São José de Itaborai (Eocene of Brazil)

Also known as Companhia Nacional de Cimento Portland quarry, 1968 fissure

Where: Rio de Janeiro, Brazil (22.8° S, 42.9° W: paleocoordinates 25.6° S, 31.7° W)

• coordinate stated in text

• local area-level geographic resolution

When: Itaboraian mammal zone, Itaboraí Formation, Early/Lower Eocene (55.8 - 48.6 Ma)

• Late Paleocene marl fillings in Early Eocene karst

•Itaboraian SALMA

•Middle sequence (S2) of the Itaborai Basin sediments

•Oliveira and Goin argue that it is more likely to be late Paleocene-early Eocene in age

•Early Eocene (Itaboraian SALMA); 53-50 Ma according to Woodburne et al. (2014) and Goin et al. (2018).

• group of beds-level stratigraphic resolution

Environment/lithology: fissure fill; marl

• Fissure filling deposits with marls and collapsed breccias whose composition is similar to the gray limestone facies, although poorly consolidated, and transported into these cavities by heavy rains and gravitational flows

Size classes: macrofossils, mesofossils

Collected by Carlos de Paula Couto, Ney Vidal and others in 1948-1950

• Quarry under the management of the Companhia Nacional de Cimento Portland (Maui)

Primary reference: E. V. Oliveira and F. J. Goin. 2011. A reassessment of bunodont metatherians from the Paleogene of Itaborai (Brazil): Systematics and the age of the Itaborian SALMA. Revista Brasileira de Paleontologia 14(2):105-136 [P. Mannion/P. Mannion]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 118314: authorized by Philip Mannion, entered by Philip Mannion on 12.10.2011, edited by Mark Uhen, Gustavo Ballen, Richard Butler, Jelle Zijlstra, Grace Varnham and Kateryn Pino

Creative Commons license: CC BY (attribution)

