Oeningen (Zurich collection) (Miocene of Germany)

Where: Baden-Württemberg, Germany (47.7° N, 8.9° E: paleocoordinates 47.9° N, 7.8° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 7 mammal zone, Upper Oehningen beds Member (Upper Freshwater-Molasse Formation), Sarmatian (12.7 - 11.6 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified limestone

• "Rising into massive mountains, as in the well-known Righi and Rossberg, they attain a thickness of more than 6000 feet. While they include proofs of the presenceof the sea, they have preserved with marvellous perfection a large number of the plants which clothed the Alps, and of the insects which flitted through the woodlands. They form part of a great series of deposits which have been termed “Molasse” by the Swiss geologists." Geikie, 1882

•"...deposited in the Northern Alpine Foreland basin..." Uhl et al., 2006

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Collection methods: Collection of Zurich University

Primary reference: O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 117252: authorized by Matthew Clapham, entered by Matthew Clapham on 18.09.2011, edited by Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Hemiptera -
Halys spectabilis n. sp.10 Heer 1865 true bug
Halys bruckmanni6 Heer 1853 true bug
from the Kesselstein
 Hemiptera - Cercopidae
Cercopis herrichi n. sp.10, Cercopis hageni n. sp.10, Cercopis germari n. sp.10
Cercopis herrichi n. sp.10 Heer 1865 froghopper
Cercopis hageni n. sp.10 Heer 1865 froghopper
Cercopis germari n. sp.10 Heer 1865 froghopper
 Hemiptera - Alydidae
Alydus pulchellus n. sp.6 Heer 1853 broad-headed bug
lower quarry
 Hemiptera - Lygaeidae
Heterogaster tristis n. sp.10 Heer 1865 milkweed bug
Pachymerus obsoletus n. sp.6 Heer 1853 milkweed bug
lower quarry
Cephalocoris pilosus n. gen. n. sp.6 Heer 1853 milkweed bug
lower quarry
 Hemiptera - Scutelleridae
Pachycoris protogaeus n. sp.6 Heer 1853 metallic shield bug
lower quarry
Pachycoris escheri6 Heer 1853 metallic shield bug
Pachycoris burmeisteri n. sp.10 Heer 1865 metallic shield bug
 Hemiptera - Pentatomidae
Eurydema arcuata n. sp.6 Heer 1853 stink bug
from the Kesselstein
Eurydema impudica6 Heer 1853 stink bug
Pentatoma pictum n. sp.10 Heer 1865 stink bug
 Hemiptera - Cydnidae
Cydnus oeningensis6 Heer 1853 burrowing bug
lower quarry
Cydnopsis pygmaea n. sp.6 Heer 1853 burrowing bug
upper quarry
Cydnopsis exilis n. sp.6 Heer 1853 burrowing bug
Cydnopsis deleta n. sp.6 Heer 1853 burrowing bug
lower quarry
 Hemiptera - Tingidae
"Monanthia wollastoni n. sp." = Dictyla wollastoni10
"Monanthia wollastoni n. sp." = Dictyla wollastoni10 Heer 1865 lace bug
 Hemiptera - Reduviidae
Prostemma oeningensis n. sp.6 Heer 1853 assassin bug
lower quarry
Harpactor obsoletus n. sp.6 Heer 1853 assassin bug
from the Kesselstein
Harpactor maculipes n. sp.6 Heer 1853 assassin bug
lower quarry
Nabis gracillima n. sp.10 Heer 1865 assassin bug
 Hemiptera - Gerridae
Limnobates prodromus n. sp.10 Heer 1865 water strider
 Hemiptera - Belostomatidae
"Belostoma speciosum n. sp." = Belostomates speciosum10
"Belostoma speciosum n. sp." = Belostomates speciosum10 Heer 1865 giant water bug
