Klein Lehmhagen pit, Grimmen (LGA collection) (Jurassic of Germany)

Where: Mecklenburg-Vorpommern, Germany (54.1° N, 13.1° E: paleocoordinates 43.7° N, 18.9° E)

• coordinate based on political unit

When: Dactylioceras tenuicostatum zone, Early/Lower Toarcian (183.0 - 182.0 Ma)

• unnamed "bituminous shale bed" equivalent to Posidonian Shale (Posidonienschiefer); Lias epsilon level = lowest Toarcian; also lower part of Harpoceras falcifer Zone

• bed-level stratigraphic resolution

Environment/lithology: offshore; lithified, carbonaceous shale

• bituminous shale ("bituminösen Schiefer")

Size classes: macrofossils, mesofossils

Preservation: adpression, concretion

Collected by Ansorge; reposited in the MfN

Primary reference: W. Zessin. 1987. Variabilität, Merkmalswandel und Phylogenie der Elcanidae im Jungpaläozoikum und Mesozoikum und die Phylogenie der Ensifera. Deutsche Entomologische Zeitschrift 34(1-3):1-76 [M. Clapham/J. Karr/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 114993: authorized by Matthew Clapham, entered by Jered Karr on 26.08.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Diptera - Heterorhyphidae
Heterorhyphus triangularis n. sp.2
Heterorhyphus triangularis n. sp.2 Ansorge 1996 fly
LGA 380
 Diptera - Protorhyphidae
Protorhyphus simplex2, Archirhyphus geinitzi2
Protorhyphus simplex2 Geinitz 1888 fly
LGA 533, 571, 744, 998, 1002, 1007, 1294
Archirhyphus geinitzi2 Handlirsch 1939 fly
LGA 999
 Diptera - Pleciofungivoridae
Pleciofungivora liassica n. sp.2
Pleciofungivora liassica n. sp.2 Ansorge 1996 fly
LGA 1045
 Diptera - Anisopodidae
Mesorhyphus zirzipanus n. sp.2
Mesorhyphus zirzipanus n. sp.2 Ansorge 1996 wood gnat
LGA 579
 Diptera - Antefungivoridae
Antefungivora germanica n. sp.2, Archibio mycetophilinus2
Antefungivora germanica n. sp.2 Ansorge 1996 fly
LGA 394
Archibio mycetophilinus2 Handlirsch 1939 fly
LGA 214, 840, 988
 Diptera - Rhagionidae
Palaeobrachyceron willmanni n. sp.2, Grimmyia baltica n. gen. n. sp.2
Palaeobrachyceron willmanni n. sp.2 Ansorge 1996 snipe fly
LGA 269
Grimmyia baltica n. gen. n. sp.2 Ansorge 1996 snipe fly
LGA 950
 Diptera -
Asilomorpha "sp 1"2, Asilomorpha "sp 2"2
Asilomorpha "sp 1"2 Rohdendorf 1961 fly
LGA 690
Asilomorpha "sp 2"2 Rohdendorf 1961 fly
LGA 429
 Diptera - Chironomidae
Aenne liasina n. gen. n. sp.3, Podonominae indet.2, ? Podonomius tumidus n. sp.2, ? Oryctochlus toarciensis n. sp.2
Aenne liasina n. gen. n. sp.3 Ansorge 1999 non biting midge
LGA 1474, 1486, 1894, 1956
Podonominae indet.2 Thienemann and Edwards 1837 non biting midge
LGA 1003
? Podonomius tumidus n. sp.2 Ansorge 1996 non biting midge
LGA 1034
? Oryctochlus toarciensis n. sp.2 Ansorge 1996 non biting midge
LGA 835
 Diptera - Ptychopteridae
"Eoptychoptera liasina" = Eoptychoptera simplex2
"Eoptychoptera liasina" = Eoptychoptera simplex2 Handlirsch 1906 phantom crane fly
