Oeningen (Carlsruhe collection) (Miocene of Germany)

Where: Baden-Württemberg, Germany (47.7° N, 8.9° E: paleocoordinates 47.9° N, 7.8° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 7 mammal zone, Upper Oehningen beds Member (Upper Freshwater-Molasse Formation), Sarmatian (12.7 - 11.6 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: crater lake; lithified limestone

• "meromictic maar lake of approximately 1000 m in diameter and perhaps up to 100 m in depth" (Lutz, 1997). "Deposited in the Northern Alpine Foreland basin" (Uhl et al., 2006).

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Primary reference: O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 115305: authorized by Matthew Clapham, entered by Matthew Clapham on 01.09.2011

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names
Insecta
 Diptera - Bibionidae
Bibio moestus n. sp., Bibio angustatus n. sp., "Bibio brevis" = Bibio brevis, "Bibio oblongus" = Bibio brevis4, Bibio fusiformis, Bibio obsoletus n. sp., Bibio elongatus n. sp., "Protomyia amoena n. sp." = Penthetria amoena, "Protomyia jucunda n. sp." = Plecia jucunda
Bibio moestus n. sp. Heer 1849 march fly
SMNK Oe. 163, 157
Bibio angustatus n. sp. Heer 1849 march fly
SMNK Oe. 179 Inv. 1274, Oe. 151-153
"Bibio brevis" = Bibio brevis Heer 1849 march fly
SMNK Oe. 160 Inv. 1193
"Bibio oblongus" = Bibio brevis4 Heer 1849 march fly
SMNK Oe. 156
Bibio fusiformis Heer 1849 march fly
SMNK Oe. 142-145, 148, 161(x2), 194
Bibio obsoletus n. sp. Heer 1849 march fly
SMNK Oe. 161
Bibio elongatus n. sp. Heer 1849 march fly
SMNK Oe. 154 Inv. 1189
"Protomyia amoena n. sp." = Penthetria amoena Heer 1849 march fly
SMNK Oe. 170 Inv. 1247; Oe. 159 in Skartveit & Pika 2014
"Protomyia jucunda n. sp." = Plecia jucunda Heer 1849 love bug
SMNK Oe. 168 Inv. 1279
 Diptera - Asilidae
Asilus antiquus
Asilus antiquus Heer 1849 robber fly
SMNK Inv. 1195
 Diptera - Chironomidae
Chironomus obsoletus n. sp.
Chironomus obsoletus n. sp. Heer 1849 non biting midge
 Odonata - Corduliidae
Cordulia sp.3, ? Cordulia sp.3
Cordulia sp.3 Leach 1815 emerald
SMNK Inv. 1219 (Libellula doris by Heer)
? Cordulia sp.3 Leach 1815 emerald
SMNH Inv. 1257, 1263 (Libellula thoe by Heer)
 Odonata - Aeshnidae
"Aeschna polydore n. sp." = Gynacantha polydore
"Aeschna polydore n. sp." = Gynacantha polydore Heer 1849 darner dragonfly
SMNK Inv. 1265
 Odonata -
"Libellula" thetis, "Aeschna" eudore n. sp.
"Libellula" thetis Heer 1849 true dragonfly
larvae
"Aeschna" eudore n. sp. Heer 1849 true dragonfly
larva
 Coleoptera - Scarabaeidae
Aphodius meyeri n. sp.1, Rhizotrogus longimanus n. sp.1
Aphodius meyeri n. sp.1 Heer 1847 scarab beetle
Rhizotrogus longimanus n. sp.1 Heer 1847 scarab beetle
 Coleoptera - Staphylinidae
Scaphidium deletum n. sp.1
Scaphidium deletum n. sp.1 Heer 1847 rove beetle
 Coleoptera - Hydrophilidae
"Hydrophilus braunii n. sp." = Hydrobiomorpha braunii1, "Hydrophilus knorrii n. sp." = Hydrophilus spectabilis1, Hydrophilus vexatorius1
"Hydrophilus braunii n. sp." = Hydrobiomorpha braunii1 Heer 1847 water scavenger beetle
"Hydrophilus knorrii n. sp." = Hydrophilus spectabilis1 Heer 1847 water scavenger beetle
Hydrophilus vexatorius1 Heer 1847 water scavenger beetle
 Coleoptera -
Malachius vertummni1