 Hemiptera - Corixidae
"Corisa fasciolata" = Corixa fasciolata6
"Corisa fasciolata" = Corixa fasciolata6 Heer 1853 water boatman
from the Libellen-schicht
 Thysanoptera -
Thrips annosa n. sp.10, Thrips oeningensis n. sp.10
Thrips annosa n. sp.10 Heer 1865 thrips
Thrips oeningensis n. sp.10 Heer 1865 thrips
 Blattodea - Rhinotermitidae
"Termes hartungi n. sp." = Reticulitermes hartungi10
"Termes hartungi n. sp." = Reticulitermes hartungi10 Heer 1865 termite
 Blattodea - Termitidae
Termes buchii n. sp.10
Termes buchii n. sp.10 Heer 1865 termite
 Blattodea - Blattoidae
"Blattidium coloratum n. sp." = Blatta colorata9
"Blattidium coloratum n. sp." = Blatta colorata9 Heer 1864 cockroach
from Kesselstein
 Dermaptera - Forficulidae
Forficula recta n. sp.10, Forficula minuta n. sp.10
Forficula recta n. sp.10 Heer 1865 common earwig
Forficula minuta n. sp.10 Heer 1865 common earwig
 Orthoptera - Acrididae
Oedipoda germari n. sp.10 Heer 1865 short-horned grasshopper
 Orthoptera - Pyrgomorphidae
"Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri10
"Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri10 Heer 1865 gaudy grasshopper
 Orthoptera - Tetrigidae
"Tetrix gracilis n. sp." = Tettigidea gracilis10
"Tetrix gracilis n. sp." = Tettigidea gracilis10 Heer 1865 pygmy grasshopper
 Orthoptera - Gryllidae
Gryllus troglodytes n. sp.10 Heer 1865 true crickets
 Orthoptera - Gryllotalpidae
Gryllotalpa stricta n. sp.10 Heer 1865 mole cricket
 Orthoptera - Tettigoniidae
"Decticus speciosus n. sp." = Platycleis speciosa10
"Decticus speciosus n. sp." = Platycleis speciosa10 Heer 1865 katydid
 Hymenoptera - Scoliidae
Scolia saussureana n. sp.10 Heer 1865 wasp
 Hymenoptera - Vespidae
Polistes kirbyanus n. sp.2 Cockerell 1915 wasp
ETH Pl. I, 681; Osmia kirbyana in Heer's label (unpublished)
Polistes primitiva n. sp.10 Heer 1865 wasp
 Hymenoptera - Formicidae
Formica orbata, "Formica lignitum" = Camponotus lignitus, "Formica heraclea" = Camponotus heracleus, Imhoffia pallida n. sp.11, Poneropsis stygia n. sp.11, "Ponera veneraria n. sp." = Poneropsis veneraria10, "Ponera globosa n. sp." = Poneropsis globosa
Formica orbata Heer 1849 ant
"Formica lignitum" = Camponotus lignitus Germar 1837 ant
"Formica heraclea" = Camponotus heracleus Heer 1849 ant
Imhoffia pallida n. sp.11 Heer 1865 ant
Poneropsis stygia n. sp.11 Heer 1867 ant
"Ponera veneraria n. sp." = Poneropsis veneraria10 Heer 1865 ant
"Ponera globosa n. sp." = Poneropsis globosa Heer 1849 ant
 Hymenoptera - Megachilidae
Megachilidae indet.16, "Apis adamitica n. sp." = Lithurge adamitica10, Osmia nigra n. sp.16
Megachilidae indet.16 Latreille 1802 mason bee
ETH Pl. I, 687; Pl. I, 688(689); Pl. I, 718
"Apis adamitica n. sp." = Lithurge adamitica10 Heer 1865 mason bee
ETH V.S. 4 (Heer's holotype); SN.200 (apparently different from Heer specimens) in Zeuner & Manning, 1976
Osmia nigra n. sp.16 Zeuner and Manning 1976 mason bee
ETH Pl. I, 719; Pl. I, 685
 Hymenoptera - Apidae
"Bombus atavus n. sp." = Xylocopa atavus11, Xylocopa senilis16, "Bombus jurinei n. sp." = Xylocopa jurinei11, Bombus abavus n. sp.16, Anthophorites mellona n. gen. n. sp., Anthophorites longaeva16, Anthophorites veterana16, Bombus grandaevus11
