PBDB

Oeningen (Zurich collection) (Miocene of Germany)

Where: Baden-Württemberg, Germany (47.7° N, 8.9° E: paleocoordinates 47.9° N, 7.8° E)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: MN 7 mammal zone, Upper Oehningen beds Member (Upper Freshwater-Molasse Formation), Sarmatian (12.7 - 11.6 Ma)

• group of beds-level stratigraphic resolution

Environment/lithology: lacustrine - large; lithified limestone

• "Rising into massive mountains, as in the well-known Righi and Rossberg, they attain a thickness of more than 6000 feet. While they include proofs of the presenceof the sea, they have preserved with marvellous perfection a large number of the plants which clothed the Alps, and of the insects which flitted through the woodlands. They form part of a great series of deposits which have been termed “Molasse” by the Swiss geologists." Geikie, 1882

•"...deposited in the Northern Alpine Foreland basin..." Uhl et al., 2006

Size classes: macrofossils, mesofossils

Preservation: mold/impression, adpression

Collection methods: Collection of Zurich University

Primary reference: O. Heer. 1849. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien. Zweiter Theil: Henschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen 1-264 [M. Clapham/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 117252: authorized by Matthew Clapham, entered by Matthew Clapham on 18.09.2011, edited by Jered Karr and Amy Delelli

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae -	Schellenbergia rotundata n. gen. n. sp.⁹, Clubiona eseri n. sp.⁹ Schellenbergia rotundata n. gen. n. sp.⁹ Heer 1864 spider Clubiona eseri n. sp.⁹ Heer 1864 spider
Araneae - Thomisidae	Thomisus sulzeri n. sp.⁹, Thomisus lividus n. sp.⁹, "Thomisus oeningensis n. sp." = Xysticus oeningensis⁹ Thomisus sulzeri n. sp.⁹ Heer 1864 crab spider Thomisus lividus n. sp.⁹ Heer 1864 crab spider "Thomisus oeningensis n. sp." = Xysticus oeningensis⁹ Heer 1864 crab spider
Araneae - Gnaphosidae	"Macaria tenella n. sp." = Micaria tenella⁹ "Macaria tenella n. sp." = Micaria tenella⁹ Heer 1864 spider
Araneae - Araneidae	"Epeira molassica n. sp." = Araneus molassicus⁹ "Epeira molassica n. sp." = Araneus molassicus⁹ Heer 1864 orb-weaving spider
Araneae - Theridiidae	Theridion annulipes n. sp.⁹, Theridion maculipes n. sp.⁹, Theridion globulus n. sp.⁹ Theridion annulipes n. sp.⁹ Heer 1864 cobweb spider Theridion maculipes n. sp.⁹ Heer 1864 cobweb spider Theridion globulus n. sp.⁹ Heer 1864 cobweb spider
Araneae - Dictynidae	? Argyroneta longipes n. sp.⁹ ? Argyroneta longipes n. sp.⁹ Heer 1864 spider
Malacostraca
Isopoda - Armadillidiidae	"Armadillo molassicus n. sp." = Armadillidium molassicum⁹ "Armadillo molassicus n. sp." = Armadillidium molassicum⁹ Heer 1864 isopod
Insecta
Odonata -	"Libellula" thetis "Libellula" thetis Heer 1849 true dragonfly larva
Odonata - Sieblosiidae	"Agrion iris n. sp." = Stenolestes iris¹³ "Agrion iris n. sp." = Stenolestes iris¹³ Heer 1879 damselfly
Odonata -	"Agrion (Sterope) parthenope n. sp." = Steropoides parthenope "Agrion (Sterope) parthenope n. sp." = Steropoides parthenope Heer 1849 damselfly
Ephemeroptera -	Ephemera oeningensis n. sp.⁹ Ephemera oeningensis n. sp.⁹ Heer 1864 mayfly
Orthoptera - Pyrgomorphidae	"Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri⁹ "Oedipoda fischeri n. sp." = Miopyrgomorpha fischeri⁹ Heer 1864 gaudy grasshopper
Orthoptera - Acrididae	Oedipoda germari n. sp.⁹ Oedipoda germari n. sp.⁹ Heer 1864 short-horned grasshopper
Orthoptera - Tetrigidae	"Tetrix gracilis n. sp." = Tettigidea gracilis⁹ "Tetrix gracilis n. sp." = Tettigidea gracilis⁹ Heer 1864 pygmy grasshopper