Malachius vertummni1 Heer 1847 beetle
 Coleoptera - Dermestidae
Dermestes pauper n. sp.1
Dermestes pauper n. sp.1 Heer 1847 skin beetle
 Coleoptera - Elateridae
Cardiophorus braunii n. sp.1, "Diacanthus sutor n. sp." = Ctenicera sutor1
Cardiophorus braunii n. sp.1 Heer 1847 click beetle
"Diacanthus sutor n. sp." = Ctenicera sutor1 Heer 1847 click beetle
 Coleoptera - Cantharidae
"Telephorus germari n. sp." = Rhagonycha (Rhagonycha) germari1, "Telephorus fragilis n. sp." = Cantharis (Cantharis) fragilis1
"Telephorus germari n. sp." = Rhagonycha (Rhagonycha) germari1 Heer 1847 soldier beetle
"Telephorus fragilis n. sp." = Cantharis (Cantharis) fragilis1 Heer 1847 soldier beetle
 Coleoptera - Buprestidae
Capnodis antiqua1, "Ancylochira deleta n. sp." = Buprestis deleta1, "Ancylochira gracilis n. sp." = Buprestis gracilis1, "Ancylochira heydenii n. sp." = Buprestis heydeni1, "Buprestites exstincta n. sp." = Buprestites exstinctus1
Capnodis antiqua1 Heer 1847 jewel beetle
"Ancylochira deleta n. sp." = Buprestis deleta1 Heer 1847 jewel beetle
"Ancylochira gracilis n. sp." = Buprestis gracilis1 Heer 1847 jewel beetle
Carlsruhe museum 279/285
"Ancylochira heydenii n. sp." = Buprestis heydeni1 Heer 1847 jewel beetle
"Buprestites exstincta n. sp." = Buprestites exstinctus1 Heer 1847 jewel beetle
 Coleoptera - Anthribidae
Anthribites moussonii n. sp.1
Anthribites moussonii n. sp.1 Heer 1847 fungus weevil
 Coleoptera - Curculionidae
Cleonus pyrrhae n. sp.1, Cleonus deucalionis n. sp.1, Cleonus larinoides n. sp.1, Brachycerus germanus n. sp.1
Cleonus pyrrhae n. sp.1 Heer 1847 snout beetle
Cleonus deucalionis n. sp.1 Heer 1847 snout beetle
Cleonus larinoides n. sp.1 Heer 1847 snout beetle
Brachycerus germanus n. sp.1 Heer 1847 snout beetle
 Coleoptera - Cerambycidae
Mesosa jasonis n. sp.1
Mesosa jasonis n. sp.1 Heer 1847 longhorned beetle
 Coleoptera - Chrysomelidae
Bruchus striolatus n. sp.1, Gonioctena clymene n. sp.1
Bruchus striolatus n. sp.1 Heer 1847 leaf beetle
Gonioctena clymene n. sp.1 Heer 1847 leaf beetle
 Coleoptera - Trogossitidae
"Trogossita koellikeri n. sp." = Trogosita koellikeri1
"Trogossita koellikeri n. sp." = Trogosita koellikeri1 Heer 1847 bark gnawing beetle
 Coleoptera - Coccinellidae
Coccinella hesione n. sp.1
Coccinella hesione n. sp.1 Heer 1847 ladybird beetle
 Coleoptera - Meloidae
Lytta aesculapii1
Lytta aesculapii1 Heer 1847 blister beetle
 Coleoptera - Tenebrionidae
Helops meissneri n. sp.1
Helops meissneri n. sp.1 Heer 1847 darkling beetle
 Coleoptera - Carabidae
Argutor antiquus n. sp.1, Harpalus sinis n. sp.1
Argutor antiquus n. sp.1 Heer 1847 ground beetle
Harpalus sinis n. sp.1 Heer 1847 ground beetle
 Hymenoptera - Apidae
Anthophorites tonsa n. sp., Anthophorites titania n. sp., Bombus abavus5, Xylocopa senilis n. sp.
Anthophorites tonsa n. sp. Heer 1849 bee
Anthophorites titania n. sp. Heer 1849 bee
Holotype Inv.1273 (Oe.92), paratype now at ETH Zurich Pl. I, 686
Bombus abavus5 Heer 1867 bumble bee
Inv. 1268 (Oe.90)
Xylocopa senilis n. sp. Heer 1849 large carpenter bee
type and 1 poorly preserved in Heer; Inv.1267, 270 (Pl 5 fig 6 in Pongracz, 1931), 1175 (Oe.89, no. 145), 1175 (Oe.89, no. 146), 1228 (Oe.93) in Zeuner & Manning, 1976
 Hymenoptera - Megachilidae
Lithurge adamitica5, Osmia antiqua n. sp.