"Bombus atavus n. sp." = Xylocopa atavus11 Heer 1867 large carpenter bee
Xylocopa senilis16 Heer 1849 large carpenter bee
ETH 21a(b), doutbfully 21c
"Bombus jurinei n. sp." = Xylocopa jurinei11 Heer 1865 large carpenter bee
ETH Pl. I, 682a (type in Heer, 1865), 682b (specimen in heer, 1867)
Bombus abavus n. sp.16 Heer 1867 bumble bee
Holotype (lost), ETH Pl. I, 808 (examined by Cockerell), SN.201
Anthophorites mellona n. gen. n. sp. Heer 1849 bee
Anthophorites longaeva16 Heer 1867 bee
ETH Pl. I, 683, 712, 809 (not Heer's types)
Anthophorites veterana16 Heer 1849 bee
ETH Pl. I, 684
Bombus grandaevus11 Heer 1849 bee
 Hymenoptera - Andrenidae
Andrena primaeva n. sp.1 Cockerell 1909 bee
SN.202
 Hymenoptera - Sphecidae
Ammophila annosa n. sp.10 Heer 1865 digger wasp
Ammophila inferna n. sp.10 Heer 1865 digger wasp
 Hymenoptera - Ichneumonidae
Ichneumonites bellus n. sp.11 Heer 1867 ichneumon wasp
Ichneumon infernalis n. sp.10 Heer 1865 ichneumon wasp
 Hymenoptera -
Pteromalites oeningensis n. gen. n. sp.10
Pteromalites oeningensis n. gen. n. sp.10 Heer 1865 chalcid wasp
 Coleoptera - Buprestidae
Protogenia escheri n. gen. n. sp.5, Anthaxia pallida n. sp.8, ? Anthaxia doris n. sp.8, "Ancylocheira concinna n. sp." = Buprestis concinna13, "Ancylochira tincta n. sp." = Buprestis tincta8, "Ancylocheira alemanica n. sp." = Buprestis alemanica13, Buprestites agriloides n. sp.8, Acmaeodera brevicollis n. sp.8, Acmaeodera antholitha n. sp.8, Capnodis puncticollis8, Capnodis spectabilis n. sp.8, Capnodis antiqua8, Chalcophora laevigata8, Chalcophora pulchella n. sp.8
Protogenia escheri n. gen. n. sp.5 Heer 1847 jewel beetle
Anthaxia pallida n. sp.8 Heer 1862 jewel beetle
from Kesselstein
? Anthaxia doris n. sp.8 Heer 1862 jewel beetle
from Kesselstein
"Ancylocheira concinna n. sp." = Buprestis concinna13 Heer 1879 jewel beetle
"Ancylochira tincta n. sp." = Buprestis tincta8 Heer 1862 jewel beetle
from Kesselstein,includes var. b. elytris majoribus
"Ancylocheira alemanica n. sp." = Buprestis alemanica13 Heer 1879 jewel beetle
Buprestites agriloides n. sp.8 Heer 1862 jewel beetle
Acmaeodera brevicollis n. sp.8 Heer 1862 jewel beetle
from Kesselstein
Acmaeodera antholitha n. sp.8 Heer 1862 jewel beetle
from Kesselstein
Capnodis puncticollis8 Heer 1847 jewel beetle
Capnodis spectabilis n. sp.8 Heer 1862 jewel beetle
from Kesselstein
Capnodis antiqua8 Heer 1847 jewel beetle
"several" specimens
Chalcophora laevigata8 Heer 1862 jewel beetle
Chalcophora pulchella n. sp.8 Heer 1862 jewel beetle
from Kesselstein
 Coleoptera - Byrrhidae
Byrrhus oeningensis8 Heer 1847 pill beetle
several from Kesselstein
 Coleoptera - Lampyridae
Lampyris orciluca n. sp.10 Heer 1865 beetle
 Coleoptera - Cantharidae
"Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta10
"Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta10 Heer 1865 soldier beetle
 Coleoptera - Elateridae
"Elater (Alaus) spectabilis n. sp." = Alaus spectabilis10
"Elater (Alaus) spectabilis n. sp." = Alaus spectabilis10 Heer 1865 click beetle
 Coleoptera - Hydrophilidae
