PBDB Collection

Oeningen (Zurich collection) (Miocene of Germany)

Where: Baden-Württemberg, Germany (47.7° N, 8.9° E: paleocoordinates 47.9° N, 7.8° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 7 mammal zone, Upper Oehningen beds Member (Upper Freshwater-Molasse Formation), Sarmatian (12.7 - 11.6 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: crater lake; lithified limestone

• "meromictic maar lake of approximately 1000 m in diameter and perhaps up to 100 m in depth" (Lutz, 1997). "Deposited in the Northern Alpine Foreland basin" (Uhl et al., 2006).

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Collection methods: Collection of Zurich University

Primary reference: O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 117252: authorized by Matthew Clapham, entered by Matthew Clapham on 18.09.2011, edited by Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae -	Clubiona eseri n. sp.¹⁰ Clubiona eseri n. sp.¹⁰ Heer 1864 spider
Araneae - Dictynidae	? Argyroneta longipes n. sp.¹⁰ ? Argyroneta longipes n. sp.¹⁰ Heer 1864 spider
Araneae - Gnaphosidae	"Macaria tenella n. sp." = Micaria tenella¹⁰ "Macaria tenella n. sp." = Micaria tenella¹⁰ Heer 1864 spider
Araneae - Thomisidae	Thomisus sulzeri n. sp.¹⁰, Thomisus lividus n. sp.¹⁰, "Thomisus oeningensis n. sp." = Xysticus oeningensis¹⁰ Thomisus sulzeri n. sp.¹⁰ Heer 1864 crab spider Thomisus lividus n. sp.¹⁰ Heer 1864 crab spider "Thomisus oeningensis n. sp." = Xysticus oeningensis¹⁰ Heer 1864 crab spider
Araneae - Araneidae	"Epeira molassica n. sp." = Araneus molassicus¹⁰ "Epeira molassica n. sp." = Araneus molassicus¹⁰ Heer 1864 orb-weaving spider
Araneae - Theridiidae	Theridion maculipes n. sp.¹⁰, Theridion annulipes n. sp.¹⁰, Theridion globulus n. sp.¹⁰ Theridion maculipes n. sp.¹⁰ Heer 1864 cobweb spider Theridion annulipes n. sp.¹⁰ Heer 1864 cobweb spider Theridion globulus n. sp.¹⁰ Heer 1864 cobweb spider
Araneae -	Schellenbergia rotundata n. gen. n. sp.¹⁰ Schellenbergia rotundata n. gen. n. sp.¹⁰ Heer 1864 spider
Malacostraca
Isopoda - Armadillidiidae	"Armadillo molassicus n. sp." = Armadillidium molassicum¹⁰ "Armadillo molassicus n. sp." = Armadillidium molassicum¹⁰ Heer 1864 isopod
Insecta
Ephemeroptera -	Ephemera oeningensis n. sp.¹⁰ Ephemera oeningensis n. sp.¹⁰ Heer 1864 mayfly
Odonata - Gomphidae	Lindenia heeri n. sp.¹ Lindenia heeri n. sp.¹ Boderau et al. 2024 clubtail dragonfly ETH S.N. 228 (labeled "Aeschna tyche")
Odonata -	"Libellula" thetis, "Agrion (Sterope) parthenope n. sp." = Steropoides parthenope "Libellula" thetis Heer 1849 true dragonfly larva "Agrion (Sterope) parthenope n. sp." = Steropoides parthenope Heer 1849 damselfly
Odonata - Sieblosiidae	"Agrion iris n. sp." = Stenolestes iris¹⁴ "Agrion iris n. sp." = Stenolestes iris¹⁴ Heer 1879 damselfly
Orthoptera - Tetrigidae	"Tetrix gracilis n. sp." = Tettigidea gracilis¹⁰ "Tetrix gracilis n. sp." = Tettigidea gracilis¹⁰ Heer 1864 pygmy grasshopper
Orthoptera - Acrididae	Oedipoda germari n. sp.¹⁰ Oedipoda germari n. sp.¹⁰ Heer 1864 short-horned grasshopper
Orthoptera - Pyrgomorphidae	"Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri¹⁰ "Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri¹⁰ Heer 1864 gaudy grasshopper
Orthoptera - Gryllidae	Gryllus troglodytes n. sp.¹⁰ Gryllus troglodytes n. sp.¹⁰ Heer 1864 true crickets
Orthoptera - Gryllotalpidae	Gryllotalpa stricta n. sp.¹⁰ Gryllotalpa stricta n. sp.¹⁰ Heer 1864 mole cricket
Orthoptera - Tettigoniidae	"Decticus speciosus n. sp." = Platycleis speciosa¹⁰ "Decticus speciosus n. sp." = Platycleis speciosa¹⁰ Heer 1864 katydid