Taxonomic list

Aves
 Aves -
Aves "sp. A"14 bird
MN 4115V (right carpometacarpus)
Aves "sp. B"14 bird
MN 4119−V (distal left tibiotarsus)
Aves "sp. C"14 bird
MN 4116−V (distal right tibiotarsus)
Aves "sp. D"14 bird
MN 4117−V (distal left tibiotarsus). An additional fragmentary distal end of left tarsometatarsus (MN 4120−V) may belong to this species or species E.
Aves "sp. E"14 bird
MN 4118−V (distal left tibiotarsus). An additional fragmentary distal end of left tarsometatarsus (MN 4120−V) may belong to this species or species D.
 Cuculiformes - Cuculidae
Eutreptodactylus itaboraiensis n. gen. n. sp.2 Baird and Vickers-Rich 1997 cuckoo
MNB 4083v - holotype [now lost - only a cast and original illustrations remain] (Incomplete distal fragment of a right tarsometatarsus)
 Gruiformes - Phorusrhacidae
Paleopsilopterus itaboraiensis n. gen. n. sp.1 Alvarenga 1985 terror bird
MNRJ-4040-V - holotype (proximal portion of the right tarsometatarsus); paratypes: DGM-1431-R (left and right tibiotarsus, proximally deformed and incomplete)
 Gruiformes -
Itaboravis elaphrocnemoides n. gen. n. sp.14
Itaboravis elaphrocnemoides n. gen. n. sp.14 Mayr et al. 2011 bird
MN 4114V - holotype (left coracoid); referred specimens: MN 4113V (right humerus) and MN 4121V (distal end of left humerus)
Reptilia
 Crocodylia -
Eocaiman itaboraiensis n. sp.26 Pinheiro et al. 2013 crocodilian
MCT 1791-R - type (a well-preserved small left anterior dentary fragment, broken at the level of the eleventh alveolus and bearing four fully preserved teeth); referred specimens: MCT 1792-R (small left anterior dentary fragment with no teeth); MCT 1793-R (small left anterior dentary fragment with no preserved teeth); tentatively referred specimen: MCT 1794-R (an isolated small blunt tooth [root and crown]). The specimens referred to E. itaboraiensis sp. nov. have been kept associated in the MCT collection but lack information regarding their precise provenance. However, the different colouration of these fossils and the carbonate matrix (varying from yellowish to greyish-white) suggests these specimens were recovered from different S2 fissures infills.
Mammalia
 Theriamorpha - Jaskhadelphyidae
Minusculodelphis modicum n. sp.18 Oliveira et al. 2016 metatherian
DGM 921a-M, an incomplete right dentary with complete m1–4 (holotype). MCT 2815-M, a complete right M3; MCN-PV 1792, a complete right M3; MN 2479-V, an incomplete right maxilla with complete M2 and incomplete M3; DGM 921b-M, an incomplete right dentary with roots of m1, m2, portion of m4, and with complete m3; DGM921c-M, a complete left femur.
Minusculodelphis minimus n. sp.24 Paula Couto 1962 metatherian
MNRJ 2342-V (holotype), 2503-V
 Theriamorpha -
Marmosopsis juradoi n. sp.24 Paula Couto 1962 metatherian
All the specimens housed at MNRJ: MNRJ 2343-V (holotype), 2344-V, 2345-V, 2346-V, 2348-V:2350-V, 2356-V:2357-V, 2359-V:2360-V, 2460-V: 2473-V, 2434-V (this record must be a mistyping for 2474 from the order of the hypodigm report), 2475-V:2481-V (MNRJ 2482-V is reported in the measurements section. However, it is missing in the hypodigmia section. Among the specimens reported as examined, only MNRJ 2481-V shows upper M1-4 present, so it is very likely that 2482 is but a mistyping for 2481. This requires confirmation by direct measurements of those specimens housed at MNRJ), 2483-V:2487-V, 2495-V:2502-V.
 Theriamorpha - Pucadelphyidae
Itaboraidelphys camposi13 Marshall and de Muizon 1984 metatherian
Holotype (DGM 817-M, left partial dentary with p3-m4); DGM 804-M, right dentary fragment with m2-3 and m1, m4 alveoli; DGM 814-M, right dentary fragment with m1-4; DGM 923-M, right dentary fragment with p2-m1 and m2 alveoli; DGM 926-M, maxillary fragment with M2-3 and M4 alveolus; MNRJ 2878-V a, isolated M2; MNRJ 2878-V b, isolated M3
 Theriamorpha - Peradectidae
Procaroloameghinia pricei n. gen. n. sp. Marshall 1982 marsupial
DGM 805-M and 924-M - holotype (two pieces of the same right dentary)
 Theriamorpha - Herpetotheriidae
 Theriamorpha -
Australidelphia indet.11 Szalay 1982 marsupial
"stem Australidelphia"
 Dasyuromorphia - Malleodectidae
Gaylordia mater n. sp.17, "Gaylordia macrocynodonta n. gen. n. sp." = Gaylordia macrocynodonta19, "Xenodelphis doelloi n. sp." = Gaylordia macrocynodonta24
Gaylordia mater n. sp.17 Oliveira and Goin 2015 marsupial
