Letopala River (PIN collection 117) (Permian of Russian Federation)

Where: Arkangelsk, Russian Federation (65.6° N, 42.6° E: paleocoordinates 33.0° N, 32.6° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Iva-Gora Beds Formation, Roadian (272.3 - 268.8 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lagoonal; lithified siliciclastic sediments

• Probable brackish lagoon with some access to sea

Size classes: macrofossils, mesofossils

Preservation: mold/impression

Collected by Yu. D. Zekkel in 1935; reposited in the PIN

Primary reference: A. V. Martynov. 1938. Permian fossil Insects from the Arkhangelsk district. Part V. The family Euthygrammidae and its relationships (with the description of a new genus and family from Chekarda). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7(3):69-80 [M. Clapham/J. Karr/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 116933: authorized by Matthew Clapham, entered by Jered Karr on 14.09.2011, edited by Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Dicondylia - Ampelipteridae
Tshekardobia lurida9 Novokshonov 2004 winged insect
PIN 117/2416
Tshekardobia maculitincta n. sp.9 Novokshonov 2004 winged insect
PIN 117/2285
 Hemiptera - Coleoscytidae
Coleoscyta rotundata3 Martynov 1935 planthopper
PIN 117/1993
 Hemiptera - Scytinopteridae
Scytinoptera cubitalis n. sp.2 Becker-Migdisova 1948 true bug
PIN 117/1939, 1978, 2090
Scytinoptera obliquoovata2 Martynov 1928 true bug
PIN 117/1979, 1745, 1818
Scytinoptera kokeni2 Handlirsch 1904 true bug
PIN 117/1850
Scytinoptera reducta2 Martynov 1928 true bug
PIN 117/1984, 1957, 2110, 2089, 1715, 1920, 1806, 1786, 1956, 2098, 1994
 Hemiptera - Ingruidae
Ingruo inopinata n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1826
Ingruo insperata n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1737
 Hemiptera - Archescytinidae
Archescytina fusca n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1813(1923)
Archescytina zekkeli n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1725
Archescytina gunderseni3 Becker-Migdisova 1960 true bug
PIN 117/2055, 1975
Archescytina carpenteri n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1766
Permopsylla melanostigmata n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1846
Sojanoscytina minor3 Martynov 1933 true bug
PIN 117/1724(1921)
Sojanoscytina difficilis3 Martynov 1933 true bug
PIN 117/1965
Sojanoscytina grandis3 Martynov 1933 true bug
PIN 117/2005
 Hemiptera - Permopsyllidiidae
Permaphidopsis sojanensis n. gen. n. sp.3, Propatrix psylloides n. gen. n. sp.3
Permaphidopsis sojanensis n. gen. n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1810
Propatrix psylloides n. gen. n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/2011
 Hemiptera - Boreoscytidae
"Coleoscyta ramosa n. sp." = Boreoscyta nefasta3
"Coleoscyta ramosa n. sp." = Boreoscyta nefasta3 Becker-Migdisova 1949 true bug
PIN 117/1964
 Hemiptera - Prosbolopseidae
Anaprosbole ivensis3 Becker-Migdisova 1960 true bug
PIN 117/2096
Sojanopsylla brevipennis n. gen. n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/1741(1742)
Scytoneura elliptica3 Martynov 1935 true bug
Ivaia sp.3 Becker-Migdisova 1960 true bug
PIN 117/1830
Ivaia indistincta3 Becker-Migdisova 1960 true bug
PIN 117/1824
Mundus nodosus n. gen. n. sp.3 Becker-Migdisova 1960 true bug
PIN 117/2021A
Prosbolopsis sp.3 Martynov 1935 true bug
PIN 117/2028
Prosbolopsis ovalis3 Martynov 1935 true bug
PIN 117/2006 ,1825, 1809, 1800
 Hemiptera - Prosbolidae
Permocicada simulatrix1 Martynov 1935 true bug
PIN 117/1990, 1924
Permocicada subtruncata1 Martynov 1935 true bug
PIN 117/1952
Permocicada integra1 Becker-Migdisova 1940 true bug
PIN 117/2082(2116)
Permocicada incompleta n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/1752
Permocicada idelensis1 Zalessky 1929 true bug
PIN 117/1781, 1748(1937), 2094, 1981, 2081, 2018
Permocicada pusilla1 Becker-Migdisova 1940 true bug
PIN 117/1925