Escheria bella n. sp.8, Hydrophilopsis elongata n. gen. n. sp.8, "Hydrous braunii" = Hydrobiomorpha braunii8, "Hydrous escheri n. sp." = Hydrobiomorpha braunii8, Hydrobiomorpha heeri n. sp.3, "Hydrophilus spectabilis" = Hydrophilus spectabilis5, "Hydrophilus giganteus n. sp." = Hydrophilus spectabilis8, "Hydrous rehmanni" = Hydrochara noachica8, Hydrous ovalis n. sp.8, Hydrobius couloni n. sp.8
Escheria bella n. sp.8 Heer 1862 water scavenger beetle
Hydrophilopsis elongata n. gen. n. sp.8 Heer 1862 water scavenger beetle
from Insektenschicht (lower quarry)
"Hydrous braunii" = Hydrobiomorpha braunii8 Heer 1847 water scavenger beetle
from Kesselstein and Insektenschicht (lower quarry)
"Hydrous escheri n. sp." = Hydrobiomorpha braunii8 Heer 1847 water scavenger beetle
from Insektenschicht (lower quarry); ETHZ NO-Z51.3-39-4:4-Pl.I148
Hydrobiomorpha heeri n. sp.3 Fikacek and Schmied 2013 water scavenger beetle
ETHZ NO-Z51.3-39-4:4-Pl. I 149b (label reads Hydrophilus ovalis)
"Hydrophilus spectabilis" = Hydrophilus spectabilis5 Heer 1847 water scavenger beetle
"Hydrophilus giganteus n. sp." = Hydrophilus spectabilis8 Heer 1847 water scavenger beetle
from Kesselstein; ETHZ NO-Z51.3-39-4:4-Pl.I143b
"Hydrous rehmanni" = Hydrochara noachica8 Heer 1847 water scavenger beetle
Hydrous ovalis n. sp.8 Heer 1862 water scavenger beetle
from Kesselstein
Hydrobius couloni n. sp.8 Heer 1862 water scavenger beetle
from Kesselstein
 Coleoptera - Histeridae
Hister mastodontis n. sp.8 Heer 1862 clown beetle
from Kesselstein
Hister morosus n. sp.8 Heer 1862 clown beetle
from Insektenschicht (lower quarry)
Hister coprolithorum n. sp.8 Heer 1862 clown beetle
numerous specimens from Kesselstein
Hister aemulus n. sp.8 Heer 1862 clown beetle
several from Kesselstein
Hister antiquus n. sp.8 Heer 1862 clown beetle
from Kesselstein
Hister vetustus n. sp.8 Heer 1862 clown beetle
several from Kesselstein
Hister marmoratus n. sp.8 Heer 1862 clown beetle
from Kesselstein
Hister maculigerus n. sp.8 Heer 1862 clown beetle
from Kesselstein
 Coleoptera - Silphidae
Silpha tricostata8 Heer 1847 carrion beetle
from Kesselstein
 Coleoptera - Staphylinidae
Scaphisoma gracile n. sp.8 Heer 1862 rove beetle
2 from Kesselstein, 1 from from Insektenschicht (lower quarry)
Scaphidium deletum8 Heer 1847 rove beetle
Staphylinus atavus n. sp.8 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Lathrobium oeningense n. sp.8 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
? Oxyporus seuberti n. sp.8 Heer 1862 rove beetle
Bledius speciosus n. sp.8 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Oxytelus proaevus n. sp.8 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
Protactus minor n. sp.8 Heer 1862 rove beetle
from Insektenschicht (lower quarry)
 Coleoptera - Nitidulidae
Nitidula ancora n. sp.8 Heer 1862 sap beetle
from Kesselstein
Nitidula maculigera n. sp.8 Heer 1862 sap beetle
Nitidula pallida n. sp.8 Heer 1862 sap beetle
 Coleoptera - Meloidae
Lytta aesculapii5 Heer 1847 blister beetle
Zonites vetusta n. sp.10 Heer 1865 blister beetle
 Coleoptera - Tenebrionidae
Tagenopsis brevicornis n. gen. n. sp.10
Tagenopsis brevicornis n. gen. n. sp.10 Heer 1865 darkling beetle
