris rotiformis Liang 1990 Permophricodothyris elegantula¹ Waagen 1883
Orthida - Schizophoriidae	Orthotichia altilis n. sp., Orthotichia gypidularhynchia n. sp. Orthotichia altilis n. sp. Liang 1990 Orthotichia gypidularhynchia n. sp. Liang 1990
Orthida - Enteletidae	"Peltichia transversa" = Peltichia transversa², "Peltichia planisinosa n. sp." = Peltichia transversa, "Enteletes tschernyschewi" = Enteletes tschernyscheffi "Peltichia transversa" = Peltichia transversa² Huang 1933 "Peltichia planisinosa n. sp." = Peltichia transversa Huang 1933 "Enteletes tschernyschewi" = Enteletes tschernyscheffi Diener 1897
Rhynchonellida - Psilocamaridae	"Zhejiangella sexplicata" = Cyrolexis octoplicata, "Zhejiangella quadriplicata" = Cyrolexis octoplicata, "Zhejiangella septaplicata" = Cyrolexis octoplicata, "Zhejiangella pentaplicata" = Cyrolexis octoplicata, "Zhejiangellina wangi n. gen. n. sp." = Cyrolexis octoplicata "Zhejiangella sexplicata" = Cyrolexis octoplicata Liang 1990 "Zhejiangella quadriplicata" = Cyrolexis octoplicata Liang 1990 "Zhejiangella septaplicata" = Cyrolexis octoplicata Liang 1990 "Zhejiangella pentaplicata" = Cyrolexis octoplicata Liang 1990 "Zhejiangellina wangi n. gen. n. sp." = Cyrolexis octoplicata Liang 1990
Strophomenata
Productida - Richthofeniidae	"Caninella zhinanensis" = Linnaeocaninella zhinanensis "Caninella zhinanensis" = Linnaeocaninella zhinanensis Liang 1990
Productida - Permianellidae	"Paritisteges equilateralis" = Permianella typica, "Obliqunsteges distortus n. gen. n. sp." = Permianella typica, "Paritisteges latesulcata" = Dicystoconcha lapparenti, "Tenerella usualisa n. gen." = Dicystoconcha lapparenti, ""Fabulasteges" pumilis n. sp." = Dicystoconcha lapparenti, "Dipunctella stenosulcata n. sp." = Dicystoconcha lapparenti, "Fabulasteges planata n. gen. n. sp." = Dicystoconcha lapparenti "Paritisteges equilateralis" = Permianella typica He and Zhu 1979 "Obliqunsteges distortus n. gen. n. sp." = Permianella typica He and Zhu 1979 "Paritisteges latesulcata" = Dicystoconcha lapparenti Termier et al. 1974 "Tenerella usualisa n. gen." = Dicystoconcha lapparenti Termier et al. 1974 ""Fabulasteges" pumilis n. sp." = Dicystoconcha lapparenti Termier et al. 1974 "Dipunctella stenosulcata n. sp." = Dicystoconcha lapparenti Termier et al. 1974 "Fabulasteges planata n. gen. n. sp." = Dicystoconcha lapparenti Termier et al. 1974
Productida - Lyttoniidae	"Collemataria tilita" = Oldhamina tilita, "Leptodus nobilis" = Leptodus nobilis, "Leptodus nobilis" = Leptodus nobilis⁴, "Semigublerina flabelata n. gen. n. sp." = Leptodus nobilis, Poikilosakos tongluensis n. sp., Keyserlingina "sp. 1", Juxoldhamina imparilisa n. gen., "Juxoldhamina" spicula n. sp. "Collemataria tilita" = Oldhamina tilita Liang 1990 "Leptodus nobilis" = Leptodus nobilis Waagen 1883 "Leptodus nobilis" = Leptodus nobilis⁴ Waagen 1883 "Semigublerina flabelata n. gen. n. sp." = Leptodus nobilis Waagen 1883 Poikilosakos tongluensis n. sp. Liang 1990 Keyserlingina "sp. 1" Tschernyschew 1902 Juxoldhamina imparilisa n. gen. Liang 1990 "Juxoldhamina" spicula n. sp. Liang 1990
Productida - Aulostegidae	"Neoedriosteges transversa n. gen. n. sp." = Edriosteges poyangensis, "Uncisteges maceus" = Urushtenoidea maceus "Neoedriosteges transversa n. gen. n. sp." = Edriosteges poyangensis Kayser 1883 "Uncisteges maceus" = Urushtenoidea maceus Jin 1963
Productida - Echinoconchidae	"Vediproductus punctatiformis" = Vediproductus punctatiformis, "Vediproductus punctatiformis" = Vediproductus punctatiformis³, "Vediproductus humifusus n. sp." = Vediproductus punctatiformis, "Vediproductus tongluensis n. sp." = Vediproductus punctatiformis "Vediproductus punctatiformis" = Vediproductus punctatiformis Chao 1927 "Vediproductus punctatiformis" = Vediproductus punctatiformis³ Chao 1927 "Vediproductus humifusus n. sp." = Vediproductus punctatiformis Chao 1927 "Vediproductus tongluensis n. sp." = Vediproductus punctatiformis Chao 1927