Dermaptera - Forficulidae	Forficula recta n. sp.¹⁰, Forficula minuta n. sp.¹⁰ Forficula recta n. sp.¹⁰ Heer 1864 common earwig Forficula minuta n. sp.¹⁰ Heer 1864 common earwig
Blattodea - Rhinotermitidae	"Termes hartungi n. sp." = Reticulitermes hartungi¹⁰ "Termes hartungi n. sp." = Reticulitermes hartungi¹⁰ Heer 1864 termite
Blattodea - Termitidae	Termes buchii n. sp.¹⁰ Termes buchii n. sp.¹⁰ Heer 1864 termite
Blattodea - Blattidae	"Blattidium coloratum n. sp." = Blatta colorata¹¹ "Blattidium coloratum n. sp." = Blatta colorata¹¹ Heer 1864 cockroach from Kesselstein
Hemiptera - Cercopidae	Cercopis hageni n. sp.¹⁰, Cercopis germari n. sp.¹⁰, Cercopis herrichi n. sp.¹⁰ Cercopis hageni n. sp.¹⁰ Heer 1864 froghopper Cercopis germari n. sp.¹⁰ Heer 1864 froghopper Cercopis herrichi n. sp.¹⁰ Heer 1864 froghopper
Hemiptera - Corixidae	"Corisa fasciolata" = Corixa fasciolata⁷ "Corisa fasciolata" = Corixa fasciolata⁷ Heer 1853 water boatman from the Libellen-schicht
Hemiptera - Belostomatidae	"Belostoma speciosum n. sp." = Belostomates speciosum¹⁰ "Belostoma speciosum n. sp." = Belostomates speciosum¹⁰ Heer 1864 giant water bug
Hemiptera - Lygaeidae	Cephalocoris pilosus n. gen. n. sp.⁷ Cephalocoris pilosus n. gen. n. sp.⁷ Heer 1853 milkweed bug lower quarry
Hemiptera - Rhyparochromidae	Pachymerus obsoletus n. sp.⁷ Pachymerus obsoletus n. sp.⁷ Heer 1853 seed bug lower quarry
Hemiptera - Heterogastridae	Heterogaster tristis n. sp.¹⁰ Heterogaster tristis n. sp.¹⁰ Heer 1864 true bug
Hemiptera - Alydidae	Alydus pulchellus n. sp.⁷, "Nabis gracillima n. sp." = Licremytra gracillima¹⁰ Alydus pulchellus n. sp.⁷ Heer 1853 broad-headed bug lower quarry "Nabis gracillima n. sp." = Licremytra gracillima¹⁰ Heer 1864 broad-headed bug
Hemiptera - Pentatomidae	Halys bruckmanni⁷, Halys spectabilis n. sp.¹⁰, Eurydema arcuata n. sp.⁷, Eurydema impudica⁷, Pentatoma pictum n. sp.¹⁰ Halys bruckmanni⁷ Heer 1853 stink bug from the Kesselstein Halys spectabilis n. sp.¹⁰ Heer 1864 stink bug Eurydema arcuata n. sp.⁷ Heer 1853 stink bug from the Kesselstein Eurydema impudica⁷ Heer 1853 stink bug Pentatoma pictum n. sp.¹⁰ Heer 1864 stink bug
Hemiptera - Scutelleridae	Pachycoris burmeisteri n. sp.¹⁰, Pachycoris protogaeus n. sp.⁷, Pachycoris escheri⁷ Pachycoris burmeisteri n. sp.¹⁰ Heer 1864 metallic shield bug Pachycoris protogaeus n. sp.⁷ Heer 1853 metallic shield bug lower quarry Pachycoris escheri⁷ Heer 1853 metallic shield bug
Hemiptera - Cydnidae	Cydnus oeningensis⁷, Cydnopsis deleta n. sp.⁷, Cydnopsis pygmaea n. sp.⁷, Cydnopsis exilis n. sp.⁷ Cydnus oeningensis⁷ Heer 1853 burrowing bug lower quarry Cydnopsis deleta n. sp.⁷ Heer 1853 burrowing bug lower quarry Cydnopsis pygmaea n. sp.⁷ Heer 1853 burrowing bug upper quarry Cydnopsis exilis n. sp.⁷ Heer 1853 burrowing bug
Hemiptera - Tingidae	"Monanthia wollastoni n. sp." = Dictyla wollastoni¹⁰ "Monanthia wollastoni n. sp." = Dictyla wollastoni¹⁰ Heer 1864 lace bug
Hemiptera - Reduviidae	Harpactor maculipes n. sp.⁷, Harpactor obsoletus n. sp.⁷ Harpactor maculipes n. sp.⁷ Heer 1853 assassin bug lower quarry Harpactor obsoletus n. sp.⁷ Heer 1853 assassin bug from the Kesselstein
Hemiptera - Nabidae	Prostemma oeningensis n. sp.⁷ Prostemma oeningensis n. sp.⁷ Heer 1853 true bug lower quarry
Hemiptera - Gerridae	Limnobates prodromus n. sp.¹⁰ Limnobates prodromus n. sp.¹⁰ Heer 1864 water strider
Thysanoptera -	Thrips oeningensis n. sp.¹⁰, Thrips annosa n. sp.¹⁰ Thrips oeningensis n. sp.¹⁰ Heer 1864 thrips Thrips annosa n. sp.¹⁰ Heer 1864 thrips
Hymenoptera -	Pteromalites oeningensis n. gen. n. sp.¹⁰ Pteromalites oeningensis n. gen. n. sp.¹⁰ Heer 1864 chalcid wasp
