Elmo, MCZ 1935 collection Upper Limestone (Permian of the United States)

Where: Kansas (38.6° N, 97.2° W: paleocoordinates 3.6° N, 25.8° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Wellington Formation, Artinskian (290.1 - 279.3 Ma)

• Stratigraphy is essentially the same as other Elmo collections except for new upper layer of non extensive limestone. The floor of the quarry where fossils taken is the "Stump layer", resting upon this floor is the white, blocky limestone from which the other collection of insects were taken. In the new quarry the blocky layer is surmounted by about two inches of soft calcareous shale, which in turn is overlain by another blocky layer of limestone. New layer can not be more than 60 feet in diameter. Above this new layer is the shaly limestone found elsewhere in the field. Upper layer thins laterally and pinches out within 20 feet of edge of new quarry.

• group of beds-level stratigraphic resolution

Environment/lithology: pond; lithified, lenticular limestone

• Presence of Myalinella may suggest brackish-marine conditions, but stratigraphic position and presence of conchostracans and freshwater ostracods suggest a lacustrine/pond setting.
• Much harder than lower Elmo limestone, sulphur odor when struck by hammer, heavier as well with specific gravity being 12 % higher than lower bed. The new layer contains spots of limonite & numerous dendrites of combined limonite & manganese.

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by F.M. Carpenter in 1935; reposited in the MCZ

Collection methods: quarrying,

Primary reference: F. M. Carpenter. 1939. The Lower Permian Insects of Kansas. Part 8: Additional Megasecoptera, Protodonata, Odonata, Homoptera, Psocoptera, Protelytroptera, Plectoptera and Protoperlaria. Proceedings of the American Academy of Arts and Sciences 73(3):29-70 [C. Labandeira/C. Labandeira/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 114151: authorized by Matthew Clapham, entered by Matthew Clapham on 11.08.2011, edited by Conrad Labandeira

Creative Commons license: CC BY (attribution)

Taxonomic list

 Protelytroptera - Protelytridae
Protelytron permianum Tillyard 1931 winged insect
Protelytron furcatum n. sp. Carpenter 1939 winged insect
 Protelytroptera - Permelytridae
Permelytron schucherti Tillyard 1931 winged insect
Bed (upper/lower) not given
Parablattelytron subincisum Tillyard 1931 winged insect
Parablattelytron rectum Tillyard 1931 winged insect
 Grylloblattodea - Lemmatophoridae
Lisca minuta Sellards 1909 ice crawler
Paraprisca grandis Carpenter 1935 ice crawler
Lecorium elongatum Sellards 1909 ice crawler
Artinska clara Sellards 1909 ice crawler
Artinska ovata Sellards 1909 ice crawler
Lemmatophora typa Sellards 1909 ice crawler
 Caloneurodea - Caloneuridae
Anomalogramma parva n. gen. n. sp.
Anomalogramma parva n. gen. n. sp. Carpenter 1943 winged insect
 Miomoptera - Palaeomanteidae
Delopterum minutum, "Delopterum kansanum n. sp." = Delopsocus kansanum, "Delopterum elongatum" = Delopsocus elongatus
Delopterum minutum Sellards 1909 winged insect
"Delopterum kansanum n. sp." = Delopsocus kansanum Carpenter 1939 winged insect
"Delopterum elongatum" = Delopsocus elongatus Sellards 1909 winged insect
 Permopsocida - Permopsocidae
Lithopsocidium permianum Carpenter 1932 winged insect
Permopsocus ovatus n. sp. Carpenter 1939 winged insect
Permopsocus latipennis Tillyard 1926 winged insect
 Permopsocida - Psocidiidae
Dichentomum tinctum Tillyard 1926 winged insect
Bed (upper/lower) not given
 Hemiptera - Archescytinidae
"Permoscytina kansasensis" = Archescytina kansasensis Tillyard 1926 true bug
Bed (upper/lower) not given
Archescytina permiana Tillyard 1926 true bug
Permopsylla americana Tillyard 1926 true bug
Permopsylla grandis Carpenter 1933 true bug
Permopsylla minuta Carpenter 1931 true bug
Permopsylla permiana Carpenter 1931 true bug
 Megasecoptera - Protohymenidae
Permohymen schucherti Tillyard 1924 winged insect
Protohymen permianus Tillyard 1924 winged insect
Protohymen readi Carpenter 1933 winged insect
Protohymen elongatus Carpenter 1930 winged insect
 Diaphanopterodea - Asthenohymenidae
Asthenohymen dunbari Tillyard 1924 winged insect
Asthenohymen pusillus Tillyard 1926 winged insect
 Ephemerida - Misthodotidae
Misthodotes ovalis Tillyard 1932 mayfly
Bed (upper/lower) not given
Misthodotes obtusus Sellards 1907 mayfly
At least three from upper layer
 Ephemerida - Protereismatidae
Protereisma insigne Tillyard 1932 mayfly
 Meganisoptera - Meganeuridae
"Typus permianus" = Tupus permianus, Meganeuropsis permiana n. gen. n. sp., "Typus readi" = Shenzhousia readi
"Typus permianus" = Tupus permianus Sellards 1906 griffinfly
Meganeuropsis permiana n. gen. n. sp. Carpenter 1939 griffinfly
Possibly from same individual
"Typus readi" = Shenzhousia readi Carpenter 1933 griffinfly
 Dicondylia - Ditaxineuridae
Ditaxineura anomalostigma Tillyard 1926 winged insect
 Dicondylia - Progoneuridae
Progoneura minuta, "Kennedya reducta n. sp." = Progoneura reducta
Progoneura minuta Carpenter 1931 winged insect
"Kennedya reducta n. sp." = Progoneura reducta Carpenter 1939 winged insect
 Dicondylia - Kennedyidae
Kennedya tillyardi n. sp. Carpenter 1939 winged insect
Kennedya mirabilis Tillyard 1925 winged insect