PBDB Collection

Bon Tsagaan (PIN collection 3559, outcrop 87/8) (Cretaceous of Mongolia)

Where: Mongolia (45.5° N, 99.1° E: paleocoordinates 46.1° N, 99.1° E)

• coordinate based on nearby landmark

• local area-level geographic resolution

When: Khurilt Member (Dzun-Bain Formation), Aptian (125.0 - 113.0 Ma)

• Possibly late Neocomian (Barremian)-early Aptian but typically treated as Aptian by paleoentomologists.

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine; lithified siliciclastic sediments

Size class: macrofossils

Preservation: adpression

Primary reference: J. M. Carpenter and A. P. Rasnitsyn. 1990. Mesozoic Vespidae. Psyche 97(1-2):1-20 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 113355: authorized by Matthew Clapham, entered by Matthew Clapham on 28.07.2011, edited by Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Insecta
Blattaria - Caloblattinidae	Caloblattina "sp. 2"²¹, Nuurcala popovi n. gen. n. sp.²¹ Caloblattina "sp. 2"²¹ Handlirsch 1906 cockroach PIN 3559/8017 Nuurcala popovi n. gen. n. sp.²¹ Vršanský 2003 cockroach
Hymenoptera - Sepulcidae	Neoxyelula reducta¹⁴, Neoxyelula oligophlebia n. sp.¹⁴, Thoracotrema oculatum n. sp.¹⁰, Ghilarella mercurialis n. gen. n. sp.¹⁰, Ghilarella alexialis n. sp.⁴ Neoxyelula reducta¹⁴ Rasnitsyn 1993 stem sawfly PIN 3559/4433, 4446, ?4440, ?4441 Neoxyelula oligophlebia n. sp.¹⁴ Rasnitsyn 1993 stem sawfly PIN 3559/4432, 4437, 4445 Thoracotrema oculatum n. sp.¹⁰ Rasnitsyn 1988 stem sawfly PIN 3559/4572 Ghilarella mercurialis n. gen. n. sp.¹⁰ Rasnitsyn 1988 stem sawfly PIN 3559/4461, 644, 647, 649, 665, 4462, 4565 (not 646) Ghilarella alexialis n. sp.⁴ Kopylov and Rasnitsyn 2017 stem sawfly PIN 3559/646 (formerly paratype of G. mercurialis)
Hymenoptera - Cephidae	"Mesocephus ghilarovi n. sp." = Cuspilongus ghilarovi¹⁰ "Mesocephus ghilarovi n. sp." = Cuspilongus ghilarovi¹⁰ Rasnitsyn 1988 stem sawfly PIN 3559/652
Hymenoptera - Ichneumonidae	Rudimentifera mora n. gen. n. sp.³, Dischysma maculata n. gen. n. sp.³, Dischysma similis n. sp.³, Palaeoichneumon ornatus n. sp.³, Palaeoichneumon danu n. sp.³ Rudimentifera mora n. gen. n. sp.³ Kopylov 2009 ichneumon wasp PIN 3559/4486 Dischysma maculata n. gen. n. sp.³ Kopylov 2009 ichneumon wasp PIN 3559/4492 Dischysma similis n. sp.³ Kopylov 2009 ichneumon wasp PIN 3559/4490 Palaeoichneumon ornatus n. sp.³ Kopylov 2009 ichneumon wasp PIN 3559/4488 Palaeoichneumon danu n. sp.³ Kopylov 2009 ichneumon wasp PIN 3559/4489
Hymenoptera - Angarosphecidae	"Bestiola subpetiolata n. sp." = Cretobestiola subpetiolata¹⁶, "Bestiola communis n. sp." = Cretobestiola communis¹⁶ "Bestiola subpetiolata n. sp." = Cretobestiola subpetiolata¹⁶ Pulawski and Rasnitsyn 1999 apoid wasp PIN 3559/4529 "Bestiola communis n. sp." = Cretobestiola communis¹⁶ Pulawski and Rasnitsyn 1999 apoid wasp PIN 3559/4525, 4528, 4530, 4526