Hymenoptera - Ichneumonidae	Ichneumonites bellus n. sp.¹², Ichneumon infernalis n. sp.¹⁰ Ichneumonites bellus n. sp.¹² Heer 1867 ichneumon wasp Ichneumon infernalis n. sp.¹⁰ Heer 1864 ichneumon wasp
Hymenoptera - Scoliidae	Scolia saussureana n. sp.¹⁰ Scolia saussureana n. sp.¹⁰ Heer 1864 hairy flower wasp
Hymenoptera - Formicidae	Imhoffia pallida n. sp.¹², "Formica heraclea" = Camponotites heracleus, "Formica lignitum" = Camponotites lignitus, Formica orbata, "Ponera veneraria n. sp." = Poneropsis veneraria¹⁰, Poneropsis stygia n. sp.¹², "Ponera globosa n. sp." = Poneropsis globosa Imhoffia pallida n. sp.¹² Heer 1864 ant "Formica heraclea" = Camponotites heracleus Heer 1849 ant "Formica lignitum" = Camponotites lignitus Germar 1837 ant Formica orbata Heer 1849 ant "Ponera veneraria n. sp." = Poneropsis veneraria¹⁰ Heer 1864 ant Poneropsis stygia n. sp.¹² Heer 1867 ant "Ponera globosa n. sp." = Poneropsis globosa Heer 1849 ant
Hymenoptera - Sphecidae	Ammophila annosa n. sp.¹⁰, Ammophila inferna n. sp.¹⁰ Ammophila annosa n. sp.¹⁰ Heer 1864 digger wasp Ammophila inferna n. sp.¹⁰ Heer 1864 digger wasp
Hymenoptera - Megachilidae	Megachilidae indet.¹⁷, "Apis adamitica n. sp." = Lithurge adamitica¹⁰, Osmia nigra n. sp.¹⁷ Megachilidae indet.¹⁷ Latreille 1802 mason bee ETH Pl. I, 687; Pl. I, 688(689); Pl. I, 718 "Apis adamitica n. sp." = Lithurge adamitica¹⁰ Heer 1864 mason bee ETH V.S. 4 (Heer's holotype); SN.200 (apparently different from Heer specimens) in Zeuner & Manning, 1976 Osmia nigra n. sp.¹⁷ Zeuner and Manning 1976 mason bee ETH Pl. I, 719; Pl. I, 685
Hymenoptera -	Andrena primaeva n. sp.² Andrena primaeva n. sp.² Cockerell 1909 bee SN.202
Hymenoptera - Apidae	"Bombus atavus n. sp." = Xylocopa atavus¹², "Bombus jurinei n. sp." = Xylocopa jurinei¹², Xylocopa senilis¹⁷, Bombus grandaevus¹², Anthophorites veterana¹⁷, Anthophorites mellona n. gen. n. sp., Anthophorites longaeva¹⁷, Bombus abavus n. sp.¹⁷ "Bombus atavus n. sp." = Xylocopa atavus¹² Heer 1867 large carpenter bee "Bombus jurinei n. sp." = Xylocopa jurinei¹² Heer 1864 large carpenter bee ETH Pl. I, 682a (type in Heer, 1865), 682b (specimen in heer, 1867) Xylocopa senilis¹⁷ Heer 1849 large carpenter bee ETH 21a(b), doutbfully 21c Bombus grandaevus¹² Heer 1849 bee Anthophorites veterana¹⁷ Heer 1849 bee ETH Pl. I, 684 Anthophorites mellona n. gen. n. sp. Heer 1849 bee Anthophorites longaeva¹⁷ Heer 1867 bee ETH Pl. I, 683, 712, 809 (not Heer's types) Bombus abavus n. sp.¹⁷ Heer 1867 bumble bee Holotype (lost), ETH Pl. I, 808 (examined by Cockerell), SN.201
Hymenoptera - Vespidae	Polistes kirbyanus n. sp.³, Polistes primitiva n. sp.¹⁰ Polistes kirbyanus n. sp.³ Cockerell 1915 wasp ETH Pl. I, 681; Osmia kirbyana in Heer's label (unpublished) Polistes primitiva n. sp.¹⁰ Heer 1864 wasp
Coleoptera -	Hydrobius godeti n. sp.⁹, Helophorus magnus n. sp.⁹, Cistelites spectabilis n. sp.¹³, Hydrophilus stenopterus n. sp.⁹ Hydrobius godeti n. sp.⁹ Heer 1862 beetle from Libellenschicht (upper quarry) Helophorus magnus n. sp.⁹ Heer 1862 beetle from Kesselstein Cistelites spectabilis n. sp.¹³ Heer 1869 beetle Hydrophilus stenopterus n. sp.⁹ Heer 1862 beetle from Kesselstein
Coleoptera - Elateridae	"Elater (Alaus) spectabilis n. sp." = Alaus spectabilis¹⁰ "Elater (Alaus) spectabilis n. sp." = Alaus spectabilis¹⁰ Heer 1864 click beetle
Coleoptera - Cantharidae	"Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta¹⁰ "Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta¹⁰ Heer 1864 soldier beetle
Coleoptera - Lampyridae	Lampyris orciluca n. sp.¹⁰ Lampyris orciluca n. sp.¹⁰ Heer 1864 beetle
Coleoptera - Byrrhidae	Byrrhus oeningensis⁹ Byrrhus oeningensis⁹ Heer 1847 pill beetle several from Kesselstein