Orthoptera - Tettigoniidae	"Decticus speciosus n. sp." = Platycleis speciosa⁹ "Decticus speciosus n. sp." = Platycleis speciosa⁹ Heer 1864 katydid
Orthoptera - Gryllotalpidae	Gryllotalpa stricta n. sp.⁹ Gryllotalpa stricta n. sp.⁹ Heer 1864 mole cricket
Orthoptera - Gryllidae	Gryllus troglodytes n. sp.⁹ Gryllus troglodytes n. sp.⁹ Heer 1864 true crickets
Blattodea - Rhinotermitidae	"Termes hartungi n. sp." = Reticulitermes hartungi⁹ "Termes hartungi n. sp." = Reticulitermes hartungi⁹ Heer 1864 termite
Blattodea - Termitidae	Termes buchii n. sp.⁹ Termes buchii n. sp.⁹ Heer 1864 termite
Blattodea - Blattidae	"Blattidium coloratum n. sp." = Blatta colorata¹⁰ "Blattidium coloratum n. sp." = Blatta colorata¹⁰ Heer 1864 cockroach from Kesselstein
Dermaptera - Forficulidae	Forficula minuta n. sp.⁹, Forficula recta n. sp.⁹ Forficula minuta n. sp.⁹ Heer 1864 common earwig Forficula recta n. sp.⁹ Heer 1864 common earwig
Hemiptera - Cercopidae	Cercopis germari n. sp.⁹, Cercopis herrichi n. sp.⁹, Cercopis hageni n. sp.⁹ Cercopis germari n. sp.⁹ Heer 1864 froghopper Cercopis herrichi n. sp.⁹ Heer 1864 froghopper Cercopis hageni n. sp.⁹ Heer 1864 froghopper
Hemiptera - Gerridae	Limnobates prodromus n. sp.⁹ Limnobates prodromus n. sp.⁹ Heer 1864 water strider
Hemiptera - Reduviidae	Harpactor maculipes n. sp.⁶, Harpactor obsoletus n. sp.⁶ Harpactor maculipes n. sp.⁶ Heer 1853 assassin bug lower quarry Harpactor obsoletus n. sp.⁶ Heer 1853 assassin bug from the Kesselstein
Hemiptera - Tingidae	"Monanthia wollastoni n. sp." = Dictyla wollastoni⁹ "Monanthia wollastoni n. sp." = Dictyla wollastoni⁹ Heer 1864 lace bug
Hemiptera - Nabidae	Prostemma oeningensis n. sp.⁶ Prostemma oeningensis n. sp.⁶ Heer 1853 true bug lower quarry
Hemiptera - Belostomatidae	"Belostoma speciosum n. sp." = Belostomates speciosum⁹ "Belostoma speciosum n. sp." = Belostomates speciosum⁹ Heer 1864 giant water bug
Hemiptera - Corixidae	"Corisa fasciolata" = Corixa fasciolata⁶ "Corisa fasciolata" = Corixa fasciolata⁶ Heer 1853 water boatman from the Libellen-schicht
Hemiptera - Alydidae	Alydus pulchellus n. sp.⁶, "Nabis gracillima n. sp." = Licremytra gracillima⁹ Alydus pulchellus n. sp.⁶ Heer 1853 broad-headed bug lower quarry "Nabis gracillima n. sp." = Licremytra gracillima⁹ Heer 1864 broad-headed bug
Hemiptera - Pentatomidae	Pentatoma pictum n. sp.⁹, Halys bruckmanni⁶, Halys spectabilis n. sp.⁹, Eurydema impudica⁶, Eurydema arcuata n. sp.⁶ Pentatoma pictum n. sp.⁹ Heer 1864 stink bug Halys bruckmanni⁶ Heer 1853 stink bug from the Kesselstein Halys spectabilis n. sp.⁹ Heer 1864 stink bug Eurydema impudica⁶ Heer 1853 stink bug Eurydema arcuata n. sp.⁶ Heer 1853 stink bug from the Kesselstein
Hemiptera - Scutelleridae	Pachycoris escheri⁶, Pachycoris burmeisteri n. sp.⁹, Pachycoris protogaeus n. sp.⁶ Pachycoris escheri⁶ Heer 1853 metallic shield bug Pachycoris burmeisteri n. sp.⁹ Heer 1864 metallic shield bug Pachycoris protogaeus n. sp.⁶ Heer 1853 metallic shield bug lower quarry
Hemiptera - Cydnidae	Cydnus oeningensis⁶, Cydnopsis deleta n. sp.⁶, Cydnopsis exilis n. sp.⁶, Cydnopsis pygmaea n. sp.⁶ Cydnus oeningensis⁶ Heer 1853 burrowing bug lower quarry Cydnopsis deleta n. sp.⁶ Heer 1853 burrowing bug lower quarry Cydnopsis exilis n. sp.⁶ Heer 1853 burrowing bug Cydnopsis pygmaea n. sp.⁶ Heer 1853 burrowing bug upper quarry
Hemiptera - Heterogastridae	Heterogaster tristis n. sp.⁹ Heterogaster tristis n. sp.⁹ Heer 1864 true bug
Hemiptera - Rhyparochromidae	Pachymerus obsoletus n. sp.⁶ Pachymerus obsoletus n. sp.⁶ Heer 1853 seed bug lower quarry
Hemiptera - Lygaeidae	Cephalocoris pilosus n. gen. n. sp.⁶ Cephalocoris pilosus n. gen. n. sp.⁶ Heer 1853 milkweed bug lower quarry