 Coleoptera - Lymexylidae
Hylecoetus cylindricus n. sp.10 Heer 1865 ship-timber beetle
 Coleoptera - Trogossitidae
Trogosita bella n. sp.8 Heer 1862 bark gnawing beetle
Trogosita amissa n. sp.8 Heer 1862 bark gnawing beetle
from Kesselstein and Insektenschicht (lower quarry)
Trogosita longicollis n. sp.8 Heer 1862 bark gnawing beetle
from Kesselstein
Trogosita koellikeri8 Heer 1847 bark gnawing beetle
Trogosita assimilis n. sp.8 Heer 1862 bark gnawing beetle
from Kesselstein
Trogosita sculpturata n. sp.8 Heer 1862 bark gnawing beetle
from Kesselstein
"Gymnochila obesa n. sp." = Gymnocheilis obesa8 Heer 1862 bark gnawing beetle
from Kesselstein
 Coleoptera - Coccinellidae
"Coccinella spectabilis n. sp." = Coccinella ponomarenkoi10, Coccinella amabilis n. sp.10, "Coccinella decempustulata n. sp." = Coccinella heeri13, Coccinella colorata n. sp.10
"Coccinella spectabilis n. sp." = Coccinella ponomarenkoi10 Ukrainsky 2010 ladybug or ladybird
Coccinella amabilis n. sp.10 Heer 1865 ladybug or ladybird
"Coccinella decempustulata n. sp." = Coccinella heeri13 Ukrainsky 2010 ladybug or ladybird
Written "Coccinella 10-pustulata"
Coccinella colorata n. sp.10 Heer 1865 ladybug or ladybird
 Coleoptera - Cerambycidae
"Prionus polyphemus n. sp." = Ergates polyphemus10, "Prionus spectabilis n. sp." = Ergates spectabilis10, Callidium escheri n. sp.10, Callidium procerum n. sp.10, Clytus pulcher n. sp.10, Lamia antiqua n. sp.13, Acanthoderus lepidus n. sp.10, Acanthoderus sepultus n. sp.10
"Prionus polyphemus n. sp." = Ergates polyphemus10 Heer 1865 longhorned beetle
"Prionus spectabilis n. sp." = Ergates spectabilis10 Heer 1865 longhorned beetle
Callidium escheri n. sp.10 Heer 1865 longhorned beetle
Callidium procerum n. sp.10 Heer 1865 longhorned beetle
Clytus pulcher n. sp.10 Heer 1865 longhorned beetle
Lamia antiqua n. sp.13 Heer 1879 longhorned beetle
Acanthoderus lepidus n. sp.10 Heer 1865 longhorned beetle
Acanthoderus sepultus n. sp.10 Heer 1865 longhorned beetle
 Coleoptera - Chrysomelidae
"Lema vetusta n. sp." = Crioceris vetusta10, Galeruca gemmifera n. sp.13, Galeruca buchi n. sp.10, Gonioctena japeti n. sp.5
"Lema vetusta n. sp." = Crioceris vetusta10 Heer 1865 leaf beetle
Galeruca gemmifera n. sp.13 Heer 1879 leaf beetle
Galeruca buchi n. sp.10 Heer 1865 leaf beetle
Gonioctena japeti n. sp.5 Heer 1847 leaf beetle
 Coleoptera - Curculionidae
Cleonus speciosus n. sp.10 Heer 1865 snout beetle
Brachycerus nanus n. sp.10 Heer 1865 snout beetle
Sitona atavina n. sp.10 Heer 1865 snout beetle
Naupactus crassirostris n. sp.10 Heer 1865 snout beetle
Lixus oeningensis n. sp.10 Heer 1865 snout beetle
 Coleoptera - Attelabidae
"Attelabus durus n. sp." = Phialodes durus10
"Attelabus durus n. sp." = Phialodes durus10 Heer 1865 leaf rolling weevil
 Coleoptera - Brentidae
Miocenocylas heeri n. gen. n. sp.14, Apion antiquum n. sp.10, Antliarhinites gracilis n. gen. n. sp.10, "Rhynchites dionysus n. sp." = Omphalapion dionysus10
Miocenocylas heeri n. gen. n. sp.14 Legalov 2018 weevil
Apion antiquum n. sp.10 Heer 1865 weevil
Antliarhinites gracilis n. gen. n. sp.10 Heer 1865 weevil