Coleoptera - Buprestidae	Capnodis antiqua⁹, Capnodis spectabilis n. sp.⁹, Capnodis puncticollis⁹, Chalcophora pulchella n. sp.⁹, Chalcophora laevigata⁹, Buprestites agriloides n. sp.⁹, ? Anthaxia doris n. sp.⁹, Anthaxia pallida n. sp.⁹, "Ancylocheira concinna n. sp." = Buprestis concinna¹⁴, "Ancylochira tincta n. sp." = Buprestis tincta⁹, "Ancylocheira alemanica n. sp." = Buprestis alemanica¹⁴, Protogenia escheri n. gen. n. sp.⁶, Acmaeodera brevicollis n. sp.⁹, Acmaeodera antholitha n. sp.⁹ Capnodis antiqua⁹ Heer 1847 jewel beetle "several" specimens Capnodis spectabilis n. sp.⁹ Heer 1862 jewel beetle from Kesselstein Capnodis puncticollis⁹ Heer 1847 jewel beetle Chalcophora pulchella n. sp.⁹ Heer 1862 jewel beetle from Kesselstein Chalcophora laevigata⁹ Heer 1862 jewel beetle Buprestites agriloides n. sp.⁹ Heer 1862 jewel beetle ? Anthaxia doris n. sp.⁹ Heer 1862 jewel beetle from Kesselstein Anthaxia pallida n. sp.⁹ Heer 1862 jewel beetle from Kesselstein "Ancylocheira concinna n. sp." = Buprestis concinna¹⁴ Heer 1879 jewel beetle "Ancylochira tincta n. sp." = Buprestis tincta⁹ Heer 1862 jewel beetle from Kesselstein,includes var. b. elytris majoribus "Ancylocheira alemanica n. sp." = Buprestis alemanica¹⁴ Heer 1879 jewel beetle Protogenia escheri n. gen. n. sp.⁶ Heer 1847 jewel beetle Acmaeodera brevicollis n. sp.⁹ Heer 1862 jewel beetle from Kesselstein Acmaeodera antholitha n. sp.⁹ Heer 1862 jewel beetle from Kesselstein
Coleoptera - Glaphyridae	Glaphyrus antiquus n. sp.⁹ Glaphyrus antiquus n. sp.⁹ Heer 1862 bumble bee scarab beetle from Insektenschicht (lower quarry)
Coleoptera - Geotrupidae	Geotrupes germari n. sp.⁹ Geotrupes germari n. sp.⁹ Heer 1862 earth-boring dung beetle from Kesselstein
Coleoptera -	Gymnopleurus deperditus n. sp.⁹ Gymnopleurus deperditus n. sp.⁹ Heer 1862 scarab beetle from Kesselstein
Coleoptera - Scarabaeidae	Aphodius brevipennis n. sp.⁹, Valgus oeningensis n. sp.⁹, "Trichius aedilis n. sp." = Gnorimus aedilis⁹, Trichius lugubris n. sp.⁹, Trichius unifasciatus n. sp.⁹, ? Trichius rotundatus n. sp.⁹, "Scarabaeus proserpinae n. sp." = Pentodon proserpinae⁹, Serica minutula n. sp.⁹, Lepitrix germanica n. sp.⁹, Copris druidum n. sp.⁹, Onthophagus crassus n. sp.⁹, Onthophagus prodromus n. sp.⁹, Onthophagus bisontinus n. sp.⁹, Gymnopleurus rotundatus n. sp.⁹, Gymnopleurus sisyphus n. sp.⁶, Anomala fugax n. sp.⁹, Oniticellus amplicollis n. sp.⁹ Aphodius brevipennis n. sp.⁹ Heer 1862 scarab beetle from Kesselstein Valgus oeningensis n. sp.⁹ Heer 1862 flower chafer from Kesselstein "Trichius aedilis n. sp." = Gnorimus aedilis⁹ Heer 1862 flower chafer from Kesselstein Trichius lugubris n. sp.⁹ Heer 1862 flower chafer Trichius unifasciatus n. sp.⁹ Heer 1862 flower chafer from Kesselstein ? Trichius rotundatus n. sp.⁹ Heer 1862 flower chafer from Kesselstein "Scarabaeus proserpinae n. sp." = Pentodon proserpinae⁹ Heer 1862 rhinoceras beetle from Kesselstien Serica minutula n. sp.⁹ Heer 1862 scarab beetle from Kesselstein Lepitrix germanica n. sp.⁹ Heer 1862 scarab beetle Copris druidum n. sp.⁹ Heer 1862 scarab beetle from Kesselstein Onthophagus crassus n. sp.⁹ Heer 1862 scarab beetle from Kesselstein Onthophagus prodromus n. sp.⁹ Heer 1862 scarab beetle several from Kesselstein Onthophagus bisontinus n. sp.⁹ Heer 1862 scarab beetle from Insektenschicht (lower quarry) Gymnopleurus rotundatus n. sp.⁹ Heer 1862 scarab beetle from Kesselstein Gymnopleurus sisyphus n. sp.⁶ Heer 1847 scarab beetle Anomala fugax n. sp.⁹ Heer 1862 scarab beetle from Kesselstein Oniticellus amplicollis n. sp.⁹ Heer 1862 scarab beetle from Kesselstein