Permocicadopsis angustata1 Martynov 1935 true bug
PIN 117/1729, 1738(1740), 1936, 2099, 1807, 1795(1794), 1812; 1728, 1954, 1815, 1950, 1708(2029), 1777 (var. typica); 1986, 1822, 1843(2109), 2402, 1731 (var. brevifurcillata); 1721, 1802, 2073, 2022, 1754 (var. devia); 1775, 1757(1732) (var. remota)
Permocicadopsis letopalensis n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/2084(2119), 1987(1744)
Permocicadopsis aberrans1 Martynov 1935 true bug
PIN 117/1852(1717), 1841, 1988, 1730
Permocicadopsis insignis1 Becker-Migdisova 1940 true bug
PIN 117/1805
Sojanoneura edemskii1 Martynov 1928 true bug
PIN 117/2003, 1848(1808)
Sojanoneura bimaculata1 Martynov 1935 true bug
PIN 117/1948, 2023, 2107, 1831, 2112, 2111, 1759, 2095, 1967, 2103
Sojanoneura dubiosa1 Martynov 1935 true bug
PIN 117/1968
Sojanoneura furcatula1 Martynov 1935 true bug
PIN 117/2105(2083)
Sojanoneura innominata1 Becker-Migdisova 1940 true bug
PIN 117/2024(1829)
Sojanoneura triangularis1 Martynov 1935 true bug
PIN 117/1960(2104), 2080(2086), 1821(1743), 1735
Sojanoneura proxima1 Martynov 1928 true bug
PIN 117/1959
Sojanoneura kazanensis1 Zalessky 1929 true bug
PIN 117/2085(1768), 1734(1983), 1940(1944), 2124, 1844
Sojanoneura zekkeliana n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/2000(1827), 1966
Sojanoneura tincta1 Becker-Migdisova 1940 true bug
PIN 117/2122
Prosbole dilatata1 Martynov 1935 true bug
PIN 117/1682, 2117
Prosbole jucunda1 Martynov 1935 true bug
PIN 117/1838
Prosbole propinqua n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/2126(2001)
Prosbole ivagorae n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/2078(2079)
Prosbole cellulifera n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/1980(1946)
Prosbole similis n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/1774(1969)
Prosbole reducta1 Martynov 1935 true bug
PIN 117/1982
Prosbole affinis n. sp.1 Becker-Migdisova 1940 true bug
PIN 117/1970
 Dicondylia - Permosialidae
Permosialis latiformis6 Martynova 1952 winged insect
PIN 117/2603(2601)
Permosialis brevifurcata n. sp.6 Martynova 1952 winged insect
PIN 117/1756
Permosialis lata6 Martynov 1928 winged insect
PIN 117/1722, 1943, 2578, 2590, 2612, 2690
 Caloneurodea - Caloneuridae
Euthygramma parallellum Martynov 1928 winged insect
PIN 117/451, 447, 390, 380, 402, 317, 432, 510, 385
 Paoliida - Paoliidae
Sojanidelia striata n. sp.12 Storozhenko 1992 winged insect
PIN 117/845(856), 796, 800
 Dicondylia - Permembiidae
Letopaloptera albardiana n. gen. n. sp.7
Letopaloptera albardiana n. gen. n. sp.7 Martynova 1961 winged insect
PIN 117/2258(2664),2370
 Glosselytrodea - Glosselytridae
Glosselytron multivenosum n. gen. n. sp.4
Glosselytron multivenosum n. gen. n. sp.4 Martynov 1938 winged insect
PIN 117/676, 677
 Glosselytrodea - Jurinidae
"Jurina reducta n. sp." = Jurina reducta4, "Anorthoneura insignis n. sp." = Jurina reducta4, "Jurina sojanensis n. sp." = Jurina sojanensis4, "Anorthoneura asymmetrica n. sp." = Jurina sojanensis4, Jurina marginata n. sp.4
"Jurina reducta n. sp." = Jurina reducta4 Martynov 1938 winged insect
PIN 117/680, 667, 670, 673, 675, 678
"Anorthoneura insignis n. sp." = Jurina reducta4 Martynov 1938 winged insect
PIN 117/669 (same number given for both Anorthoneura spp.)
"Jurina sojanensis n. sp." = Jurina sojanensis4 Martynov 1938 winged insect
PIN 117/681, 666, 679; 668 form radialis
"Anorthoneura asymmetrica n. sp." = Jurina sojanensis4 Martynov 1938 winged insect
PIN 117/669
Jurina marginata n. sp.4 Martynov 1938 winged insect
PIN 117/682, 233, 663, 667, 671, 672, 677, 678, 685
 Coleoptera - Permocupedidae
Permocupes grandis n. sp.10 Ponomarenko 1963 beetle
PIN 117/394
Permocupes sojanensis n. sp.10 Ponomarenko 1963 beetle
PIN 117/2627
 Mecoptera - Permochoristidae