Hymenoptera - Stigmaphronidae	Aphrostigmon priapulus n. sp.¹², Aphrostigmon bontsaganicum n. sp.¹² Aphrostigmon priapulus n. sp.¹² Rasnitsyn 1991 wasp PIN 3559/4472 Aphrostigmon bontsaganicum n. sp.¹² Rasnitsyn 1991 wasp PIN 3559/4473
Hymenoptera - Dryinidae	? Deinodryinus aptianus n. sp.⁵ ? Deinodryinus aptianus n. sp.⁵ Olmi et al. 2010 wasp PIN 3559/4586
Hymenoptera - Embolemidae	Baissobius carolianus n. sp.¹⁵ Baissobius carolianus n. sp.¹⁵ Rasnitsyn 1996 wasp PIN 3559/4523
Hymenoptera - Vespidae	Priorvespa quadrata n. gen. n. sp., Priorvespa recidiva n. gen. n. sp., Curiosivespa derivata n. sp. Priorvespa quadrata n. gen. n. sp. Carpenter and Rasnitsyn 1990 wasp PIN 3559/4537 Priorvespa recidiva n. gen. n. sp. Carpenter and Rasnitsyn 1990 wasp PIN 3559/4532 Curiosivespa derivata n. sp. Carpenter and Rasnitsyn 1990 wasp PIN 3559/4527
Hymenoptera - Evaniidae	Cretevania meridionalis n. sp.¹² Cretevania meridionalis n. sp.¹² Rasnitsyn 1991 ensign wasp PIN 3559/4484
Hymenoptera - Praeaulacidae	Westratia gobiensis n. sp.¹¹, Miniwestratia minima n. gen. n. sp.¹¹ Westratia gobiensis n. sp.¹¹ Rasnitsyn 1990 wasp PIN 3559/10168 Miniwestratia minima n. gen. n. sp.¹¹ Rasnitsyn 1990 wasp PIN 3559/4485
Hymenoptera - Baissidae	Baissa mongolica n. sp.¹³ Baissa mongolica n. sp.¹³ Rasnitsyn 1991 wasp PIN 3559/4479(4482), 4480(4481)
Trichoptera - Vitimotauliidae	Multimodus parcus¹⁸ Multimodus parcus¹⁸ Sukacheva 1992 caddisfly PIN 3559/5892
Coleoptera - Haliplidae	Haliplus cretaceus n. sp.⁹ Haliplus cretaceus n. sp.⁹ Prokin and Ponomarenko 2013 crawling water beetle PIN 3559/6371
Coleoptera -	Notocupes elegans⁶, Notocupes mongolicus⁶ Notocupes elegans⁶ Ponomarenko 1994 beetle PIN 3669/6060 Notocupes mongolicus⁶ Ponomarenko 1994 beetle PIN 3559/1656, 6038
Coleoptera - Cupedidae	Anaglyphites mongolicus⁷, "Priacmopsis minimus n. sp." = Anaglyphites minimus⁷ Anaglyphites mongolicus⁷ Ponomarenko 1997 reticulated beetle PIN 3559/6055 "Priacmopsis minimus n. sp." = Anaglyphites minimus⁷ Ponomarenko 1997 reticulated beetle PIN 3559/6052
Coleoptera - Ommatidae	"Tetraphalerus bontsaganensis n. sp." = Allophalerus bontsaganensis⁷, Brochocoleus planus n. sp.⁶, Brochocoleus alatus⁶, Brochocoleus cossyphus⁶, Tetraphalerus notatus n. sp.⁷, "Tetraphalerus longicollis" = Omma longicolle⁷, Omma antennatum n. sp.⁷ "Tetraphalerus bontsaganensis n. sp." = Allophalerus bontsaganensis⁷ Ponomarenko 1997 beetle PIN 3559/6051 Brochocoleus planus n. sp.⁶ Ponomarenko 1994 beetle PIN 3559/6058 Brochocoleus alatus⁶ Ponomarenko 1994 beetle PIN 3559/6057 Brochocoleus cossyphus⁶ Ponomarenko 1994 beetle PIN 3559/6046 Tetraphalerus notatus n. sp.⁷ Ponomarenko 1997 beetle PIN 3559/6053, 6029, 6056 "Tetraphalerus longicollis" = Omma longicolle⁷ Ponomarenko 1997 beetle PIN 3559/6033, 6042 Omma antennatum n. sp.⁷ Ponomarenko 1997 beetle PIN 3559/6048