Coleoptera -	"Onthophagus urus" = Onthophagus urusheeri⁹, "Copris subterranea n. sp." = Copris subterraneus⁹ "Onthophagus urus" = Onthophagus urusheeri⁹ Krell 2000 scarab beetle "several" specimens "Copris subterranea n. sp." = Copris subterraneus⁹ Heer 1862 scarab beetle from Kesselstein
Coleoptera - Hybosoridae	Hybosorus lividus n. sp.⁹ Hybosorus lividus n. sp.⁹ Heer 1862 scavenger scarab beetle
Coleoptera - Staphylinidae	Silpha tricostata⁹, Lathrobium oeningense n. sp.⁹, Bledius speciosus n. sp.⁹, Oxytelus proaevus n. sp.⁹, Scaphisoma gracile n. sp.⁹, Scaphidium deletum⁹, Staphylinus atavus n. sp.⁹, ? Oxyporus seuberti n. sp.⁹, Protactus minor n. sp.⁹ Silpha tricostata⁹ Heer 1847 rove beetle from Kesselstein Lathrobium oeningense n. sp.⁹ Heer 1862 rove beetle from Insektenschicht (lower quarry) Bledius speciosus n. sp.⁹ Heer 1862 rove beetle from Insektenschicht (lower quarry) Oxytelus proaevus n. sp.⁹ Heer 1862 rove beetle from Insektenschicht (lower quarry) Scaphisoma gracile n. sp.⁹ Heer 1862 rove beetle 2 from Kesselstein, 1 from from Insektenschicht (lower quarry) Scaphidium deletum⁹ Heer 1847 rove beetle Staphylinus atavus n. sp.⁹ Heer 1862 rove beetle from Insektenschicht (lower quarry) ? Oxyporus seuberti n. sp.⁹ Heer 1862 rove beetle Protactus minor n. sp.⁹ Heer 1862 rove beetle from Insektenschicht (lower quarry)
Coleoptera - Histeridae	Hister maculigerus n. sp.⁹, Hister aemulus n. sp.⁹, Hister marmoratus n. sp.⁹, Hister mastodontis n. sp.⁹, Hister coprolithorum n. sp.⁹, Hister vetustus n. sp.⁹, Hister antiquus n. sp.⁹, Hister morosus n. sp.⁹ Hister maculigerus n. sp.⁹ Heer 1862 clown beetle from Kesselstein Hister aemulus n. sp.⁹ Heer 1862 clown beetle several from Kesselstein Hister marmoratus n. sp.⁹ Heer 1862 clown beetle from Kesselstein Hister mastodontis n. sp.⁹ Heer 1862 clown beetle from Kesselstein Hister coprolithorum n. sp.⁹ Heer 1862 clown beetle numerous specimens from Kesselstein Hister vetustus n. sp.⁹ Heer 1862 clown beetle several from Kesselstein Hister antiquus n. sp.⁹ Heer 1862 clown beetle from Kesselstein Hister morosus n. sp.⁹ Heer 1862 clown beetle from Insektenschicht (lower quarry)
Coleoptera - Hydrophilidae	Hydrobius couloni n. sp.⁹, Hydrous ovalis n. sp.⁹, "Hydrous rehmanni" = Hydrochara noachica⁹, "Hydrous braunii" = Hydrobiomorpha braunii⁹, "Hydrous escheri n. sp." = Hydrobiomorpha braunii⁹, Hydrobiomorpha heeri n. sp.⁴, "Hydrophilus spectabilis" = Hydrophilus spectabilis⁶, "Hydrophilus giganteus n. sp." = Hydrophilus spectabilis⁹, Hydrophilopsis elongata n. gen. n. sp.⁹, Escheria bella n. sp.⁹ Hydrobius couloni n. sp.⁹ Heer 1862 water scavenger beetle from Kesselstein Hydrous ovalis n. sp.⁹ Heer 1862 water scavenger beetle from Kesselstein "Hydrous rehmanni" = Hydrochara noachica⁹ Heer 1847 water scavenger beetle "Hydrous braunii" = Hydrobiomorpha braunii⁹ Heer 1847 water scavenger beetle from Kesselstein and Insektenschicht (lower quarry) "Hydrous escheri n. sp." = Hydrobiomorpha braunii⁹ Heer 1847 water scavenger beetle from Insektenschicht (lower quarry); ETHZ NO-Z51.3-39-4:4-Pl.I148 Hydrobiomorpha heeri n. sp.⁴ Fikáček and Schmied 2013 water scavenger beetle ETHZ NO-Z51.3-39-4:4-Pl. I 149b (label reads Hydrophilus ovalis) "Hydrophilus spectabilis" = Hydrophilus spectabilis⁶ Heer 1847 water scavenger beetle "Hydrophilus giganteus n. sp." = Hydrophilus spectabilis⁹ Heer 1847 water scavenger beetle from Kesselstein; ETHZ NO-Z51.3-39-4:4-Pl.I143b Hydrophilopsis elongata n. gen. n. sp.⁹ Heer 1862 water scavenger beetle from Insektenschicht (lower quarry) Escheria bella n. sp.⁹ Heer 1862 water scavenger beetle