LGA 1368
 Diptera - Trichoceridae
Trichoceridae indet.9, Mailotrichocera zessini n. sp.9, Mailotrichocera mikereichi9, Mailotrichocera variabilis9
Trichoceridae indet.9 Kertész 1902 winter crane fly
LGA 1900
Mailotrichocera zessini n. sp.9 Krzemińska et al. 2009 winter crane fly
LGA 611
Mailotrichocera mikereichi9 Krzemińska et al. 2009 winter crane fly
LGA 79, 2005
Mailotrichocera variabilis9 Krzemińska et al. 2009 winter crane fly
LGA 288, 681, 1100, 1163, 2309
 Diptera - Psychodidae
Tanypsycha connexa n. gen. n. sp.1, Nannotanyderus krzeminskii n. gen. n. sp.1, Nannotanyderus grimmenensis n. sp.6, Liassopsychodina pommerana n. gen. n. sp.1
Tanypsycha connexa n. gen. n. sp.1 Ansorge 1994 moth fly
LGA 1233
Nannotanyderus krzeminskii n. gen. n. sp.1 Ansorge 1994 moth fly
LGA 1145, 544
Nannotanyderus grimmenensis n. sp.6 Ansorge and Krzemiński 2002 moth fly
LGA 2222
Liassopsychodina pommerana n. gen. n. sp.1 Ansorge 1994 moth fly
LGA 457, 752
 Diptera - Eoditomyidae
Eoditomyia primitiva n. gen. n. sp.2
Eoditomyia primitiva n. gen. n. sp.2 Ansorge 1996 fly
LGA 1305, 836
 Diptera - Protopleciidae
Protoplecia liasina2, Protoplecia klafacki n. sp.2
Protoplecia liasina2 Geinitz 1884 fly
LGA 1180
Protoplecia klafacki n. sp.2 Ansorge 1996 fly
LGA 892, 924
 Diptera -
Archipleciomima sp.2, Archipleciomima germanica n. sp.8
Archipleciomima sp.2 Rohdendorf 1962 fly
LGA 1203
Archipleciomima germanica n. sp.8 Krzemiński and Ansorge 2018 fly
LGA 1550, 1803, 2282
 Diptera - Chaoboridae
Rhaetomyia herrigi n. sp.2
Rhaetomyia herrigi n. sp.2 Ansorge 1996 phantom midge
LGA 1189
 Odonata - Asiopteridae
"Sphenophlebia pommerana" = Turanopteron pommerana2, "Oreopteridae indet." = Asiopteridae2
"Sphenophlebia pommerana" = Turanopteron pommerana2 Ansorge 1996 dragonfly
LGA 480, 510, 1197
"Oreopteridae indet." = Asiopteridae2 Pritykina 1968 dragonfly
LGA 250, 712, 1358
 Odonata - Liassogomphidae
Liassogomphidae "indet. 1"2, Liassogomphidae "indet. 2"2
Liassogomphidae "indet. 1"2 Tillyard 1935 dragonfly
LGA 922
Liassogomphidae "indet. 2"2 Tillyard 1935 dragonfly
LGA 349
 Odonata - Heterophlebiidae
Trigonophlebia zessini n. gen. n. sp.2, Heterophlebia buckmani2
Trigonophlebia zessini n. gen. n. sp.2 Ansorge 1996 dragonfly
LGA 739
Heterophlebia buckmani2 Brodie 1845 dragonfly
LGA 46
 Odonata - Liassophlebiidae
Grimmenopteron elegantulum n. gen. n. sp.2
Grimmenopteron elegantulum n. gen. n. sp.2 Ansorge 1996 damsel-dragonflies
LGA 345
 Dicondylia - Protomyrmeleontidae
Obotritagrion tenuiformum2, Protomyrmeleon brunonis2, "Zirzipanagrion quadriordinum n. gen. n. sp." = Protomyrmeleon quadriordinum14
Obotritagrion tenuiformum2 Zessin 1991 winged insect
LGA 876
Protomyrmeleon brunonis2 Geinitz 1888 winged insect
LGA 621, 815, 988, 1000
"Zirzipanagrion quadriordinum n. gen. n. sp." = Protomyrmeleon quadriordinum14 Zessin 1991 winged insect
LGA 162/1
