PBDB Collection

Chekarda, Sylva River (PU collection 1) (Permian of Russian Federation)

Where: Russian Federation (57.0° N, 57.7° E)

• Paleocoordinates: 29.0° N, 45.2° E (Wright 2013)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Koshelevka Formation, Kungurian (279.3 - 272.3 Ma)

• Late Kungurian

• group of beds-level stratigraphic resolution

Environment/lithology: deltaic; lithified siliciclastic sediments

• Uralian foreland basin. Alluvial to marine delta-fan environment, close to the sea, deposited under arid climate conditions.

Size class: mesofossils

Preservation: mold/impression

Reposited in the PIN

• Collection of Perm State University (holotypes stored at PIN according to Vilesov, 1995)

Primary reference: S. Y. Storozhenko and D. S. Aristov. 1999. New genus of the family Liomopteridae (Insecta: Grylloblattida) from Lower Permian of Russia. Far Eastern Entomologist 76:6-8 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 114207: authorized by Matthew Clapham, entered by Matthew Clapham on 12.08.2011, edited by Lindy Edwards and Jered Karr

Creative Commons license: CC0 (CC0)

Taxonomic list

Show authors, comments, and common names

Insecta
Diaphanopterodea - Asthenohymenidae	Asthenohymen uralicum¹⁷ Asthenohymen uralicum¹⁷ Zalessky 1939 winged insect PU 1/259, 262, 267
Megasecoptera - Scytohymenidae	Scytohymen extremus¹⁸ Scytohymen extremus¹⁸ Martynov 1937 winged insect PU 1/143 (now PIN 4987/113)
Megasecoptera - Vorkutiidae	Fragmohymen submissus n. gen. n. sp.⁸ Fragmohymen submissus n. gen. n. sp.⁸ Novokshonov 1995 winged insect PU 1/500
Palaeodictyoptera - Calvertiellidae	Sharovia tshekardica n. sp.² Sharovia tshekardica n. sp.² Ivanov 2000 winged insect PU 1/11
Miomoptera - Palaeomanteidae	Palaeomantis aestiva¹⁴ Palaeomantis aestiva¹⁴ Novokshonov 2000 winged insect PU 1/271
Protomeroptera - Protomeropidae	Marimerobius sp.¹⁰, Marimerobius sukatchevae n. sp.¹⁰ Marimerobius sp.¹⁰ Zalessky 1946 winged insect PU 1/445, 447 Marimerobius sukatchevae n. sp.¹⁰ Novokshonov 1997 winged insect PU 1/163 (or 463?), 1/444
Permotrichoptera - Microptysmatidae	Kamopanorpa uralensis⁴ Kamopanorpa uralensis⁴ Martynov 1940 winged insect PU 1/146, 1, 144, 145, 147, 149, 166, 1848 (1/444 later assigned to Marimerobius sukatchevae)
Mecoptera - Permochoristidae	Agetopanorpa sp.⁷, "Agetochorista tillyardi" = Agetopanorpa tillyardi⁶, Protopanorpa sp.⁷, Pulcheripanorpa aliena n. gen. n. sp.⁶ Agetopanorpa sp.⁷ Carpenter 1930 scorpionfly PU 1/416, 158 "Agetochorista tillyardi" = Agetopanorpa tillyardi⁶ Martynov 1940 scorpionfly PU 1/27 Protopanorpa sp.⁷ Tillyard 1926 scorpionfly PU 1/419 Pulcheripanorpa aliena n. gen. n. sp.⁶ Novokshonov 1993 scorpionfly PU 1/164
Dicondylia - Permithonidae	Uralisyra prolubnikovi n. gen. n. sp.²⁰, Uralisyra angusta n. sp.²⁰, Okolpania captiosa n. sp.²⁰, Tshekardithonopsis oblivius n. sp.²⁰ Uralisyra prolubnikovi n. gen. n. sp.²⁰ Vilesov 1995 winged insect PU 1/167 Uralisyra angusta n. sp.²⁰ Vilesov 1995 winged insect PU 1/435 Okolpania captiosa n. sp.²⁰ Vilesov 1995 winged insect PU 1/446 Tshekardithonopsis oblivius n. sp.²⁰ Vilesov 1995 winged insect PU 1/437
Raphidioptera - Parasialidae	Parasialis rozhkovi n. sp.⁵ Parasialis rozhkovi n. sp.⁵ Novokshonov 1993 snakefly PU 1/58
Dicondylia - Permembiidae	Permindigena lientericus¹² Permindigena lientericus¹² Novokshonov 1998 winged insect PU 1/107
Cnemidolestodea - Protembiidae	Sojanoraphidia martynovae¹⁹, Barmaleus dentatus n. gen. n. sp.⁹, Tshekardembia sharovi n. sp.⁸, Sylvardembia tamaena n. gen. n. sp.⁹, Aibolitus medicinus¹⁶ Sojanoraphidia martynovae¹⁹ Storozhenko and Novokshonov 1995 winged insect PU 1/397 Barmaleus dentatus n. gen. n. sp.⁹ Novokshonov 1997 winged insect PU 1/396 Tshekardembia sharovi n. sp.⁸ Novokshonov 1995 winged insect PGU N 1/246 Sylvardembia tamaena n. gen. n. sp.⁹ Novokshonov 1997 winged insect PU 1/395 Aibolitus medicinus¹⁶ Novokshonov and Storozhenko 1996 winged insect PU 1/91
Paoliida - Anthracoptilidae	Mycteroptila armipotens n. sp.¹¹ Mycteroptila armipotens n. sp.¹¹ Novokshonov 1998 winged insect PU 1/506
Paoliida - Paoliidae	Tshekardelia media n. gen. n. sp.¹ Tshekardelia media n. gen. n. sp.¹ Aristov 2002 winged insect PU 1/38
Grylloblattodea -	Sylvonympha tshekardensis n. gen. n. sp.¹⁵ Sylvonympha tshekardensis n. gen. n. sp.¹⁵ Novokshonov and Pan'kov 1999 ice crawler PU 1/7
Grylloblattodea - Lemmatophoridae	Artinska larisae¹³ Artinska larisae¹³ Novokshonov 1999 ice crawler 60 specimens in three collections (all 45 unspecified are placed here arbitrarily)
Grylloblattodea - Liomopteridae	Uralioma variabilis Uralioma variabilis Storozhenko and Aristov 1999 ice crawler PU 1/334,362,363
Dicondylia - Kennedyidae	Kennedya azari n. sp.³ Kennedya azari n. sp.³ Nel et al. 2012 winged insect PIN (sic = PU) 1/276
Dicondylia - Miracopteridae	Miracopteron mirabile⁵ Miracopteron mirabile⁵ Novokshonov 1993 winged insect PU 1/6

¹Aristov 2002; ²Ivanov 2000; ³Nel et al. 2012; ⁴Novokshonov 1992; ⁵Novokshonov 1993; ⁶Novokshonov 1993; ⁷Novokshonov 1994; ⁸Novokshonov 1995; ⁹Novokshonov 1997; ¹⁰Novokshonov 1997; ¹¹Novokshonov 1998; ¹²Novokshonov 1998; ¹³Novokshonov 1999; ¹⁴Novokshonov 2000; ¹⁵Novokshonov and Pan'kov 1999; ¹⁶Novokshonov and Storozhenko 1996; ¹⁷Novokshonov and Willmann 1999; ¹⁸Pecharová and Prokop 2018; ¹⁹Storozhenko and Novokshonov 1995; ²⁰Vilesov 1995