Coleoptera - Trogossitidae	"Gymnochila obesa n. sp." = Gymnocheilis obesa⁹, Trogosita koellikeri⁹, Trogosita longicollis n. sp.⁹, Trogosita assimilis n. sp.⁹, Trogosita sculpturata n. sp.⁹, Trogosita amissa n. sp.⁹, Trogosita bella n. sp.⁹ "Gymnochila obesa n. sp." = Gymnocheilis obesa⁹ Heer 1862 bark gnawing beetle from Kesselstein Trogosita koellikeri⁹ Heer 1847 bark gnawing beetle Trogosita longicollis n. sp.⁹ Heer 1862 bark gnawing beetle from Kesselstein Trogosita assimilis n. sp.⁹ Heer 1862 bark gnawing beetle from Kesselstein Trogosita sculpturata n. sp.⁹ Heer 1862 bark gnawing beetle from Kesselstein Trogosita amissa n. sp.⁹ Heer 1862 bark gnawing beetle from Kesselstein and Insektenschicht (lower quarry) Trogosita bella n. sp.⁹ Heer 1862 bark gnawing beetle
Coleoptera - Tenebrionidae	Tagenopsis brevicornis n. gen. n. sp.¹⁰ Tagenopsis brevicornis n. gen. n. sp.¹⁰ Heer 1864 darkling beetle
Coleoptera - Meloidae	Lytta aesculapii⁶, Zonites vetusta n. sp.¹⁰ Lytta aesculapii⁶ Heer 1847 blister beetle Zonites vetusta n. sp.¹⁰ Heer 1864 blister beetle
Coleoptera - Nitidulidae	Nitidula pallida n. sp.⁹, Nitidula ancora n. sp.⁹, Nitidula maculigera n. sp.⁹ Nitidula pallida n. sp.⁹ Heer 1862 sap beetle Nitidula ancora n. sp.⁹ Heer 1862 sap beetle from Kesselstein Nitidula maculigera n. sp.⁹ Heer 1862 sap beetle
Coleoptera - Lymexylidae	Hylecoetus cylindricus n. sp.¹⁰ Hylecoetus cylindricus n. sp.¹⁰ Heer 1864 ship-timber beetle
Coleoptera - Coccinellidae	"Coccinella decempustulata n. sp." = Coccinella heeri¹⁴, Coccinella amabilis n. sp.¹⁰, "Coccinella spectabilis n. sp." = Coccinella ponomarenkoi¹⁰, Coccinella colorata n. sp.¹⁰ "Coccinella decempustulata n. sp." = Coccinella heeri¹⁴ Ukrainsky 2010 ladybird beetle Written "Coccinella 10-pustulata" Coccinella amabilis n. sp.¹⁰ Heer 1864 ladybird beetle "Coccinella spectabilis n. sp." = Coccinella ponomarenkoi¹⁰ Ukrainsky 2010 ladybird beetle Coccinella colorata n. sp.¹⁰ Heer 1864 ladybird beetle
Coleoptera - Cerambycidae	"Prionus polyphemus n. sp." = Ergates polyphemus¹⁰, "Prionus spectabilis n. sp." = Ergates spectabilis¹⁰, Acanthoderus lepidus n. sp.¹⁰, Acanthoderus sepultus n. sp.¹⁰, Lamia antiqua n. sp.¹⁴, Callidium escheri n. sp.¹⁰, Callidium procerum n. sp.¹⁰, Clytus pulcher n. sp.¹⁰ "Prionus polyphemus n. sp." = Ergates polyphemus¹⁰ Heer 1864 longhorned beetle "Prionus spectabilis n. sp." = Ergates spectabilis¹⁰ Heer 1864 longhorned beetle Acanthoderus lepidus n. sp.¹⁰ Heer 1864 longhorned beetle Acanthoderus sepultus n. sp.¹⁰ Heer 1864 longhorned beetle Lamia antiqua n. sp.¹⁴ Heer 1879 longhorned beetle Callidium escheri n. sp.¹⁰ Heer 1864 longhorned beetle Callidium procerum n. sp.¹⁰ Heer 1864 longhorned beetle Clytus pulcher n. sp.¹⁰ Heer 1864 longhorned beetle
Coleoptera - Chrysomelidae	Galeruca gemmifera n. sp.¹⁴, Galeruca buchi n. sp.¹⁰, "Lema vetusta n. sp." = Crioceris vetusta¹⁰, Gonioctena japeti n. sp.⁶ Galeruca gemmifera n. sp.¹⁴ Heer 1879 leaf beetle Galeruca buchi n. sp.¹⁰ Heer 1864 leaf beetle "Lema vetusta n. sp." = Crioceris vetusta¹⁰ Heer 1864 leaf beetle Gonioctena japeti n. sp.⁶ Heer 1847 leaf beetle
Coleoptera - Attelabidae	"Attelabus durus n. sp." = Phialodes durus¹⁰ "Attelabus durus n. sp." = Phialodes durus¹⁰ Heer 1864 leaf rolling weevil
Coleoptera - Brentidae	Antliarhinites gracilis n. gen. n. sp.¹⁰, Apion antiquum n. sp.¹⁰, "Rhynchites dionysus n. sp." = Omphalapion dionysus¹⁰, Miocenocylas heeri n. gen. n. sp.¹⁵ Antliarhinites gracilis n. gen. n. sp.¹⁰ Heer 1864 weevil Apion antiquum n. sp.¹⁰ Heer 1864 weevil "Rhynchites dionysus n. sp." = Omphalapion dionysus¹⁰ Heer 1864 weevil Miocenocylas heeri n. gen. n. sp.¹⁵ Legalov 2018 weevil