Productida - Paucispiniferidae	"Asioproductus parvus" = Transennatia gratiosa "Asioproductus parvus" = Transennatia gratiosa Waagen 1884
Productida - Monticuliferidae	"Haydenella" "sp. 1", "Haydenella chianensis" = Haydenella kiangsiensis, "Lamiproductus typica n. gen. n. sp." = Haydenella kiangsiensis, "Lamiproductus usualis n. sp." = Haydenella kiangsiensis, Compressoproductus compressus¹, "Monticulifera cassidula n. sp." = Monticulifera cassidula, "Monticulifera cassidula" = Monticulifera cassidula¹, "Monticulifera shizilingensis n. sp." = Monticulifera cassidula, "Zhejiangoproductus zhejiangensis" = Zhejiangoproductus zhejiangensis¹, "Tongluella basilica n. gen. n. sp." = Zhejiangoproductus zhejiangensis "Haydenella" "sp. 1" Reed 1944 "Haydenella chianensis" = Haydenella kiangsiensis Kayser 1883 "Lamiproductus typica n. gen. n. sp." = Haydenella kiangsiensis Kayser 1883 "Lamiproductus usualis n. sp." = Haydenella kiangsiensis Kayser 1883 Compressoproductus compressus¹ Waagen 1884 "Monticulifera cassidula n. sp." = Monticulifera cassidula Liang 1990 "Monticulifera cassidula" = Monticulifera cassidula¹ Liang 1990 "Monticulifera shizilingensis n. sp." = Monticulifera cassidula Liang 1990 "Zhejiangoproductus zhejiangensis" = Zhejiangoproductus zhejiangensis¹ Liang 1990 "Tongluella basilica n. gen. n. sp." = Zhejiangoproductus zhejiangensis Liang 1990
Productida - Productidae	Tyloplecta yangtzeensis, Tyloplecta yangtzeensis¹, Tyloplecta huadongensis n. sp. Tyloplecta yangtzeensis Chao 1927 Tyloplecta yangtzeensis¹ Chao 1927 Tyloplecta huadongensis n. sp. Liang 1990
Productida - Productellidae	Cathaysia subpusilla, Cathaysia chonetoides, Ogbinia dzhagrensis, Echinauris zhexiensis n. sp., Zhuaconcha hirsutispina n. gen. n. sp., "Costispinifera elongata" = Neoplicatifera elongata, Costispinifera shizilingensis n. sp. Cathaysia subpusilla Licharew 1937 Cathaysia chonetoides Chao 1927 Ogbinia dzhagrensis Sarycheva 1965 Echinauris zhexiensis n. sp. Liang 1990 Zhuaconcha hirsutispina n. gen. n. sp. Liang 1990 "Costispinifera elongata" = Neoplicatifera elongata Huang 1932 Costispinifera shizilingensis n. sp. Liang 1990
Orthotetida - Derbyiidae	Derbyia sp.¹ Derbyia sp.¹ Waagen 1884
Orthotetida - Orthotetidae	Orthotetes abruptus n. sp., "Liberella lamella" = Derbyoides lamella, "Liberella cassidula n. gen. n. sp." = Derbyoides cassidula, "Derbyoides cassidula" = Derbyoides cassidula¹, "Liberella elegans n. sp." = Derbyoides cassidula Orthotetes abruptus n. sp. Liang 1990 "Liberella lamella" = Derbyoides lamella Liang 1990 "Liberella cassidula n. gen. n. sp." = Derbyoides cassidula Liang 1990 "Derbyoides cassidula" = Derbyoides cassidula¹ Liang 1990 "Liberella elegans n. sp." = Derbyoides cassidula Liang 1990
Orthotetida - Schuchertellidae	"Schuchertella" "sp. 1" "Schuchertella" "sp. 1" Girty 1904
Orthotetida - Meekellidae	Meekella "sp. 4", Meekella "sp. 3", "Meekella" "sp. 2", "Meekella" "sp. 1", Meekella zhejiangensis, Meekella heterofolda n. sp. Meekella "sp. 4" Meekella "sp. 3" "Meekella" "sp. 2" "Meekella" "sp. 1" Meekella zhejiangensis Liang 1982 Meekella heterofolda n. sp. Liang 1990
Strophomenida - Rugosochonetidae	"Neochonetes millecostata n. sp." = Tenuichonetes tenuilirata, "Mesolobus plicatiformis" = Tenuichonetes plicatiformis, Tenuichonetes lateala, Mesolobus inflata, Chonetinella lengwuensis n. sp., Alatochonetes alata n. gen. n. sp., Neochonetes gabata n. sp., "Neochonetes leurosinosa n. sp." = Neochonetes (Sommeriella) strophomenoides "Neochonetes millecostata n. sp." = Tenuichonetes tenuilirata Chao 1928 "Mesolobus plicatiformis" = Tenuichonetes plicatiformis Lee 1962 Tenuichonetes lateala Liang 1990 Mesolobus inflata Liang 1990 Chonetinella lengwuensis n. sp. Liang 1990 Alatochonetes alata n. gen. n. sp. Liang 1990 Neochonetes gabata n. sp. Liang 1990 "Neochonetes leurosinosa n. sp." = Neochonetes (Sommeriella) strophomenoides Waagen 1884
Lingulata
Lingulida - Lingulidae	"Lingula" foliaceusa n. sp. "Lingula" foliaceusa n. sp. Liang 1990

¹Clapham 2015; ²Shen and Shi 2007; ³Shen et al. 2016; ⁴Verna and Angiolini 2010