Thysanoptera -	Thrips oeningensis n. sp.⁹, Thrips annosa n. sp.⁹ Thrips oeningensis n. sp.⁹ Heer 1864 thrips Thrips annosa n. sp.⁹ Heer 1864 thrips
Hymenoptera - Ichneumonidae	Ichneumonites bellus n. sp.¹¹, Ichneumon infernalis n. sp.⁹ Ichneumonites bellus n. sp.¹¹ Heer 1867 ichneumon wasp Ichneumon infernalis n. sp.⁹ Heer 1864 ichneumon wasp
Hymenoptera -	Pteromalites oeningensis n. gen. n. sp.⁹ Pteromalites oeningensis n. gen. n. sp.⁹ Heer 1864 chalcid wasp
Hymenoptera - Scoliidae	Scolia saussureana n. sp.⁹ Scolia saussureana n. sp.⁹ Heer 1864 wasp
Hymenoptera - Sphecidae	Ammophila inferna n. sp.⁹, Ammophila annosa n. sp.⁹ Ammophila inferna n. sp.⁹ Heer 1864 digger wasp Ammophila annosa n. sp.⁹ Heer 1864 digger wasp
Hymenoptera - Megachilidae	Megachilidae indet.¹⁶, Osmia nigra n. sp.¹⁶, "Apis adamitica n. sp." = Lithurge adamitica⁹ Megachilidae indet.¹⁶ Latreille 1802 mason bee ETH Pl. I, 687; Pl. I, 688(689); Pl. I, 718 Osmia nigra n. sp.¹⁶ Zeuner and Manning 1976 mason bee ETH Pl. I, 719; Pl. I, 685 "Apis adamitica n. sp." = Lithurge adamitica⁹ Heer 1864 mason bee ETH V.S. 4 (Heer's holotype); SN.200 (apparently different from Heer specimens) in Zeuner & Manning, 1976
Hymenoptera - Apidae	Bombus grandaevus¹¹, Anthophorites veterana¹⁶, Anthophorites longaeva¹⁶, Anthophorites mellona n. gen. n. sp., Bombus abavus n. sp.¹⁶, "Bombus jurinei n. sp." = Xylocopa jurinei¹¹, "Bombus atavus n. sp." = Xylocopa atavus¹¹, Xylocopa senilis¹⁶ Bombus grandaevus¹¹ Heer 1849 bee Anthophorites veterana¹⁶ Heer 1849 bee ETH Pl. I, 684 Anthophorites longaeva¹⁶ Heer 1867 bee ETH Pl. I, 683, 712, 809 (not Heer's types) Anthophorites mellona n. gen. n. sp. Heer 1849 bee Bombus abavus n. sp.¹⁶ Heer 1867 bumble bee Holotype (lost), ETH Pl. I, 808 (examined by Cockerell), SN.201 "Bombus jurinei n. sp." = Xylocopa jurinei¹¹ Heer 1864 large carpenter bee ETH Pl. I, 682a (type in Heer, 1865), 682b (specimen in heer, 1867) "Bombus atavus n. sp." = Xylocopa atavus¹¹ Heer 1867 large carpenter bee Xylocopa senilis¹⁶ Heer 1849 large carpenter bee ETH 21a(b), doutbfully 21c
Hymenoptera -	Andrena primaeva n. sp.¹ Andrena primaeva n. sp.¹ Cockerell 1909 bee SN.202
Hymenoptera - Formicidae	Imhoffia pallida n. sp.¹¹, "Formica lignitum" = Camponotus lignitus, "Formica heraclea" = Camponotus heracleus, Formica orbata, "Ponera globosa n. sp." = Poneropsis globosa, "Ponera veneraria n. sp." = Poneropsis veneraria⁹, Poneropsis stygia n. sp.¹¹ Imhoffia pallida n. sp.¹¹ Heer 1864 ant "Formica lignitum" = Camponotus lignitus Germar 1837 ant "Formica heraclea" = Camponotus heracleus Heer 1849 ant Formica orbata Heer 1849 ant "Ponera globosa n. sp." = Poneropsis globosa Heer 1849 ant "Ponera veneraria n. sp." = Poneropsis veneraria⁹ Heer 1864 ant Poneropsis stygia n. sp.¹¹ Heer 1867 ant
Hymenoptera - Vespidae	Polistes kirbyanus n. sp.², Polistes primitiva n. sp.⁹ Polistes kirbyanus n. sp.² Cockerell 1915 wasp ETH Pl. I, 681; Osmia kirbyana in Heer's label (unpublished) Polistes primitiva n. sp.⁹ Heer 1864 wasp
Coleoptera -	Hydrophilus stenopterus n. sp.⁸, Helophorus magnus n. sp.⁸, Cistelites spectabilis n. sp.¹², Hydrobius godeti n. sp.⁸ Hydrophilus stenopterus n. sp.⁸ Heer 1862 beetle from Kesselstein Helophorus magnus n. sp.⁸ Heer 1862 beetle from Kesselstein Cistelites spectabilis n. sp.¹² Heer 1869 beetle Hydrobius godeti n. sp.⁸ Heer 1862 beetle from Libellenschicht (upper quarry)
Coleoptera - Meloidae	Zonites vetusta n. sp.⁹, Lytta aesculapii⁵ Zonites vetusta n. sp.⁹ Heer 1864 blister beetle Lytta aesculapii⁵ Heer 1847 blister beetle
Coleoptera - Tenebrionidae	Tagenopsis brevicornis n. gen. n. sp.⁹ Tagenopsis brevicornis n. gen. n. sp.⁹ Heer 1864 darkling beetle