"Rhynchites dionysus n. sp." = Omphalapion dionysus10 Heer 1865 weevil
 Coleoptera - Glaphyridae
Glaphyrus antiquus n. sp.8 Heer 1862 bumble bee scarab beetle
from Insektenschicht (lower quarry)
 Coleoptera -
Gymnopleurus deperditus n. sp.8, "Onthophagus urus" = Onthophagus urusheeri8
Gymnopleurus deperditus n. sp.8 Heer 1862 scarab beetle
from Kesselstein
"Onthophagus urus" = Onthophagus urusheeri8 Krell 2000 scarab beetle
"several" specimens
 Coleoptera - Hybosoridae
Hybosorus lividus n. sp.8 Heer 1862 scavenger scarab beetle
 Coleoptera - Scarabaeidae
Aphodius brevipennis n. sp.8, Onthophagus prodromus n. sp.8, Onthophagus bisontinus n. sp.8, Onthophagus crassus n. sp.8, Gymnopleurus sisyphus n. sp.5, Gymnopleurus rotundatus n. sp.8, Copris druidum n. sp.8, Lepitrix germanica n. sp.8, Serica minutula n. sp.8, Oniticellus amplicollis n. sp.8, Anomala fugax n. sp.8, "Scarabaeus proserpinae n. sp." = Pentodon proserpinae8, ? Trichius rotundatus n. sp.8, Trichius unifasciatus n. sp.8, Trichius lugubris n. sp.8, "Trichius aedilis n. sp." = Gnorimus aedilis8, Valgus oeningensis n. sp.8
Aphodius brevipennis n. sp.8 Heer 1862 scarab beetle
from Kesselstein
Onthophagus prodromus n. sp.8 Heer 1862 scarab beetle
several from Kesselstein
Onthophagus bisontinus n. sp.8 Heer 1862 scarab beetle
from Insektenschicht (lower quarry)
Onthophagus crassus n. sp.8 Heer 1862 scarab beetle
from Kesselstein
Gymnopleurus sisyphus n. sp.5 Heer 1847 scarab beetle
Gymnopleurus rotundatus n. sp.8 Heer 1862 scarab beetle
from Kesselstein
Copris druidum n. sp.8 Heer 1862 scarab beetle
from Kesselstein
Lepitrix germanica n. sp.8 Heer 1862 scarab beetle
Serica minutula n. sp.8 Heer 1862 scarab beetle
from Kesselstein
Oniticellus amplicollis n. sp.8 Heer 1862 scarab beetle
from Kesselstein
Anomala fugax n. sp.8 Heer 1862 scarab beetle
from Kesselstein
"Scarabaeus proserpinae n. sp." = Pentodon proserpinae8 Heer 1862 rhinoceras beetle
from Kesselstien
? Trichius rotundatus n. sp.8 Heer 1862 flower chafer
from Kesselstein
Trichius unifasciatus n. sp.8 Heer 1862 flower chafer
from Kesselstein
Trichius lugubris n. sp.8 Heer 1862 flower chafer
"Trichius aedilis n. sp." = Gnorimus aedilis8 Heer 1862 flower chafer
from Kesselstein
Valgus oeningensis n. sp.8 Heer 1862 flower chafer
from Kesselstein
 Coleoptera - Geotrupidae
Geotrupes germari n. sp.8 Heer 1862 earth-boring dung beetle
from Kesselstein
 Coleoptera -
"Copris subterranea n. sp." = Copris subterraneus8
"Copris subterranea n. sp." = Copris subterraneus8 Heer 1862 scarab beetle
from Kesselstein
 Coleoptera - Dytiscidae
Dytiscus lavateri8 Heer 1847 predaceous diving beetle
from Kesselstein
"Hydaticus areolatus n. sp." = Graphoderes areolatus8 Heer 1862 predaceous diving beetle
Cybister nicoleti8 Heer 1862 predaceous diving beetle
from Kesselstein
Cybister agassizi n. sp.8 Heer 1862 predaceous diving beetle
from Kesselstein
Cybister atavus n. sp.8 Heer 1862 predaceous diving beetle
from Kesselstein
Colymbetes aemulus n. sp.8 Heer 1862 predaceous diving beetle
from Kesselstein
Hydroporus benzeli n. sp.8 Heer 1862 predaceous diving beetle
 Coleoptera - Carabidae