Holotype. DGM 810-M, a partial right dentary with complete m2-4. Hypodigm. The type and MCT 2383-M (DGM 644-M), incomplete left dentary with m1-2 and m4; MCT 2384-M (DGM 645-M), incomplete left dentary with p3, m1-2, incomplete m3 and complete m4; MCT 2386-M, incomplete right dentary with m3 and alveoli of p3 (only the posterior alveolus is complete), m1, m2 and m4; MCT 2387-M, incomplete right dentary with complete p3 and m1, alveoli of p2, and m2; MCT 2774-M, complete left m4; MCT 2775- M, incomplete right m3; MCT 2776-M, complete right m3; MCT 2777-M, complete right M2; MCT 2778-M, incomplete left M2; MCT 2779-M, incomplete right M2; MCT 2780-M, complete left M3; MCT 2781-M, incomplete right M3.
"Gaylordia macrocynodonta n. gen. n. sp." = Gaylordia macrocynodonta19 Paula Couto 1952 marsupial
DGM 329-M - holotype (anterior part of skull and mesial fragment of left mandibular ramus); referred material: MNRJ 1366-V (median fragment of left lower jaw), 1357-1358 (left and right mandivular rami), AMNH 49801 (median portion of right lower jaw) and DGM 186-M (left mandibular ramus)
"Xenodelphis doelloi n. sp." = Gaylordia macrocynodonta24 Paula Couto 1952 marsupial
MNRJ 2504-V (holotype)
 Theriamorpha -
Monodelphopsis travassosi n. gen. n. sp.19
Monodelphopsis travassosi n. gen. n. sp.19 Paula Couto 1952 marsupial
MNRJ 1365-V - holotype (fragment of left lower jaw)
 Microbiotheria - Microbiotheriidae
Microbiotheriidae indet.11, "Mirandatherium alipioi" = Mirandatherium alipioi24, "Mirandaia ribeiroi n. gen. n. sp." = Mirandatherium alipioi19
Microbiotheriidae indet.11 Ameghino 1887 marsupial
"Mirandatherium alipioi" = Mirandatherium alipioi24 Paula Couto 1952 marsupial
=Mirandaia ribeiroi, replacement name following Paula Couto (1952)
"Mirandaia ribeiroi n. gen. n. sp." = Mirandatherium alipioi19 Paula Couto 1952 marsupial
MNRJ 1359-V - holotype (median fragment of right mandibular ramus); paratypes: DGM 361 and AMNH 49859 (two partial right mandibular rami); referred material: MNRJ 1362 (median fragment of right lower jaw), 1363 (part of right lower jaw), AMNH 49802 (median portion of right lower jaw), DGM 360-M (posterior part of right mandibular ramus) and 362-M (posterior fragment of right lower jaw)
 Theriamorpha -
Gashternia carioca n. sp.10 Goin and Oliveira 2007 marsupial
Holotype only. MCN-PV 1801, a fragment of left maxilla with complete P3 and M1, and roots M2.
 Theriamorpha - Sternbergiidae
Carolopaulacoutoia itaboraiensis17 Paula Couto 1970 marsupial
Didelphopsis cabrerai n. gen. n. sp.19 Paula Couto 1952 marsupial
MNRJ 1429-V - holotype (left maxilla); paratypes: DGM 243-M (right mandibular ramus) and MNRJ 1345-V (right m4); referred material: MNRJ 1346-V (m3?) and 1347-V (m4)
 Theriamorpha - Derorhynchidae
Derorhynchus singularis n. gen. n. sp.19, Diogenesia brevirostris n. gen. n. sp.16
Derorhynchus singularis n. gen. n. sp.19 Paula Couto 1952 marsupial
MNRJ 1348-V - holotype (incomplete right mandibular ramus); paratype: AMNH 49826 (fragment of right mandibular ramus)
Diogenesia brevirostris n. gen. n. sp.16 Oliveira et al. 2021 marsupial
Holotype: DGM 650-M, incomplete right dentary with alveoli of p1-2, roots of p3-m1, and complete m2-4.
 Didelphimorphia - Didelphidae
Didelphidae indet.19 opossum
MNRJ 1367-V and1368-V (two very small posterior fragments of right lower jaws)
 Theriamorpha - Protodidelphidae
Carolocoutoia ferigoloi n. gen. n. sp., Bergqvistherium primigenia n. gen. n. sp.5, Protodidelphis vanzolinii n. gen. n. sp.19, "Robertbutleria mastodontoidea n. gen. n. sp." = Protodidelphis mastodontoides, Periprotodidelphis bergqvistae n. gen. n. sp., Guggenheimia brasiliensis n. gen. n. sp.19, Guggenheimia crocheti n. sp.
Carolocoutoia ferigoloi n. gen. n. sp. Goin et al. 1998 marsupial
MCN-PV 1802 - holotype (upper left molar, M3?)
Bergqvistherium primigenia n. gen. n. sp.5 Carneiro 2018 marsupial
Protodidelphis vanzolinii n. gen. n. sp.19 Paula Couto 1952 marsupial
DGM 271-M - holotype (facial and palatal portion of an incomplete skull); paratype: DGM 250-M (right mandibular ramus); referred specimens: DGM 319-M (left mandibular ramus), 303-M (fragment of right maxilla), AMNH 49857 (right mandibular ramus), MNRJ 1371-V (left m1 or m2), 1426-V (left m1), 1427-V (right m3), AMNH 49803 (right m2) and 49858 (portion of left mandibular ramus)
"Robertbutleria mastodontoidea n. gen. n. sp." = Protodidelphis mastodontoides Marshall 1987 marsupial
DGM 896-M - holotype (left M1); referred material: MCT 2769-M (ex DGM 896b-M; complete left M2), MCN-PV 1799 (complete left dP3), MCN-PV 1806 (complete left M3), MNRJ 2897(b)-V (nearly complete right M2), MNRJ 2897(c)-V (incomplete right M2), MNRJ 2897(d)-V (incomplete left M3), AMNH 49858 (incomplete left dentary with p2-m2), MCT 2817-M (nearly complete left m2), MCT 2818-M (nearly complete left m2), MCT 2819-M (incomplete left m2), MNRJ 1371-V (complete left m1), MCN-PV 1819 (nearly complete left m3), MCN-PV 1816 (a nearly complete left m3), MCN-PV 1817 (incomplete left m3), MCN-PV 1820 (complete left m4)