Coleoptera - Chrysomelidae	Gonioctena japeti n. sp.⁵, "Lema vetusta n. sp." = Crioceris vetusta⁹, Galeruca gemmifera n. sp.¹³, Galeruca buchi n. sp.⁹ Gonioctena japeti n. sp.⁵ Heer 1847 leaf beetle "Lema vetusta n. sp." = Crioceris vetusta⁹ Heer 1864 leaf beetle Galeruca gemmifera n. sp.¹³ Heer 1879 leaf beetle Galeruca buchi n. sp.⁹ Heer 1864 leaf beetle
Coleoptera - Cerambycidae	"Prionus spectabilis n. sp." = Ergates spectabilis⁹, "Prionus polyphemus n. sp." = Ergates polyphemus⁹, Callidium procerum n. sp.⁹, Callidium escheri n. sp.⁹, Clytus pulcher n. sp.⁹, Lamia antiqua n. sp.¹³, Acanthoderus lepidus n. sp.⁹, Acanthoderus sepultus n. sp.⁹ "Prionus spectabilis n. sp." = Ergates spectabilis⁹ Heer 1864 longhorned beetle "Prionus polyphemus n. sp." = Ergates polyphemus⁹ Heer 1864 longhorned beetle Callidium procerum n. sp.⁹ Heer 1864 longhorned beetle Callidium escheri n. sp.⁹ Heer 1864 longhorned beetle Clytus pulcher n. sp.⁹ Heer 1864 longhorned beetle Lamia antiqua n. sp.¹³ Heer 1879 longhorned beetle Acanthoderus lepidus n. sp.⁹ Heer 1864 longhorned beetle Acanthoderus sepultus n. sp.⁹ Heer 1864 longhorned beetle
Coleoptera - Brentidae	Apion antiquum n. sp.⁹, Antliarhinites gracilis n. gen. n. sp.⁹, "Rhynchites dionysus n. sp." = Omphalapion dionysus⁹, Miocenocylas heeri n. gen. n. sp.¹⁴ Apion antiquum n. sp.⁹ Heer 1864 weevil Antliarhinites gracilis n. gen. n. sp.⁹ Heer 1864 weevil "Rhynchites dionysus n. sp." = Omphalapion dionysus⁹ Heer 1864 weevil Miocenocylas heeri n. gen. n. sp.¹⁴ Legalov 2018 weevil
Coleoptera - Curculionidae	Naupactus crassirostris n. sp.⁹, Sitona atavina n. sp.⁹, Cleonus speciosus n. sp.⁹, Lixus oeningensis n. sp.⁹, Brachycerus nanus n. sp.⁹ Naupactus crassirostris n. sp.⁹ Heer 1864 snout beetle Sitona atavina n. sp.⁹ Heer 1864 snout beetle Cleonus speciosus n. sp.⁹ Heer 1864 snout beetle Lixus oeningensis n. sp.⁹ Heer 1864 snout beetle Brachycerus nanus n. sp.⁹ Heer 1864 snout beetle
Coleoptera - Attelabidae	"Attelabus durus n. sp." = Phialodes durus⁹ "Attelabus durus n. sp." = Phialodes durus⁹ Heer 1864 leaf rolling weevil
Coleoptera - Nitidulidae	Nitidula maculigera n. sp.⁸, Nitidula ancora n. sp.⁸, Nitidula pallida n. sp.⁸ Nitidula maculigera n. sp.⁸ Heer 1862 sap beetle Nitidula ancora n. sp.⁸ Heer 1862 sap beetle from Kesselstein Nitidula pallida n. sp.⁸ Heer 1862 sap beetle
Coleoptera - Trogossitidae	Trogosita amissa n. sp.⁸, Trogosita longicollis n. sp.⁸, Trogosita koellikeri⁸, Trogosita bella n. sp.⁸, Trogosita assimilis n. sp.⁸, Trogosita sculpturata n. sp.⁸, "Gymnochila obesa n. sp." = Gymnocheilis obesa⁸ Trogosita amissa n. sp.⁸ Heer 1862 bark gnawing beetle from Kesselstein and Insektenschicht (lower quarry) Trogosita longicollis n. sp.⁸ Heer 1862 bark gnawing beetle from Kesselstein Trogosita koellikeri⁸ Heer 1847 bark gnawing beetle Trogosita bella n. sp.⁸ Heer 1862 bark gnawing beetle Trogosita assimilis n. sp.⁸ Heer 1862 bark gnawing beetle from Kesselstein Trogosita sculpturata n. sp.⁸ Heer 1862 bark gnawing beetle from Kesselstein "Gymnochila obesa n. sp." = Gymnocheilis obesa⁸ Heer 1862 bark gnawing beetle from Kesselstein
Coleoptera - Lymexylidae	Hylecoetus cylindricus n. sp.⁹ Hylecoetus cylindricus n. sp.⁹ Heer 1864 ship-timber beetle
Coleoptera - Coccinellidae	Coccinella amabilis n. sp.⁹, Coccinella colorata n. sp.⁹, "Coccinella decempustulata n. sp." = Coccinella heeri¹³, "Coccinella spectabilis n. sp." = Coccinella ponomarenkoi⁹ Coccinella amabilis n. sp.⁹ Heer 1864 ladybird beetle Coccinella colorata n. sp.⁹ Heer 1864 ladybird beetle "Coccinella decempustulata n. sp." = Coccinella heeri¹³ Ukrainsky 2010 ladybird beetle Written "Coccinella 10-pustulata" "Coccinella spectabilis n. sp." = Coccinella ponomarenkoi⁹ Ukrainsky 2010 ladybird beetle