Coleoptera - Curculionidae	Lixus oeningensis n. sp.¹⁰, Naupactus crassirostris n. sp.¹⁰, Sitona atavina n. sp.¹⁰, Cleonus speciosus n. sp.¹⁰, Brachycerus nanus n. sp.¹⁰ Lixus oeningensis n. sp.¹⁰ Heer 1864 snout beetle Naupactus crassirostris n. sp.¹⁰ Heer 1864 snout beetle Sitona atavina n. sp.¹⁰ Heer 1864 snout beetle Cleonus speciosus n. sp.¹⁰ Heer 1864 snout beetle Brachycerus nanus n. sp.¹⁰ Heer 1864 snout beetle
Coleoptera - Dytiscidae	"Hydroporus benzeli n. sp." = Nebrioporus benzeli⁹, "Hydaticus areolatus n. sp." = Graphoderus heeri⁹, Dytiscus lavateri⁹, Cybister agassizi n. sp.⁹, Cybister atavus n. sp.⁹, "Cybister nicoleti" = Megadytes nicoleti⁹, Colymbetes aemulus n. sp.⁹ "Hydroporus benzeli n. sp." = Nebrioporus benzeli⁹ Heer 1862 predaceous diving beetle "Hydaticus areolatus n. sp." = Graphoderus heeri⁹ Nilsson 2001 predaceous diving beetle Dytiscus lavateri⁹ Heer 1847 predaceous diving beetle from Kesselstein Cybister agassizi n. sp.⁹ Heer 1862 predaceous diving beetle from Kesselstein Cybister atavus n. sp.⁹ Heer 1862 predaceous diving beetle from Kesselstein "Cybister nicoleti" = Megadytes nicoleti⁹ Heer 1862 predaceous diving beetle from Kesselstein Colymbetes aemulus n. sp.⁹ Heer 1862 predaceous diving beetle from Kesselstein
Coleoptera - Carabidae	Calosoma escheri n. sp.⁸, "Calosoma deplanatum n. sp." = Campalita deplanatum⁸, Calosoma escrobiculatum n. sp.⁸, Calosoma nauckianum n. sp.⁸, Calosoma catenulatum n. sp.⁸, Sinis brevicollis n. gen. n. sp.⁹, Badister grandis n. sp.⁹, Badister fragilis n. sp.⁹, Badister macrocephalus n. sp.⁹, Pterostichus minutulus n. sp.⁹, Pterostichus vetustus n. sp.⁹, Dichirotrichus lividus⁹, Harpalus tardigradus n. sp.⁹, Harpalus stygius n. sp.⁹, Harpalus stierlini n. sp.⁹, Harpalus constrictus n. sp.⁹, Amara princeps n. sp.⁹, Amara pinguicula n. sp.⁹, Amara primigenia n. sp.⁹, Trechinites oblongus n. sp.⁹, Trechinites clairvillii n. sp.⁹, Glenopterus laevigatus n. gen. n. sp.⁶, Carabites laesicollis n. sp.⁹, Nebria pluto n. sp.⁹ Calosoma escheri n. sp.⁸ Heer 1860 ground beetle "Calosoma deplanatum n. sp." = Campalita deplanatum⁸ Heer 1860 ground beetle Calosoma escrobiculatum n. sp.⁸ Heer 1860 ground beetle Calosoma nauckianum n. sp.⁸ Heer 1860 ground beetle Calosoma catenulatum n. sp.⁸ Heer 1860 ground beetle Kesselstein Sinis brevicollis n. gen. n. sp.⁹ Heer 1862 ground beetle 1 male, 1 female from lower quarry Badister grandis n. sp.⁹ Heer 1862 ground beetle from Kesselstein Badister fragilis n. sp.⁹ Heer 1862 ground beetle from Kesselstein (2 var. paulo major) Badister macrocephalus n. sp.⁹ Heer 1862 ground beetle "multiple examples" in Kesselstein, including var. major Pterostichus minutulus n. sp.⁹ Heer 1862 ground beetle from Insektenschicht (lower quarry) Pterostichus vetustus n. sp.⁹ Heer 1862 ground beetle from Insektenschicht (lower quarry) Dichirotrichus lividus⁹ Heer 1862 ground beetle from Kesselstein Harpalus tardigradus n. sp.⁹ Heer 1862 ground beetle from Insektenschicht (lower quarry) Harpalus stygius n. sp.⁹ Heer 1862 ground beetle from Insektenschicht (lower quarry) Harpalus stierlini n. sp.⁹ Heer 1862 ground beetle Harpalus constrictus n. sp.⁹ Heer 1862 ground beetle from Kesselstein Amara princeps n. sp.⁹ Heer 1862 ground beetle 1 from Insektenschicht (lower quarry), 1 from Kesselstein Amara pinguicula n. sp.⁹ Heer 1862 ground beetle from Insektenschicht (lower quarry) Amara primigenia n. sp.⁹ Heer 1862 ground beetle from Kesselstein Trechinites oblongus n. sp.⁹ Heer 1862 ground beetle from lower quarry Trechinites clairvillii n. sp.⁹ Heer 1862 ground beetle from lower quarry Glenopterus laevigatus n. gen. n. sp.⁶ Heer 1847 ground beetle Carabites laesicollis n. sp.⁹ Heer 1862 ground beetle Nebria pluto n. sp.⁹ Heer 1862 ground beetle 1 from lower quarry, 2 from Kesselstein