Lithurge adamitica5 Heer 1864 mason bee
Inv.1216 (Oe.95) (apparently different from Heer specimens)
Osmia antiqua n. sp. Heer 1849 mason bee
 Hymenoptera - Pompilidae
"Pompilus induratus n. sp." = Pompilites induratus
"Pompilus induratus n. sp." = Pompilites induratus Heer 1849 spider wasp
 Hymenoptera - Formicidae
Formica macrophthalma, Formica pulchella, Formica procera, Formica seuberti3, Formica orbata, Formica longicollis n. sp., Formica immersa3, "Formica pinguis" = Oecophylla obesa, "Formica lignitum" = Camponotus lignitus, "Formica oeningensis" = Camponotus oeningensis, "Ponera fuliginosa" = Poneropsis fuliginosa, Myrmica macrocephala3, Cephites oeningensis n. sp.
Formica macrophthalma Heer 1849 ant
Formica pulchella Heer 1849 ant
Formica procera Heer 1849 ant
Formica seuberti3 Heer 1849 ant
SMNK Inv. 1193 (Oe 113)
Formica orbata Heer 1849 ant
Formica longicollis n. sp. Heer 1849 ant
Formica immersa3 Heer 1849 ant
SMNK Inv. 1282
"Formica pinguis" = Oecophylla obesa Heer 1849 weaver ant
ssp. oeningensis
"Formica lignitum" = Camponotus lignitus Germar 1837 ant
Includes SMNK Inv. 1182, 1230
"Formica oeningensis" = Camponotus oeningensis Heer 1849 ant
Formica obesa oeningensis
"Ponera fuliginosa" = Poneropsis fuliginosa Heer 1849 ant
ssp. oeningensis (SMNK Inv. 1235, 1236)
Myrmica macrocephala3 Heer 1849 ant
SMNK Inv. 1215
Cephites oeningensis n. sp. Heer 1849 ant
 Hymenoptera - Ichneumonidae
Anomalon protogaeum n. sp., Cryptus antiquus n. sp.
Anomalon protogaeum n. sp. Heer 1849 ichneumon wasp
Cryptus antiquus n. sp. Heer 1849 ichneumon wasp
 Orthoptera - Tettigoniidae
Phaneroptera vetusta
Phaneroptera vetusta Heer 1849 katydid
 Hemiptera -
Pseudophana amatoria n. sp.2
Pseudophana amatoria n. sp.2 Heer 1853 true bug
 Hemiptera - Membracidae
Membracites cristatus n. gen. n. sp.2
Membracites cristatus n. gen. n. sp.2 Heer 1853 treehopper
 Hemiptera - Aphrophoridae
Aphrophora spumarioides2
Aphrophora spumarioides2 Heer 1853 froghopper
 Hemiptera - Cercopidae
Cercopis oeningensis n. sp.2
Cercopis oeningensis n. sp.2 Heer 1853 froghopper
lower quarry
 Hemiptera - Reduviidae
Evagoras impressus n. sp.2
Evagoras impressus n. sp.2 Heer 1853 assassin bug
lower quarry
 Hemiptera - Belostomatidae
Diplonychus rotundatus n. sp.2
Diplonychus rotundatus n. sp.2 Heer 1853 giant water bug
 Hemiptera - Lygaeidae
Lygaeites obsoletus n. sp.2
Lygaeites obsoletus n. sp.2 Heer 1853 milkweed bug
 Hemiptera - Rhyparochromidae
Pachymerus oblongus n. sp.2
Pachymerus oblongus n. sp.2 Heer 1853 seed bug
 Hemiptera - Scutelleridae
Pachycoris escheri2, Pachycoris germari2
Pachycoris escheri2 Heer 1853 metallic shield bug
Pachycoris germari2 Heer 1853 metallic shield bug
 Hemiptera - Pentatomidae
Halys bruckmanni2, "Eusarcoris pinguis n. sp." = Eysarcoris2, Eurydema brevicollis n. sp.2, Eurydema impudica2, Pentatoma longiceps n. sp.2
Halys bruckmanni2 Heer 1853 stink bug
lower quarry
"Eusarcoris pinguis n. sp." = Eysarcoris2 stink bug
lower quarry
Eurydema brevicollis n. sp.2 Heer 1853 stink bug
lower quarry
Eurydema impudica2 Heer 1853 stink bug
Pentatoma longiceps n. sp.2 Heer 1853 stink bug
lower quarry
1Heer 1847; 2Heer 1853; 3Pongrácz 1931; 4Skartveit and Pika 2014; 5Zeuner and Manning 1976