Coleoptera - Buprestidae	Capnodis puncticollis⁸, Capnodis spectabilis n. sp.⁸, Capnodis antiqua⁸, Chalcophora laevigata⁸, Chalcophora pulchella n. sp.⁸, Acmaeodera antholitha n. sp.⁸, Acmaeodera brevicollis n. sp.⁸, Anthaxia pallida n. sp.⁸, ? Anthaxia doris n. sp.⁸, Protogenia escheri n. gen. n. sp.⁵, "Ancylocheira alemanica n. sp." = Buprestis alemanica¹³, "Ancylochira tincta n. sp." = Buprestis tincta⁸, "Ancylocheira concinna n. sp." = Buprestis concinna¹³, Buprestites agriloides n. sp.⁸ Capnodis puncticollis⁸ Heer 1847 jewel beetle Capnodis spectabilis n. sp.⁸ Heer 1862 jewel beetle from Kesselstein Capnodis antiqua⁸ Heer 1847 jewel beetle "several" specimens Chalcophora laevigata⁸ Heer 1862 jewel beetle Chalcophora pulchella n. sp.⁸ Heer 1862 jewel beetle from Kesselstein Acmaeodera antholitha n. sp.⁸ Heer 1862 jewel beetle from Kesselstein Acmaeodera brevicollis n. sp.⁸ Heer 1862 jewel beetle from Kesselstein Anthaxia pallida n. sp.⁸ Heer 1862 jewel beetle from Kesselstein ? Anthaxia doris n. sp.⁸ Heer 1862 jewel beetle from Kesselstein Protogenia escheri n. gen. n. sp.⁵ Heer 1847 jewel beetle "Ancylocheira alemanica n. sp." = Buprestis alemanica¹³ Heer 1879 jewel beetle "Ancylochira tincta n. sp." = Buprestis tincta⁸ Heer 1862 jewel beetle from Kesselstein,includes var. b. elytris majoribus "Ancylocheira concinna n. sp." = Buprestis concinna¹³ Heer 1879 jewel beetle Buprestites agriloides n. sp.⁸ Heer 1862 jewel beetle
Coleoptera - Cantharidae	"Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta⁹ "Telephorus macilentus n. sp." = Cantharis (Cantharis) macilenta⁹ Heer 1864 soldier beetle
Coleoptera - Lampyridae	Lampyris orciluca n. sp.⁹ Lampyris orciluca n. sp.⁹ Heer 1864 beetle
Coleoptera - Elateridae	"Elater (Alaus) spectabilis n. sp." = Alaus spectabilis⁹ "Elater (Alaus) spectabilis n. sp." = Alaus spectabilis⁹ Heer 1864 click beetle
Coleoptera - Byrrhidae	Byrrhus oeningensis⁸ Byrrhus oeningensis⁸ Heer 1847 pill beetle several from Kesselstein
Coleoptera - Staphylinidae	Protactus minor n. sp.⁸, ? Oxyporus seuberti n. sp.⁸, Oxytelus proaevus n. sp.⁸, Bledius speciosus n. sp.⁸, Silpha tricostata⁸, Lathrobium oeningense n. sp.⁸, Staphylinus atavus n. sp.⁸, Scaphisoma gracile n. sp.⁸, Scaphidium deletum⁸ Protactus minor n. sp.⁸ Heer 1862 rove beetle from Insektenschicht (lower quarry) ? Oxyporus seuberti n. sp.⁸ Heer 1862 rove beetle Oxytelus proaevus n. sp.⁸ Heer 1862 rove beetle from Insektenschicht (lower quarry) Bledius speciosus n. sp.⁸ Heer 1862 rove beetle from Insektenschicht (lower quarry) Silpha tricostata⁸ Heer 1847 rove beetle from Kesselstein Lathrobium oeningense n. sp.⁸ Heer 1862 rove beetle from Insektenschicht (lower quarry) Staphylinus atavus n. sp.⁸ Heer 1862 rove beetle from Insektenschicht (lower quarry) Scaphisoma gracile n. sp.⁸ Heer 1862 rove beetle 2 from Kesselstein, 1 from from Insektenschicht (lower quarry) Scaphidium deletum⁸ Heer 1847 rove beetle
Coleoptera -	Gymnopleurus deperditus n. sp.⁸ Gymnopleurus deperditus n. sp.⁸ Heer 1862 scarab beetle from Kesselstein
Coleoptera - Glaphyridae	Glaphyrus antiquus n. sp.⁸ Glaphyrus antiquus n. sp.⁸ Heer 1862 bumble bee scarab beetle from Insektenschicht (lower quarry)
Coleoptera -	"Onthophagus urus" = Onthophagus urusheeri⁸, "Copris subterranea n. sp." = Copris subterraneus⁸ "Onthophagus urus" = Onthophagus urusheeri⁸ Krell 2000 scarab beetle "several" specimens "Copris subterranea n. sp." = Copris subterraneus⁸ Heer 1862 scarab beetle from Kesselstein
Coleoptera - Geotrupidae	Geotrupes germari n. sp.⁸ Geotrupes germari n. sp.⁸ Heer 1862 earth-boring dung beetle from Kesselstein
