Oeningen (Carlsruhe collection) (Miocene of Germany)

Where: Baden-Württemberg, Germany (47.7° N, 8.9° E: paleocoordinates 47.9° N, 7.8° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 7 mammal zone, Upper Oehningen beds Member (Upper Freshwater-Molasse Formation), Sarmatian (12.7 - 11.6 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified limestone

• "Rising into massive mountains, as in the well-known Righi and Rossberg, they attain a thickness of more than 6000 feet. While they include proofs of the presenceof the sea, they have preserved with marvellous perfection a large number of the plants which clothed the Alps, and of the insects which flitted through the woodlands. They form part of a great series of deposits which have been termed “Molasse” by the Swiss geologists." Geikie, 1882

•"...deposited in the Northern Alpine Foreland basin..." Uhl et al., 2006

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Primary reference: O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 115305: authorized by Matthew Clapham, entered by Matthew Clapham on 01.09.2011

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera -
Halys bruckmanni2 Heer 1853 true bug
lower quarry
Pseudophana amatoria n. sp.2 Heer 1853 true bug
 Hemiptera - Membracidae
Membracites cristatus n. gen. n. sp.2
Membracites cristatus n. gen. n. sp.2 Heer 1853 treehopper
 Hemiptera - Aphrophoridae
Aphrophora spumarioides2 Heer 1853 spittle bug
 Hemiptera - Cercopidae
Cercopis oeningensis n. sp.2 Heer 1853 froghopper
lower quarry
 Hemiptera - Lygaeidae
Lygaeites obsoletus n. sp.2 Heer 1853 milkweed bug
Pachymerus oblongus n. sp.2 Heer 1853 milkweed bug
 Hemiptera - Scutelleridae
Pachycoris germari2 Heer 1853 metallic shield bug
Pachycoris escheri2 Heer 1853 metallic shield bug
 Hemiptera - Pentatomidae
Eusarcoris pinguis n. sp.2 Heer 1853 stink bug
lower quarry
Eurydema brevicollis n. sp.2 Heer 1853 stink bug
lower quarry
Eurydema impudica2 Heer 1853 stink bug
Pentatoma longiceps n. sp.2 Heer 1853 stink bug
lower quarry
 Hemiptera - Reduviidae
Evagoras impressus n. sp.2 Heer 1853 assassin bug
lower quarry
 Hemiptera - Belostomatidae
Diplonychus rotundatus n. sp.2 Heer 1853 giant water bug
 Orthoptera - Tettigoniidae
Phaneroptera vetusta Heer 1849 katydid
 Hymenoptera - Pompilidae
"Pompilus induratus n. sp." = Pompilites induratus
"Pompilus induratus n. sp." = Pompilites induratus Heer 1849 spider wasp
 Hymenoptera - Formicidae
Formica orbata Heer 1849 ant
Formica macrophthalma Heer 1849 ant
Formica immersa3 Heer 1849 ant
SMNK Inv. 1282
Formica seuberti3 Heer 1849 ant
SMNK Inv. 1193 (Oe 113)
Formica longicollis n. sp. Heer 1849 ant
Formica pulchella Heer 1849 ant
Formica procera Heer 1849 ant
"Formica oeningensis" = Camponotus oeningensis Heer 1849 ant
Formica obesa oeningensis
"Formica lignitum" = Camponotus lignitus Germar 1837 ant
Includes SMNK Inv. 1182, 1230
"Formica pinguis" = Oecophylla obesa Heer 1849 weaver ant
ssp. oeningensis
"Ponera fuliginosa" = Poneropsis fuliginosa Heer 1849 ant
ssp. oeningensis (SMNK Inv. 1235, 1236)
Cephites oeningensis n. sp. Heer 1849 ant
Myrmica macrocephala3 Heer 1849 ant
SMNK Inv. 1215
 Hymenoptera - Megachilidae
Lithurge adamitica5 Heer 1865 mason bee
Inv.1216 (Oe.95) (apparently different from Heer specimens)
Osmia antiqua n. sp. Heer 1849 mason bee
 Hymenoptera - Apidae
Xylocopa senilis n. sp. Heer 1849 large carpenter bee
type and 1 poorly preserved in Heer; Inv.1267, 270 (Pl 5 fig 6 in Pongracz, 1931), 1175 (Oe.89, no. 145), 1175 (Oe.89, no. 146), 1228 (Oe.93) in Zeuner & Manning, 1976
Bombus abavus5 Heer 1867 bumble bee
Inv. 1268 (Oe.90)
Anthophorites tonsa n. sp. Heer 1849 bee
Anthophorites titania n. sp. Heer 1849 bee
Holotype Inv.1273 (Oe.92), paratype now at ETH Zurich Pl. I, 686
 Hymenoptera - Ichneumonidae
Anomalon protogaeum n. sp. Heer 1849 ichneumon wasp
Cryptus antiquus n. sp. Heer 1849 ichneumon wasp
 Coleoptera - Dermestidae
Dermestes pauper n. sp.1 Heer 1847 larder beetle
 Coleoptera - Buprestidae
"Ancylochira gracilis n. sp." = Buprestis gracilis1, "Ancylochira deleta n. sp." = Buprestis deleta1, "Ancylochira heydenii n. sp." = Buprestis heydeni1, "Buprestites exstincta n. sp." = Buprestites exstinctus1, Capnodis antiqua1