 Dicondylia -
Ephemeroptera indet.2
Ephemeroptera indet.2 Hyatt and Arms 1890 mayfly
LGA 1502
 Neuroptera - Osmylidae
"Mesosmylina falciferum n. sp." = Mesosmylina falcifera2
"Mesosmylina falciferum n. sp." = Mesosmylina falcifera2 Ansorge 1996 lance lacewing
LGA 1295
 Neuroptera - Prohemerobiidae
Actinophlebia sp.2, "Sibithone prodroma" = Prohemerobius prodromus2, Prohemerobius latus2
Actinophlebia sp.2 Handlirsch 1906 lacewing
LGA 1080, 324
"Sibithone prodroma" = Prohemerobius prodromus2 Handlirsch 1906 lacewing
LGA 1001, 607, 452
Prohemerobius latus2 Handlirsch 1920 lacewing
LGA 124
 Neuroptera - Ithonidae
Polyosmylus excelsus n. gen. n. sp.2
Polyosmylus excelsus n. gen. n. sp.2 Ansorge 1996 moth lacewing
LGA 1033, 540
 Dicondylia -
Coleoptera indet.5
Coleoptera indet.5 Linnaeus 1758 beetle
LGA 1230
 Coleoptera - Trachypachidae
Liassodites obsti n. gen. n. sp.7
Liassodites obsti n. gen. n. sp.7 Kirejtshuk and Ansorge 2022 false ground beetle
LGA 2850
 Mecoptera - Orthophlebiidae
Orthophlebia brunsvicensis2, Orthophlebia germanica2, Orthophlebia cf. elongata2
Orthophlebia brunsvicensis2 Bode 1905 scorpionfly
LGA 790
Orthophlebia germanica2 Handlirsch 1906 scorpionfly
LGA 811, 1362
Orthophlebia cf. elongata2 Handlirsch 1939 scorpionfly
LGA 737
 Mecoptera - Cimbrophlebiidae
Cimbrophlebiidae indet.5
Cimbrophlebiidae indet.5 Willmann 1977 hangingfly
LGA 1608
 Mecoptera - Bittacidae
Bittacidae indet.2, Mesobittacus minutus2, Neorthophlebia maculipennis2
Bittacidae indet.2 Handlirsch 1906 hangingfly
LGA 500
Mesobittacus minutus2 Handlirsch 1939 hangingfly
LGA 133, 876, 1269, 1298
Neorthophlebia maculipennis2 Handlirsch 1906 hangingfly
LGA 1369
 Mecoptera - Pseudopolycentropodidae
Pseudopolycentropus sp.2, Pseudopolycentropus triangularis2
Pseudopolycentropus sp.2 Handlirsch 1906 scorpionfly
LGA 769, 993
Pseudopolycentropus triangularis2 Handlirsch 1920 scorpionfly
LGA 636, 925, 20, 738, 795, 1151
 Necrotrichoptera - Necrotauliidae
Mesotrichopteridium intermedium4, Necrotaulius parvulus4
Mesotrichopteridium intermedium4 Handlirsch 1906 winged insect
LGA 348, 533, 675, 789, 830, 874, 784, 1425
Necrotaulius parvulus4 Geinitz 1884 winged insect
 Lepidoptera - Eolepidopterigidae
? Eolepidopterigidae indet.15, ? Eolepidopterigidae indet.15, ? Eolepidopterigidae indet.15
? Eolepidopterigidae indet.15 Rasnitsyn 1983 moth
LGA 2150
? Eolepidopterigidae indet.15 Rasnitsyn 1983 moth
LGA 968
? Eolepidopterigidae indet.15 Rasnitsyn 1983 moth
LGA 1500
 Trichoptera - Philopotamidae
Liadotaulius maior4
Liadotaulius maior4 Handlirsch 1906 finger-net caddisfly
LGA 290, 458, 626, 649, 1995, 2040, 2188, 220, 1198, 1710, 672, 713, 934
 Hymenoptera - Karatavitidae
Grimmaratavites mirabilis n. gen. n. sp.10
Grimmaratavites mirabilis n. gen. n. sp.10 Rasnitsyn et al. 2006 wood wasp
LGA 2510
 Plecoptera - Perlariopseidae