"Petromantis variolosa" = Petromantis robusta5, "Petromantis rossica" = Petromantis rossica5, "Petromantis borealis" = Petromantis rossica5, "Permochorista maculipennis" = Petromantis maculipennis5, "Permochorista venosa" = Petromantis maculipennis5, "Petrochorista dubia" = Petromantis dubia5, "Permochorista dubiosa" = Petromantis dubiosa5, Petrochorista elegantula5, Petrochorista lepida n. sp.5, Permochorista nana n. sp.5, "Agetochorista letopalae n. sp." = Agetopanorpa letopalae5, "Agetochorista ornata" = Agetopanorpa ornata5, "Agetochorista splendida" = Agetopanorpa ornata5, "Agetochorista dubia" = Agetopanorpa dubia5, "Agetochorista tillyardiana" = Agetopanorpa tillyardiana5, "Agetochorista fasciata" = Agetopanorpa fasciata5
"Petromantis variolosa" = Petromantis robusta5 Martynov 1932 scorpionfly
PIN 117/1538
"Petromantis rossica" = Petromantis rossica5 Handlirsch 1904 scorpionfly
"Petromantis borealis" = Petromantis rossica5 Handlirsch 1904 scorpionfly
"Permochorista maculipennis" = Petromantis maculipennis5 Martynov 1932 scorpionfly
PIN 117/1239(1235), 1248(2649), 1249, 1256, 1470, 1471, 1476, 1497, 1517, 1533, 1564, 1565, 1571, 1579, 1628(1626); var. major 117/1488
"Permochorista venosa" = Petromantis maculipennis5 Martynov 1932 scorpionfly
PIN 117/1221a, 1235, 1451, 1465, 1466, 1478, 1568
"Petrochorista dubia" = Petromantis dubia5 Martynov 1932 scorpionfly
PIN 117/1206, 1511, 1535, 2692
"Permochorista dubiosa" = Petromantis dubiosa5 Martynov 1932 scorpionfly
PIN 117/1453
Petrochorista elegantula5 Martynov 1931 scorpionfly
PIN 117/1491, 1582
Petrochorista lepida n. sp.5 Martynova 1948 scorpionfly
PIN 117/1631, 1523, 2655
Permochorista nana n. sp.5 Martynova 1948 scorpionfly
PIN 117/1509, 1621
"Agetochorista letopalae n. sp." = Agetopanorpa letopalae5 Martynova 1948 scorpionfly
PIN 117/1638, 2740
"Agetochorista ornata" = Agetopanorpa ornata5 Martynov 1932 scorpionfly
PIN 117/1231, 1234, 1268, 1459(1503), 1472, 1480, 1492, 1500, 1519, 1531, 1542, 1561, 1575, 1578, 1620, 1623, 1624, 2654, 2714, 2715, 2737
"Agetochorista splendida" = Agetopanorpa ornata5 Martynov 1932 scorpionfly
PIN 117/1242, 1259, 1260, 1516(1520), 1552, 1560, 1576, 1580, 2643; var. differens 117/1532; var. furcata 117/1230
"Agetochorista dubia" = Agetopanorpa dubia5 Martynov 1932 scorpionfly
PIN 117/1434, 1518, 2667, 2730
"Agetochorista tillyardiana" = Agetopanorpa tillyardiana5 Martynova 1948 scorpionfly
PIN 117/1223(1247)
"Agetochorista fasciata" = Agetopanorpa fasciata5 Martynov 1932 scorpionfly
PIN 117/1217, 1237, 1244, 1481
 Mecoptera - Permopanorpidae
"Permopanorpa angustata" = Martynopanorpa angustata5, "Permopanorpa zekkeli n. sp." = Martynopanorpa zekkeli5
"Permopanorpa angustata" = Martynopanorpa angustata5 Martynov 1932 scorpionfly
PIN 117/1550
"Permopanorpa zekkeli n. sp." = Martynopanorpa zekkeli5 Martynova 1948 scorpionfly
PIN 117/2757
 Dicondylia - Permithonidae
"Permithonopsis ivensis" = Permithonopsis ivensis6, "Permithonopsis proxima" = Permithonopsis ivensis6, Permithonopsis enormis n. sp.6, Permosisyra latipennis6, "Permithonopsis paurovenosa n. sp." = Permosisyra paurovenosa6, "Sialidopsis sojanensis n. sp." = Permosisyra paurovenosa6, Palaemerobius proavitus6
"Permithonopsis ivensis" = Permithonopsis ivensis6 Martynov 1932 winged insect
PIN 117/2142
"Permithonopsis proxima" = Permithonopsis ivensis6 Martynov 1932 winged insect
PIN 117/1508(2553), 2570
Permithonopsis enormis n. sp.6 Martynova 1952 winged insect
PIN 117/2149, 2519, 2528
Permosisyra latipennis6 Martynov 1932 winged insect
PIN 117/2642
"Permithonopsis paurovenosa n. sp." = Permosisyra paurovenosa6 Martynova 1952 winged insect
PIN 117/2555
"Sialidopsis sojanensis n. sp." = Permosisyra paurovenosa6 Martynova 1952 winged insect
PIN 117/2147 (counterpart of P. paurovenosa)
Palaemerobius proavitus6 Martynov 1928 winged insect
PIN 117/2556
 Dicondylia - Permepallagidae
Permepallage sp.8 Martynov 1938 winged insect
PIN 117/66
 Eoblattida - Megakhosaridae
Parakhosara magna n. sp.13 Storozhenko 1994 insect
PIN 117/839
 Eoblattida - Bardapteridae
Olgaephilus emendatus11 Storozhenko 1990 insect
PIN 117/331, 375