Coleoptera - Hybosoridae	Hybosorus lividus n. sp.⁸ Hybosorus lividus n. sp.⁸ Heer 1862 scavenger scarab beetle
Coleoptera - Scarabaeidae	Aphodius brevipennis n. sp.⁸, "Scarabaeus proserpinae n. sp." = Pentodon proserpinae⁸, Anomala fugax n. sp.⁸, Serica minutula n. sp.⁸, Lepitrix germanica n. sp.⁸, Oniticellus amplicollis n. sp.⁸, Onthophagus prodromus n. sp.⁸, Onthophagus bisontinus n. sp.⁸, Onthophagus crassus n. sp.⁸, Gymnopleurus rotundatus n. sp.⁸, Gymnopleurus sisyphus n. sp.⁵, Copris druidum n. sp.⁸, Trichius lugubris n. sp.⁸, ? Trichius rotundatus n. sp.⁸, Trichius unifasciatus n. sp.⁸, "Trichius aedilis n. sp." = Gnorimus aedilis⁸, Valgus oeningensis n. sp.⁸ Aphodius brevipennis n. sp.⁸ Heer 1862 scarab beetle from Kesselstein "Scarabaeus proserpinae n. sp." = Pentodon proserpinae⁸ Heer 1862 rhinoceras beetle from Kesselstien Anomala fugax n. sp.⁸ Heer 1862 scarab beetle from Kesselstein Serica minutula n. sp.⁸ Heer 1862 scarab beetle from Kesselstein Lepitrix germanica n. sp.⁸ Heer 1862 scarab beetle Oniticellus amplicollis n. sp.⁸ Heer 1862 scarab beetle from Kesselstein Onthophagus prodromus n. sp.⁸ Heer 1862 scarab beetle several from Kesselstein Onthophagus bisontinus n. sp.⁸ Heer 1862 scarab beetle from Insektenschicht (lower quarry) Onthophagus crassus n. sp.⁸ Heer 1862 scarab beetle from Kesselstein Gymnopleurus rotundatus n. sp.⁸ Heer 1862 scarab beetle from Kesselstein Gymnopleurus sisyphus n. sp.⁵ Heer 1847 scarab beetle Copris druidum n. sp.⁸ Heer 1862 scarab beetle from Kesselstein Trichius lugubris n. sp.⁸ Heer 1862 flower chafer ? Trichius rotundatus n. sp.⁸ Heer 1862 flower chafer from Kesselstein Trichius unifasciatus n. sp.⁸ Heer 1862 flower chafer from Kesselstein "Trichius aedilis n. sp." = Gnorimus aedilis⁸ Heer 1862 flower chafer from Kesselstein Valgus oeningensis n. sp.⁸ Heer 1862 flower chafer from Kesselstein
Coleoptera - Histeridae	Hister morosus n. sp.⁸, Hister coprolithorum n. sp.⁸, Hister vetustus n. sp.⁸, Hister mastodontis n. sp.⁸, Hister aemulus n. sp.⁸, Hister maculigerus n. sp.⁸, Hister marmoratus n. sp.⁸, Hister antiquus n. sp.⁸ Hister morosus n. sp.⁸ Heer 1862 clown beetle from Insektenschicht (lower quarry) Hister coprolithorum n. sp.⁸ Heer 1862 clown beetle numerous specimens from Kesselstein Hister vetustus n. sp.⁸ Heer 1862 clown beetle several from Kesselstein Hister mastodontis n. sp.⁸ Heer 1862 clown beetle from Kesselstein Hister aemulus n. sp.⁸ Heer 1862 clown beetle several from Kesselstein Hister maculigerus n. sp.⁸ Heer 1862 clown beetle from Kesselstein Hister marmoratus n. sp.⁸ Heer 1862 clown beetle from Kesselstein Hister antiquus n. sp.⁸ Heer 1862 clown beetle from Kesselstein
Coleoptera - Hydrophilidae	Escheria bella n. sp.⁸, Hydrous ovalis n. sp.⁸, "Hydrophilus spectabilis" = Hydrophilus spectabilis⁵, "Hydrophilus giganteus n. sp." = Hydrophilus spectabilis⁸, Hydrobiomorpha heeri n. sp.³, "Hydrous braunii" = Hydrobiomorpha braunii⁸, "Hydrous escheri n. sp." = Hydrobiomorpha braunii⁸, "Hydrous rehmanni" = Hydrochara noachica⁸, Hydrophilopsis elongata n. gen. n. sp.⁸, Hydrobius couloni n. sp.⁸ Escheria bella n. sp.⁸ Heer 1862 water scavenger beetle Hydrous ovalis n. sp.⁸ Heer 1862 water scavenger beetle from Kesselstein "Hydrophilus spectabilis" = Hydrophilus spectabilis⁵ Heer 1847 water scavenger beetle "Hydrophilus giganteus n. sp." = Hydrophilus spectabilis⁸ Heer 1847 water scavenger beetle from Kesselstein; ETHZ NO-Z51.3-39-4:4-Pl.I143b Hydrobiomorpha heeri n. sp.³ Fikáček and Schmied 2013 water scavenger beetle ETHZ NO-Z51.3-39-4:4-Pl. I 149b (label reads Hydrophilus ovalis) "Hydrous braunii" = Hydrobiomorpha braunii⁸ Heer 1847 water scavenger beetle from Kesselstein and Insektenschicht (lower quarry) "Hydrous escheri n. sp." = Hydrobiomorpha braunii⁸ Heer 1847 water scavenger beetle from Insektenschicht (lower quarry); ETHZ NO-Z51.3-39-4:4-Pl.I148 "Hydrous rehmanni" = Hydrochara noachica⁸ Heer 1847 water scavenger beetle Hydrophilopsis elongata n. gen. n. sp.⁸ Heer 1862 water scavenger beetle from Insektenschicht (lower quarry) Hydrobius couloni n. sp.⁸ Heer 1862 water scavenger beetle from Kesselstein