"Calosoma deplanatum n. sp." = Campalita deplanatum7 Heer 1860 ground beetle
Calosoma nauckianum n. sp.7 Heer 1860 ground beetle
Calosoma escrobiculatum n. sp.7 Heer 1860 ground beetle
Calosoma catenulatum n. sp.7 Heer 1860 ground beetle
Kesselstein
Calosoma escheri n. sp.7 Heer 1860 ground beetle
Harpalus constrictus n. sp.8 Heer 1862 ground beetle
from Kesselstein
Harpalus tardigradus n. sp.8 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Harpalus stygius n. sp.8 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Harpalus stierlini n. sp.8 Heer 1862 ground beetle
Dichirotrichus lividus8 Heer 1862 ground beetle
from Kesselstein
Amara primigenia n. sp.8 Heer 1862 ground beetle
from Kesselstein
Amara princeps n. sp.8 Heer 1862 ground beetle
1 from Insektenschicht (lower quarry), 1 from Kesselstein
Amara pinguicula n. sp.8 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Sinis brevicollis n. gen. n. sp.8 Heer 1862 ground beetle
1 male, 1 female from lower quarry
Pterostichus minutulus n. sp.8 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Pterostichus vetustus n. sp.8 Heer 1862 ground beetle
from Insektenschicht (lower quarry)
Badister grandis n. sp.8 Heer 1862 ground beetle
from Kesselstein
Badister macrocephalus n. sp.8 Heer 1862 ground beetle
"multiple examples" in Kesselstein, including var. major
Badister fragilis n. sp.8 Heer 1862 ground beetle
from Kesselstein (2 var. paulo major)
Nebria pluto n. sp.8 Heer 1862 ground beetle
1 from lower quarry, 2 from Kesselstein
Carabites laesicollis n. sp.8 Heer 1862 ground beetle
Trechinites oblongus n. sp.8 Heer 1862 ground beetle
from lower quarry
Trechinites clairvillii n. sp.8 Heer 1862 ground beetle
from lower quarry
Glenopterus laevigatus n. gen. n. sp.5 Heer 1847 ground beetle
 Coleoptera - Gyrinidae
Miodineutes oeningensis n. gen. n. sp.4, Miodineutes heeri n. sp.4, "Dineutus insignis n. sp." = Miodineutes insignis8, "Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris8
Miodineutes oeningensis n. gen. n. sp.4 Hatch 1927 whirligig beetle
Dineutus insignis specimen in pl 2, fig 28 of Heer
Miodineutes heeri n. sp.4 Hatch 1927 whirligig beetle
Dineutus longiventris Specimen in pl 2, fig 25 of Heer
"Dineutus insignis n. sp." = Miodineutes insignis8 Heer 1862 whirligig beetle
from Kesselstein
"Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris8 Heer 1862 whirligig beetle
from Kesselstein
 Coleoptera -
Helophorus magnus n. sp.8 Heer 1862 beetle
from Kesselstein
Hydrobius godeti n. sp.8 Heer 1862 beetle
from Libellenschicht (upper quarry)
Hydrophilus stenopterus n. sp.8 Heer 1862 beetle
from Kesselstein
Cistelites spectabilis n. sp.12 Heer 1869 beetle
 Diptera - Chironomidae
Chironomus gaudini n. sp.10 Heer 1865 non biting midge
 Diptera - Syrphidae
Syrphus bremii n. sp.10 Heer 1865 hover fly
Syrphus schellenbergi n. sp.10 Heer 1865 hover fly
 Diptera - Oestridae
"Dipterites obovatus n. sp." = Adipterites obovatus10
"Dipterites obovatus n. sp." = Adipterites obovatus10 Heer 1865 bot fly
 Diptera - Tabanidae
"? Hexatoma oeningensis n. sp." = Heptatoma oeningensis10
"? Hexatoma oeningensis n. sp." = Heptatoma oeningensis10 Heer 1865 horse fly