"Ancylochira gracilis n. sp." = Buprestis gracilis1 Heer 1847 jewel beetle
Carlsruhe museum 279/285
"Ancylochira deleta n. sp." = Buprestis deleta1 Heer 1847 jewel beetle
"Ancylochira heydenii n. sp." = Buprestis heydeni1 Heer 1847 jewel beetle
"Buprestites exstincta n. sp." = Buprestites exstinctus1 Heer 1847 jewel beetle
Capnodis antiqua1 Heer 1847 jewel beetle
 Coleoptera - Cantharidae
"Telephorus fragilis n. sp." = Cantharis (Cantharis) fragilis1, "Telephorus germari n. sp." = Rhagonycha (Rhagonycha) germari1
"Telephorus fragilis n. sp." = Cantharis (Cantharis) fragilis1 Heer 1847 soldier beetle
"Telephorus germari n. sp." = Rhagonycha (Rhagonycha) germari1 Heer 1847 soldier beetle
 Coleoptera - Elateridae
Cardiophorus braunii n. sp.1 Heer 1847 click beetle
Diacanthus sutor n. sp.1 Heer 1847 click beetle
 Coleoptera - Hydrophilidae
"Hydrophilus braunii n. sp." = Hydrobiomorpha braunii1, "Hydrophilus knorrii n. sp." = Hydrophilus spectabilis1, Hydrophilus vexatorius1
"Hydrophilus braunii n. sp." = Hydrobiomorpha braunii1 Heer 1847 water scavenger beetle
"Hydrophilus knorrii n. sp." = Hydrophilus spectabilis1 Heer 1847 water scavenger beetle
Hydrophilus vexatorius1 Heer 1847 water scavenger beetle
 Coleoptera - Staphylinidae
Scaphidium deletum n. sp.1 Heer 1847 rove beetle
 Coleoptera - Meloidae
Lytta aesculapii1 Heer 1847 blister beetle
 Coleoptera - Tenebrionidae
Helops meissneri n. sp.1 Heer 1847 darkling beetle
 Coleoptera - Melyridae
Malachius vertummni1 Heer 1847 soft-winged flower beetle
 Coleoptera - Trogossitidae
"Trogossita koellikeri n. sp." = Trogosita koellikeri1
"Trogossita koellikeri n. sp." = Trogosita koellikeri1 Heer 1847 bark gnawing beetle
 Coleoptera - Coccinellidae
Coccinella hesione n. sp.1 Heer 1847 ladybug or ladybird
 Coleoptera - Cerambycidae
Mesosa jasonis n. sp.1 Heer 1847 longhorned beetle
 Coleoptera - Chrysomelidae
Bruchus striolatus n. sp.1 Heer 1847 leaf beetle
Gonioctena clymene n. sp.1 Heer 1847 leaf beetle
 Coleoptera - Curculionidae
Brachycerus germanus n. sp.1 Heer 1847 snout beetle
Cleonus pyrrhae n. sp.1 Heer 1847 snout beetle
Cleonus deucalionis n. sp.1 Heer 1847 snout beetle
Cleonus larinoides n. sp.1 Heer 1847 snout beetle
 Coleoptera - Anthribidae
Anthribites moussonii n. sp.1 Heer 1847 fungus weevil
 Coleoptera - Scarabaeidae
Aphodius meyeri n. sp.1 Heer 1847 scarab beetle
Rhizotrogus longimanus n. sp.1 Heer 1847 scarab beetle
 Coleoptera - Carabidae
Harpalus sinis n. sp.1 Heer 1847 ground beetle
Argutor antiquus n. sp.1 Heer 1847 ground beetle
 Diptera - Chironomidae
Chironomus obsoletus n. sp. Heer 1849 non biting midge
 Diptera - Asilidae
Asilus antiquus Heer 1849 robber fly
SMNK Inv. 1195
 Diptera - Bibionidae
Bibio elongatus n. sp., "Protomyia amoena n. sp." = Penthetria amoena, "Protomyia jucunda n. sp." = Plecia jucunda, Bibio obsoletus n. sp., Bibio angustatus n. sp., Bibio fusiformis, Bibio moestus n. sp., "Bibio brevis" = Bibio brevis, "Bibio oblongus" = Bibio brevis4
Bibio elongatus n. sp. Heer 1849 march fly
SMNK Oe. 154 Inv. 1189
"Protomyia amoena n. sp." = Penthetria amoena Heer 1849 march fly
SMNK Oe. 170 Inv. 1247; Oe. 159 in Skartveit & Pika 2014
"Protomyia jucunda n. sp." = Plecia jucunda Heer 1849 love bug
SMNK Oe. 168 Inv. 1279
Bibio obsoletus n. sp. Heer 1849 march fly
SMNK Oe. 161
Bibio angustatus n. sp. Heer 1849 march fly
SMNK Oe. 179 Inv. 1274, Oe. 151-153
Bibio fusiformis Heer 1849 march fly
SMNK Oe. 142-145, 148, 161(x2), 194
Bibio moestus n. sp. Heer 1849 march fly
SMNK Oe. 163, 157
"Bibio brevis" = Bibio brevis Heer 1849 march fly
SMNK Oe. 160 Inv. 1193
"Bibio oblongus" = Bibio brevis4 Heer 1849 march fly
SMNK Oe. 156
 Odonata - Aeshnidae
"Aeschna polydore n. sp." = Gynacantha polydore
"Aeschna polydore n. sp." = Gynacantha polydore Heer 1849 darner dragonfly
SMNK Inv. 1265
 Odonata - Corduliidae
Cordulia sp.3 Leach 1815 emerald
SMNK Inv. 1219 (Libellula doris by Heer)
? Cordulia sp.3 Leach 1815 emerald
SMNH Inv. 1257, 1263 (Libellula thoe by Heer)
 Odonata -
"Aeschna" eudore n. sp. Heer 1849 true dragonfly
larva
"Libellula" thetis Heer 1849 true dragonfly
larvae