Coleoptera - Carabidae	Amara princeps n. sp.⁸, Amara pinguicula n. sp.⁸, Amara primigenia n. sp.⁸, Sinis brevicollis n. gen. n. sp.⁸, Pterostichus vetustus n. sp.⁸, Pterostichus minutulus n. sp.⁸, Badister grandis n. sp.⁸, Badister macrocephalus n. sp.⁸, Badister fragilis n. sp.⁸, Harpalus constrictus n. sp.⁸, Harpalus stierlini n. sp.⁸, Harpalus stygius n. sp.⁸, Harpalus tardigradus n. sp.⁸, Dichirotrichus lividus⁸, Carabites laesicollis n. sp.⁸, Glenopterus laevigatus n. gen. n. sp.⁵, Trechinites clairvillii n. sp.⁸, Trechinites oblongus n. sp.⁸, Nebria pluto n. sp.⁸, "Calosoma deplanatum n. sp." = Campalita deplanatum⁷, Calosoma catenulatum n. sp.⁷, Calosoma nauckianum n. sp.⁷, Calosoma escrobiculatum n. sp.⁷, Calosoma escheri n. sp.⁷ Amara princeps n. sp.⁸ Heer 1862 ground beetle 1 from Insektenschicht (lower quarry), 1 from Kesselstein Amara pinguicula n. sp.⁸ Heer 1862 ground beetle from Insektenschicht (lower quarry) Amara primigenia n. sp.⁸ Heer 1862 ground beetle from Kesselstein Sinis brevicollis n. gen. n. sp.⁸ Heer 1862 ground beetle 1 male, 1 female from lower quarry Pterostichus vetustus n. sp.⁸ Heer 1862 ground beetle from Insektenschicht (lower quarry) Pterostichus minutulus n. sp.⁸ Heer 1862 ground beetle from Insektenschicht (lower quarry) Badister grandis n. sp.⁸ Heer 1862 ground beetle from Kesselstein Badister macrocephalus n. sp.⁸ Heer 1862 ground beetle "multiple examples" in Kesselstein, including var. major Badister fragilis n. sp.⁸ Heer 1862 ground beetle from Kesselstein (2 var. paulo major) Harpalus constrictus n. sp.⁸ Heer 1862 ground beetle from Kesselstein Harpalus stierlini n. sp.⁸ Heer 1862 ground beetle Harpalus stygius n. sp.⁸ Heer 1862 ground beetle from Insektenschicht (lower quarry) Harpalus tardigradus n. sp.⁸ Heer 1862 ground beetle from Insektenschicht (lower quarry) Dichirotrichus lividus⁸ Heer 1862 ground beetle from Kesselstein Carabites laesicollis n. sp.⁸ Heer 1862 ground beetle Glenopterus laevigatus n. gen. n. sp.⁵ Heer 1847 ground beetle Trechinites clairvillii n. sp.⁸ Heer 1862 ground beetle from lower quarry Trechinites oblongus n. sp.⁸ Heer 1862 ground beetle from lower quarry Nebria pluto n. sp.⁸ Heer 1862 ground beetle 1 from lower quarry, 2 from Kesselstein "Calosoma deplanatum n. sp." = Campalita deplanatum⁷ Heer 1860 ground beetle Calosoma catenulatum n. sp.⁷ Heer 1860 ground beetle Kesselstein Calosoma nauckianum n. sp.⁷ Heer 1860 ground beetle Calosoma escrobiculatum n. sp.⁷ Heer 1860 ground beetle Calosoma escheri n. sp.⁷ Heer 1860 ground beetle
Coleoptera - Gyrinidae	"Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris⁸, Miodineutes oeningensis n. gen. n. sp.⁴, "Dineutus insignis n. sp." = Miodineutes insignis⁸, Miodineutes heeri n. sp.⁴ "Dineutus longiventris n. sp." = Dineutus (Cyclinus) longiventris⁸ Heer 1862 whirligig beetle from Kesselstein Miodineutes oeningensis n. gen. n. sp.⁴ Hatch 1927 whirligig beetle Dineutus insignis specimen in pl 2, fig 28 of Heer "Dineutus insignis n. sp." = Miodineutes insignis⁸ Heer 1862 whirligig beetle from Kesselstein Miodineutes heeri n. sp.⁴ Hatch 1927 whirligig beetle Dineutus longiventris Specimen in pl 2, fig 25 of Heer