Perlariopseidae indet.2, Dicronemoura furcata n. sp.2
Perlariopseidae indet.2 Sinitshenkova 1985 stonefly
LGA 777
Dicronemoura furcata n. sp.2 Ansorge 1996 stonefly
LGA 1278
 Grylloblattodea - Bajanzhargalanidae
Nele jurassica n. gen. n. sp.2
Nele jurassica n. gen. n. sp.2 Ansorge 1996 ice crawler
LGA 1310, 1111, 211, 977
 Orthoptera - Regiatidae
Protochaeta lanceolata2, Protochaeta masculina2
Protochaeta lanceolata2 Handlirsch 1939 grasshopper
LGA 235
Protochaeta masculina2 Handlirsch 1939 grasshopper
LGA 113, 304, 324, 497, 597, 686, 780, 937, 940, 1114, 1280
 Orthoptera - Locustopsidae
"Locustopsis bernstorffi" = Plesioschwinzia bernstorffi2, "Locustopsis kruegeri" = Plesioschwinzia kruegeri13, Locustopsis " sp 1"2, Locustopsis "sp 2"2, Locustopsis "sp 3"2, Locustopsis gyra2
"Locustopsis bernstorffi" = Plesioschwinzia bernstorffi2 Geinitz 1880 grasshopper
LGA 100, 296
"Locustopsis kruegeri" = Plesioschwinzia kruegeri13 Zessin 1983 grasshopper
LGA 100
Locustopsis " sp 1"2 Handlirsch 1906 grasshopper
LGA 1272
Locustopsis "sp 2"2 Handlirsch 1906 grasshopper
LGA 1144
Locustopsis "sp 3"2 Handlirsch 1906 grasshopper
LGA 1309
Locustopsis gyra2 Zessin 1983 grasshopper
LGA 390
 Orthoptera - Protogryllidae
Protogryllus sp.2, "Protogryllus dobbertinensis" = Protogryllus (Protogryllus) dobbertinensis2
Protogryllus sp.2 Handlirsch 1906 grasshopper
LGA 32, 51, 358, 406, 511, 604, 766, 887, 889, 1040, 1172, 1196, 1238
"Protogryllus dobbertinensis" = Protogryllus (Protogryllus) dobbertinensis2 Geinitz 1880 grasshopper
LGA 94, 244, 324, 395, 813, 818, 865, 932, 1113, 1136, 1267
 Orthoptera - Elcanidae
Parelcana tenuis2, "Elcana minima" = Panorpidium minima, "Elcana magna" = Panorpidium magna, "Elcana media" = Panorpidium media, "Elcana geinitzi" = Panorpidium geinitzi2
Parelcana tenuis2 Handlirsch 1906 grasshopper
LGA 487, 1351
"Elcana minima" = Panorpidium minima Handlirsch 1906 grasshopper
LGA 61/1/2
"Elcana magna" = Panorpidium magna Handlirsch 1906 grasshopper
LGA 34/1/2, 62
"Elcana media" = Panorpidium media Handlirsch 1906 grasshopper
LGA 4/1/2, 58/1/2, 63, 402/1/2
"Elcana geinitzi" = Panorpidium geinitzi2 Heer 1880 grasshopper
LGA 445
 Blattodea - Blattulidae
Blattula dubia12, Blattula langfeldti12
Blattula dubia12 Handlirsch 1939 cockroach
LGA 1725
Blattula langfeldti12 Geinitz 1880 cockroach
LGA 86, 96, 27, 331/1(2), 380, 425, 579, 594, 888, 893, 1077, 1201, 1275, 1372, 1529, 1578, 1739, 1820, 2169, 13, 118, 154, 880, 1328, 1435, 1519, 1842, 1887, 1926, 2006, 2015, 2196, 2356, 888, 1820
 Blattodea - Mesoblattinidae
Mesoblattina protypa12
Mesoblattina protypa12 Geinitz 1880 cockroach
LGA 1320, 1942
 Dicondylia - Aerophasmidae
Durnovaria cf. parallela2
Durnovaria cf. parallela2 Whalley 1985 winged insect
LGA 1086, 1323
 Lophioneurida - Lophioneuridae
Undacypha europaea n. sp.2
Undacypha europaea n. sp.2 Ansorge 1996 winged insect