 Diptera - Bibionidae
Bibio elongatus15 Heer 1849 march fly
ETHZ S.N. 189 Pl. I 863 (labeled by Heer)
"Protomyia speciosa n. sp." = Penthetria amoena10 Heer 1849 march fly
ETHZ Pl. I 873; also Pl. I 872, 876 as P. amoena in Skartveit & Pika, 2014
Penthetria brevicollis15 Heer 1849 march fly
ETHZ Pl. I 870
Plecia hilaris15 Heer 1849 love bug
ETHZ Pl. I 850
"Protomyia jucunda" = Plecia jucunda Heer 1849 love bug
ETHZ Pl. I 871, 853(x2), 856, 861, 867, 869 (some labeled by Brongniart fide Skartveit & Pika 2014)
Plecia macilenta n. sp.15 Skartveit and Pika 2014 love bug
ETHZ Pl. I 853, 853 female
Bibio pulchellus15 Heer 1849 march fly
ETHZ Pl. I 854
Bibio angustatus15 Heer 1849 march fly
ETHZ Pl. I 851
Bibio fusiformis15 Heer 1849 march fly
ETHZ Pl. I 858 (x2), 861 (x6)
Bibio moestus15 Heer 1849 march fly
ETHZ Pl. I 855, 857(x2)
Bibio crassinervis n. sp.15 Skartveit and Pika 2014 march fly
ETHZ Pl. I 851, Pl.I 851(2), Pl. I 862, Pl. I 875, Pl. I 888
Bibio brevis15 Heer 1849 march fly
Some in lot Pl. I 856; ETHZ Pl.I 865, Pl. I 887
Dilophus pumilio n. sp.15 Skartveit and Pika 2014 march fly
ETHZ Pl. I 886
 Diptera - Mycetophilidae
Mycetophila orci n. sp.10 Heer 1865 fungus gnat
Mycetophila pusillima n. sp.10 Heer 1865 fungus gnat
 Diptera - Cecidomyiidae
Cecidomyia bremii n. sp.10 Heer 1865 gall midge
 Diptera - Sciaridae
Sciara deleta n. sp.10
Sciara deleta n. sp.10 Heer 1865 dark-winged fungus gnat
 Lepidoptera - Bombycidae
"Bombycites" oeningensis n. sp. Heer 1849 moth
pupa
Bombycites buchii n. sp.10 Heer 1865 moth
 Ephemeroptera -
Ephemera oeningensis n. sp.10 Heer 1865 mayfly
 Odonata - Sieblosiidae
"Agrion iris n. sp." = Stenolestes iris13
"Agrion iris n. sp." = Stenolestes iris13 Heer 1879 damselfly
 Odonata -
"Agrion (Sterope) parthenope n. sp." = Steropoides parthenope, "Libellula" thetis
"Agrion (Sterope) parthenope n. sp." = Steropoides parthenope Heer 1849 damselfly
"Libellula" thetis Heer 1849 true dragonfly
larva
Malacostraca
 Isopoda - Armadillidiidae
"Armadillo molassicus n. sp." = Armadillidium molassicum10
"Armadillo molassicus n. sp." = Armadillidium molassicum10 Heer 1865 isopod
Arachnida
 Araneae -
Schellenbergia rotundata n. gen. n. sp.10
Schellenbergia rotundata n. gen. n. sp.10 Heer 1865 spider
 Araneae - Gnaphosidae
"Macaria tenella n. sp." = Micaria tenella10
"Macaria tenella n. sp." = Micaria tenella10 Heer 1865 spider
 Araneae - Dictynidae
? Argyroneta longipes n. sp.10 Heer 1865 spider
 Araneae - Thomisidae
"Thomisus oeningensis n. sp." = Xysticus oeningensis10, Thomisus lividus n. sp.10, Thomisus sulzeri n. sp.10
"Thomisus oeningensis n. sp." = Xysticus oeningensis10 Heer 1865 crab spider
Thomisus lividus n. sp.10 Heer 1865 crab spider
Thomisus sulzeri n. sp.10 Heer 1865 crab spider
 Araneae - Araneidae
"Epeira molassica n. sp." = Araneus molassicus10
"Epeira molassica n. sp." = Araneus molassicus10 Heer 1865 orb-weaving spider
 Araneae - Theridiidae
Theridion globulus n. sp.10 Heer 1865 cobweb spider
Theridion annulipes n. sp.10 Heer 1865 cobweb spider
Theridion maculipes n. sp.10 Heer 1865 cobweb spider
 Araneae -
Clubiona eseri n. sp.10
Clubiona eseri n. sp.10 Heer 1865 spider