Coleoptera - Gyrinidae	"Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris⁹, Miodineutes oeningensis n. gen. n. sp.⁵, "Dineutus insignis n. sp." = Miodineutes insignis⁹, Miodineutes heeri n. sp.⁵ "Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris⁹ Heer 1862 whirligig beetle from Kesselstein Miodineutes oeningensis n. gen. n. sp.⁵ Hatch 1927 whirligig beetle Dineutus insignis specimen in pl 2, fig 28 of Heer "Dineutus insignis n. sp." = Miodineutes insignis⁹ Heer 1862 whirligig beetle from Kesselstein Miodineutes heeri n. sp.⁵ Hatch 1927 whirligig beetle Dineutus longiventris Specimen in pl 2, fig 25 of Heer
Lepidoptera -	"Bombycites" oeningensis n. sp., Bombycites buchii n. sp.¹⁰ "Bombycites" oeningensis n. sp. Heer 1849 moth pupa Bombycites buchii n. sp.¹⁰ Heer 1864 moth
Diptera - Chironomidae	Chironomus gaudini n. sp.¹⁰ Chironomus gaudini n. sp.¹⁰ Heer 1864 non biting midge
Diptera - Syrphidae	Syrphus schellenbergi n. sp.¹⁰, Syrphus bremii n. sp.¹⁰ Syrphus schellenbergi n. sp.¹⁰ Heer 1864 hover fly Syrphus bremii n. sp.¹⁰ Heer 1864 hover fly
Diptera - Oestridae	"Dipterites obovatus n. sp." = Adipterites obovatus¹⁰ "Dipterites obovatus n. sp." = Adipterites obovatus¹⁰ Heer 1864 bot fly
Diptera - Tabanidae	"? Hexatoma oeningensis n. sp." = Heptatoma oeningensis¹⁰ "? Hexatoma oeningensis n. sp." = Heptatoma oeningensis¹⁰ Heer 1864 horse fly
Diptera - Sciaridae	Sciara deleta n. sp.¹⁰ Sciara deleta n. sp.¹⁰ Heer 1864 dark-winged fungus gnat
Diptera - Cecidomyiidae	Cecidomyia bremii n. sp.¹⁰ Cecidomyia bremii n. sp.¹⁰ Heer 1864 gall midge
Diptera - Mycetophilidae	Mycetophila orci n. sp.¹⁰, Mycetophila pusillima n. sp.¹⁰ Mycetophila orci n. sp.¹⁰ Heer 1864 fungus gnat Mycetophila pusillima n. sp.¹⁰ Heer 1864 fungus gnat
Diptera - Bibionidae	Bibio elongatus¹⁶, Plecia macilenta n. sp.¹⁶, "Protomyia jucunda" = Plecia jucunda, Plecia hilaris¹⁶, Bibio pulchellus¹⁶, Bibio moestus¹⁶, Bibio brevis¹⁶, Bibio crassinervis n. sp.¹⁶, Bibio fusiformis¹⁶, Bibio angustatus¹⁶, Dilophus pumilio n. sp.¹⁶, "Protomyia speciosa n. sp." = Penthetria amoena¹⁰, Penthetria brevicollis¹⁶ Bibio elongatus¹⁶ Heer 1849 march fly ETHZ S.N. 189 Pl. I 863 (labeled by Heer) Plecia macilenta n. sp.¹⁶ Skartveit and Pika 2014 love bug ETHZ Pl. I 853, 853 female "Protomyia jucunda" = Plecia jucunda Heer 1849 love bug ETHZ Pl. I 871, 853(x2), 856, 861, 867, 869 (some labeled by Brongniart fide Skartveit & Pika 2014) Plecia hilaris¹⁶ Heer 1849 love bug ETHZ Pl. I 850 Bibio pulchellus¹⁶ Heer 1849 march fly ETHZ Pl. I 854 Bibio moestus¹⁶ Heer 1849 march fly ETHZ Pl. I 855, 857(x2) Bibio brevis¹⁶ Heer 1849 march fly Some in lot Pl. I 856; ETHZ Pl.I 865, Pl. I 887 Bibio crassinervis n. sp.¹⁶ Skartveit and Pika 2014 march fly ETHZ Pl. I 851, Pl.I 851(2), Pl. I 862, Pl. I 875, Pl. I 888 Bibio fusiformis¹⁶ Heer 1849 march fly ETHZ Pl. I 858 (x2), 861 (x6) Bibio angustatus¹⁶ Heer 1849 march fly ETHZ Pl. I 851 Dilophus pumilio n. sp.¹⁶ Skartveit and Pika 2014 march fly ETHZ Pl. I 886 "Protomyia speciosa n. sp." = Penthetria amoena¹⁰ Heer 1849 march fly ETHZ Pl. I 873; also Pl. I 872, 876 as P. amoena in Skartveit & Pika, 2014 Penthetria brevicollis¹⁶ Heer 1849 march fly ETHZ Pl. I 870

¹Boderau et al. 2024; ²Cockerell 1909; ³Cockerell 1915; ⁴Fikáček and Schmied 2013; ⁵Hatch 1927; ⁶Heer 1847; ⁷Heer 1853; ⁸Heer 1860; ⁹Heer 1862; ¹⁰Heer 1864; ¹¹Heer 1864; ¹²Heer 1867; ¹³Heer 1869; ¹⁴Heer 1879; ¹⁵Legalov 2018; ¹⁶Skartveit and Pika 2014; ¹⁷Zeuner and Manning 1976