LGA 233, 223, 292, 378, 490, 645, 746, 871, 883, 967, 983, 1274, 1333, 1342, 1339, 1364, 1365
 Hemiptera - Archiconiopterygidae
Archiconiopteryx liasina2
Archiconiopteryx liasina2 Handlirsch 1906 true bug
LGA 620, 666, 747
 Hemiptera - Protopsyllidiidae
Aphidulum ciliatum n. sp.2, Toarcopsyllidium ernsti n. gen. n. sp.2
Aphidulum ciliatum n. sp.2 Ansorge 1996 true bug
LGA 264, 641, 931
Toarcopsyllidium ernsti n. gen. n. sp.2 Ansorge 1996 true bug
LGA 1185
 Hemiptera - Liadopsyllidae
Liadopsylla obtusa n. sp.2
Liadopsylla obtusa n. sp.2 Ansorge 1996 jumping plant louse
LGA 731
 Hemiptera - Oviparosiphidae
Grimmenaphis magnifica n. gen. n. sp.2
Grimmenaphis magnifica n. gen. n. sp.2 Ansorge 1996 aphid
LGA 1313, 903
 Hemiptera - Hylicellidae
Liassocercopis schnicki n. gen. n. sp.2
Liassocercopis schnicki n. gen. n. sp.2 Ansorge 1996 true bug
LGA 814
 Hemiptera - Archijassidae
Archijassidae indet.2, Archijassus heeri2, Mesoledra pachyneura2, Ardela grimmenensis n. gen. n. sp.2
Archijassidae indet.2 Becker-Migdisova 1962 true bug
LGA 240, 380, 384, 701
Archijassus heeri2 Geinitz 1880 true bug
LGA 505, 640, 757, 1204, 1288, 1340, 1300, 108
Mesoledra pachyneura2 Handlirsch 1939 true bug
LGA 138, 206, 742, 1050, 1086, 1299, 927
Ardela grimmenensis n. gen. n. sp.2 Ansorge 1996 true bug
LGA 1348, 673, 856, 1104, 1216, 505, 350, 1325, 163, 266
 Hemiptera - Procercopidae
Procercopidae indet.2, Procercopina frenzeli n. sp.2
Procercopidae indet.2 Handlirsch 1906 froghopper
LGA 676
Procercopina frenzeli n. sp.2 Ansorge 1996 froghopper
LGA 565, 380
 Hemiptera -
Heteroptera indet.5
Heteroptera indet.5 Latreille 1810 true bug
LGA 936
 Hemiptera - Fulgoridiidae
Fulgoridium "sp 1"2, Fulgoridium "sp 2"2, Fulgoridium sp.2
Fulgoridium "sp 1"2 Handlirsch 1906 planthopper
LGA 820
Fulgoridium "sp 2"2 Handlirsch 1906 planthopper
LGA 123
Fulgoridium sp.2 Handlirsch 1906 planthopper
LGA 569, 749, 1183
 Hemiptera -
Fulgoridulum egens2
Fulgoridulum egens2 Handlirsch 1939 planthopper
LGA 70, 89, 198, 714, 1313, 876, 1206, 628, 893, 1349, 920, 899, 896, 893, 1303, 920, 896, 1313
 Permopsocida - Psocidiidae
Liassopsocus lanceolatus n. gen. n. sp.2
Liassopsocus lanceolatus n. gen. n. sp.2 Ansorge 1996 winged insect
LGA 1246, 997
 Permopsocida - Archipsyllidae
Archipsylla primitiva2
Archipsylla primitiva2 Handlirsch 1906 winged insect
LGA 1229
 Eoblattida - Blattogryllidae
Griphopteron molle2
Griphopteron molle2 Handlirsch 1939 insect
LGA 850
Arachnida
 Araneae -
Seppo koponeni n. gen. n. sp.11
Seppo koponeni n. gen. n. sp.11 Selden and Dunlop 2014 spider
MB.A 2966
1Ansorge 1994; 2Ansorge 1996; 3Ansorge 1999; 4Ansorge 2002; 5Ansorge 2003; 6Ansorge and Krzemiński 2002; 7Kirejtshuk and Ansorge 2022; 8Kopeć et al. 2018; 9Krzemińska et al. 2009; 10Rasnitsyn et al. 2006; 11Selden and Dunlop 2014; 12Vršanský and Ansorge 2007; 13Zessin 1988; 14Zessin 1991; 15Zhang et al. 2018