Coleoptera - Dytiscidae	"Cybister nicoleti" = Megadytes nicoleti⁸, Cybister atavus n. sp.⁸, Cybister agassizi n. sp.⁸, "Hydaticus areolatus n. sp." = Graphoderus heeri⁸, Dytiscus lavateri⁸, Colymbetes aemulus n. sp.⁸, "Hydroporus benzeli n. sp." = Nebrioporus benzeli⁸ "Cybister nicoleti" = Megadytes nicoleti⁸ Heer 1862 predaceous diving beetle from Kesselstein Cybister atavus n. sp.⁸ Heer 1862 predaceous diving beetle from Kesselstein Cybister agassizi n. sp.⁸ Heer 1862 predaceous diving beetle from Kesselstein "Hydaticus areolatus n. sp." = Graphoderus heeri⁸ Nilsson 2001 predaceous diving beetle Dytiscus lavateri⁸ Heer 1847 predaceous diving beetle from Kesselstein Colymbetes aemulus n. sp.⁸ Heer 1862 predaceous diving beetle from Kesselstein "Hydroporus benzeli n. sp." = Nebrioporus benzeli⁸ Heer 1862 predaceous diving beetle
Lepidoptera - Bombycidae	Bombycites buchii n. sp.⁹, "Bombycites" oeningensis n. sp. Bombycites buchii n. sp.⁹ Heer 1864 moth "Bombycites" oeningensis n. sp. Heer 1849 moth pupa
Diptera - Chironomidae	Chironomus gaudini n. sp.⁹ Chironomus gaudini n. sp.⁹ Heer 1864 non biting midge
Diptera - Syrphidae	Syrphus bremii n. sp.⁹, Syrphus schellenbergi n. sp.⁹ Syrphus bremii n. sp.⁹ Heer 1864 hover fly Syrphus schellenbergi n. sp.⁹ Heer 1864 hover fly
Diptera - Oestridae	"Dipterites obovatus n. sp." = Adipterites obovatus⁹ "Dipterites obovatus n. sp." = Adipterites obovatus⁹ Heer 1864 bot fly
Diptera - Tabanidae	"? Hexatoma oeningensis n. sp." = Heptatoma oeningensis⁹ "? Hexatoma oeningensis n. sp." = Heptatoma oeningensis⁹ Heer 1864 horse fly
Diptera - Bibionidae	Bibio fusiformis¹⁵, Bibio brevis¹⁵, Bibio crassinervis n. sp.¹⁵, Bibio moestus¹⁵, Bibio angustatus¹⁵, Bibio pulchellus¹⁵, Dilophus pumilio n. sp.¹⁵, Plecia hilaris¹⁵, "Protomyia jucunda" = Plecia jucunda, Plecia macilenta n. sp.¹⁵, "Protomyia speciosa n. sp." = Penthetria amoena⁹, Penthetria brevicollis¹⁵, Bibio elongatus¹⁵ Bibio fusiformis¹⁵ Heer 1849 march fly ETHZ Pl. I 858 (x2), 861 (x6) Bibio brevis¹⁵ Heer 1849 march fly Some in lot Pl. I 856; ETHZ Pl.I 865, Pl. I 887 Bibio crassinervis n. sp.¹⁵ Skartveit and Pika 2014 march fly ETHZ Pl. I 851, Pl.I 851(2), Pl. I 862, Pl. I 875, Pl. I 888 Bibio moestus¹⁵ Heer 1849 march fly ETHZ Pl. I 855, 857(x2) Bibio angustatus¹⁵ Heer 1849 march fly ETHZ Pl. I 851 Bibio pulchellus¹⁵ Heer 1849 march fly ETHZ Pl. I 854 Dilophus pumilio n. sp.¹⁵ Skartveit and Pika 2014 march fly ETHZ Pl. I 886 Plecia hilaris¹⁵ Heer 1849 love bug ETHZ Pl. I 850 "Protomyia jucunda" = Plecia jucunda Heer 1849 love bug ETHZ Pl. I 871, 853(x2), 856, 861, 867, 869 (some labeled by Brongniart fide Skartveit & Pika 2014) Plecia macilenta n. sp.¹⁵ Skartveit and Pika 2014 love bug ETHZ Pl. I 853, 853 female "Protomyia speciosa n. sp." = Penthetria amoena⁹ Heer 1849 march fly ETHZ Pl. I 873; also Pl. I 872, 876 as P. amoena in Skartveit & Pika, 2014 Penthetria brevicollis¹⁵ Heer 1849 march fly ETHZ Pl. I 870 Bibio elongatus¹⁵ Heer 1849 march fly ETHZ S.N. 189 Pl. I 863 (labeled by Heer)
Diptera - Sciaridae	Sciara deleta n. sp.⁹ Sciara deleta n. sp.⁹ Heer 1864 dark-winged fungus gnat
Diptera - Mycetophilidae	Mycetophila pusillima n. sp.⁹, Mycetophila orci n. sp.⁹ Mycetophila pusillima n. sp.⁹ Heer 1864 fungus gnat Mycetophila orci n. sp.⁹ Heer 1864 fungus gnat
Diptera - Cecidomyiidae	Cecidomyia bremii n. sp.⁹ Cecidomyia bremii n. sp.⁹ Heer 1864 gall midge

¹Cockerell 1909; ²Cockerell 1915; ³Fikáček and Schmied 2013; ⁴Hatch 1927; ⁵Heer 1847; ⁶Heer 1853; ⁷Heer 1860; ⁸Heer 1862; ⁹Heer 1864; ¹⁰Heer 1864; ¹¹Heer 1867; ¹²Heer 1869; ¹³Heer 1879; ¹⁴Legalov 2018; ¹⁵Skartveit and Pika 2014; ¹⁶Zeuner and Manning 1976