Coleoptera -	"Hydrophilopsia bontsaganica n. sp." = Laetopsia bontsaganica⁸ "Hydrophilopsia bontsaganica n. sp." = Laetopsia bontsaganica⁸ Prokin 2009 water scavenger beetle PIN 3559/6370
Hemiptera - Perforissidae	Tsaganema oshanini n. gen. n. sp.¹⁷ Tsaganema oshanini n. gen. n. sp.¹⁷ Shcherbakov 2007 planthopper PIN 3559/7455, 7457, 7458, 7451, 7452, 7456, 7454, 7453
Alienoptera - Umenocoleidae	Petropterix mirabilis²⁰ Petropterix mirabilis²⁰ Vršanský 2003 winged insect PIN 3559/5785, 5786, 5787, 5788, 5789, 5791, 5792, 5793, 5794, 5890
Dicondylia -	Blattodea indet.²¹ Blattodea indet.²¹ Latreille 1810 cockroach
Blattodea - Blattulidae	"Ctenoblattina tsaganica n. sp." = Elisama tsaganica¹⁹, Elisama parallela²¹ "Ctenoblattina tsaganica n. sp." = Elisama tsaganica¹⁹ Vršanský 1999 cockroach PIN 3559/8100 (count from Vrsansky 2003) Elisama parallela²¹ Vršanský 2003 cockroach PIN 3559/8018, 8031, 8167, 8038, 8053, 8058
Blattodea - Mesoblattinidae	Praeblattella dichotoma n. sp.²¹, Praeblattella zrnko n. sp.²¹, Praeblattella ponomarenkoi²¹, Tarakanula shcherbakovi n. gen. n. sp.²¹ Praeblattella dichotoma n. sp.²¹ Vršanský 2003 cockroach PIN 3559/8050(8152), 8030, 8041, 8068 Praeblattella zrnko n. sp.²¹ Vršanský 2003 cockroach PIN 3559/8065(8109) Praeblattella ponomarenkoi²¹ Vršanský 2003 cockroach PIN 3559/8149, 8188, 8114, 8150 Tarakanula shcherbakovi n. gen. n. sp.²¹ Vršanský 2003 cockroach PIN 3559/8037(8045), 8085, 8062, 8070
Mantodea -	Cretophotina mongolica²¹ Cretophotina mongolica²¹ Gratshev and Zherikhin 1993 mantis PIN 3559/8195
Diptera - Mycetophilidae	Syntemna zhuzhan¹, Syntemna tele¹, Drepanorzeckia extrunculipennis¹, Drepanorzeckia plana¹, Ipsaneusidalys shato¹ Syntemna zhuzhan¹ Blagoderov 1998 fungus gnat PIN 3559/9094, 9095, 10105 Syntemna tele¹ Blagoderov 1998 fungus gnat PIN 3559/9272 Drepanorzeckia extrunculipennis¹ Blagoderov 1997 fungus gnat PIN 3559/9113(9123) Drepanorzeckia plana¹ Blagoderov 1997 fungus gnat PIN 3559/9070, 9076 Ipsaneusidalys shato¹ Blagoderov 1998 fungus gnat PIN 3559/9196, 9200, 9246, 9247
Arachnida
Araneae - Antrodiaetidae	Cretacattyma raveni n. gen. n. sp.² Cretacattyma raveni n. gen. n. sp.² Eskov and Zonshtein 1990 folding trapdoor spider PIN 3559/5762
Araneae -	Cretomegahexura platnicki n. gen. n. sp.² Cretomegahexura platnicki n. gen. n. sp.² Eskov and Zonshtein 1990 spider PIN 3559/5763, 5760, 5761

¹Blagoderov 1998; ²Eskov and Zonshtein 1990; ³Kopylov 2009; ⁴Kopylov and Rasnitsyn 2017; ⁵Olmi et al. 2010; ⁶Ponomarenko 1994; ⁷Ponomarenko 1997; ⁸Prokin 2009; ⁹Prokin and Ponomarenko 2013; ¹⁰Rasnitsyn 1988; ¹¹Rasnitsyn 1990; ¹²Rasnitsyn 1991; ¹³Rasnitsyn 1991; ¹⁴Rasnitsyn 1993; ¹⁵Rasnitsyn 1996; ¹⁶Rasnitsyn et al. 1999; ¹⁷Shcherbakov 2007; ¹⁸Sukacheva 1992; ¹⁹Vršanský 1999; ²⁰Vršanský 2003; ²¹Vršanský 2003