Periprotodidelphis bergqvistae n. gen. n. sp. Oliveira and Goin 2011 marsupial
DGM 806-M (left M?3)
Guggenheimia brasiliensis n. gen. n. sp.19 Paula Couto 1952 marsupial
DGM 297-M - holotype (incomplete left mandibular ramus)
Guggenheimia crocheti n. sp. Oliveira and Goin 2011 marsupial
DGM 314-M - type (incomplete left dentary); hypodigm: DGM 315-M (incomplete left dentary), MNRJ 2899(a)-V (right m3), MCT 2783-M (left m3) and MCT 2782-M (left M1)
 Marsupialia -
Paucituberculata indet.11 Ameghino 1894 marsupial
 Paucituberculata -
Riolestes capricornicus n. gen. n. sp.9
Riolestes capricornicus n. gen. n. sp.9 Goin et al. 2009 marsupial
MCN-PV 1790 - holotype (an isolated m1)
 Polydolopimorphia - Polydolopidae
Polydolopidae indet.11 Ameghino 1897 metatherian
 Polydolopimorphia -
Zeusdelphys complicatus n. gen. n. sp., aff. Bobbschaefferia sp., "Schaefferia fluminensis n. gen. n. sp." = Bobbschaefferia fluminensis19
Zeusdelphys complicatus n. gen. n. sp. Marshall 1987 metatherian
MCT 2830-M - holotype (ex- DGM 896a-M, a left ?M1)
aff. Bobbschaefferia sp. Paula Couto 1970 metatherian
MCN-PV 1797 (an incomplete right maxilla) and MCT 2816-M (right m3)
"Schaefferia fluminensis n. gen. n. sp." = Bobbschaefferia fluminensis19 Paula Couto 1970 metatherian
MNRJ 1350-V - holotype (portion of left mandibular ramus); paratypes now comprise the type of Guggenheimia crocheti: DGM 314-M (incomplete left mandibular ramus) and 315 (fragment of left manibular ramus)
 Polydolopimorphia - Bonapartheriidae
"Epidolops ameghinoi n. gen. n. sp." = Epidolops ameghinoi20, "Epidolops gracilis n. sp." = Epidolops ameghinoi20
"Epidolops ameghinoi n. gen. n. sp." = Epidolops ameghinoi20 Paula Couto 1952 metatherian
DGM 321-M - holotype (almost complete but much crushed and flattened skull, lacking the upper part of the braincase and the part posterior to the basisphenoid, as well as the anterior end of the premaxillae; almost complete mandible, lacking the anterior end of the incisor border); referred material: DGM 170-197-M and 207-209-M (fragmentary right and left lower jaws with and without teeth), DGM 198-206-M (partial right and left maxillae with teeth), MNRJ 1372-1374-V, 1376-1378-V, 1381-1392-V, 1394-V and 1395-V, 1397-V to 1405-V (incomplete right and left lower jaws with and without teeth), 1407-V to 1416-V (portions of right and left maxillae with and without teeth), 1417-V to 1419-V (partial right P3), 1420-V (incomplete left P3), 1421-V (left M2), 1422-V (right Ml), 1423-V (anterior part of left M1), 1424-V (right I?2), AMNH 49806 to 49809, 49811 and 49813 (fragmentary lower jaws with teeth)
"Epidolops gracilis n. sp." = Epidolops ameghinoi20 Paula Couto 1952 metatherian
DGM 188-M - type (fragment of right lower jaw); paratype: MNRJ 1400-V (partial left lower jaw); referred material: DGM 191-M (part of left lower jaw with P3, M1 present), 198-M (portion of left maxilla with P3, M1-2 present), MNRJ 1390-V and 1391-V (anterior parts, respectively, of right lower jaw with roots of P2 and with P3 present, and of left lower jaw with P2-3 present), 1412-V (part of right maxilla with P3, M1-2 present), 1404-V (fragment of left maxilla with P3 present), 1411-V (portion of right maxilla with P3 present), AMNH 49809, 49810, 49812 to 49814 and 49860 (lower jaw fragments with teeth)
 Theriamorpha - Stagodontidae
Eobrasilia coutoi24 Simpson 1947 metatherian
AMNH 39424 (holotype); Marshall (1984) adds the following specimens: DGM 919-M, DGM 896-M(a)
 Theriamorpha - Pediomyidae
cf. Pediomyidae indet.17 Simpson 1927 metatherian
 Sparassodonta - Borhyaenidae
Borhyaenidae indet.11 metatherian
Nemolestes brasiliensis27 Rangel et al. 2023 metatherian
MNRJ 1344-V (large left lower molar)
 Sparassodonta - Hathlyacynidae
"Hathlyacynidae indet." = Hathlyacininae11
"Hathlyacynidae indet." = Hathlyacininae11 Ameghino 1894 metatherian
 Sparassodonta -
"Patene simpsoni n. sp." = Patene simpsoni20, "Ischyrodidelphis castelllanosi n. gen. n. sp." = Patene simpsoni19
"Patene simpsoni n. sp." = Patene simpsoni20 Paula Couto 1952 metatherian
MNRJ 1331-V - type (incomplete right maxilla); paratypes: DGM 324-M (fragment of right lower jaw with alveolus of P2 and roots of P3, anterior root of M1, talonid and anterior root of M2, with M3, and with roots of M4 present), MNRJ 1332-V (right M4), 1335-V (right Ml or M2); referred material: MNRJ 1336-V to 1343-V and 1428-V (eight left lower molars, and one right lower molar, reduced to the trigonid) and DGM 331-M (part of left maxilla)
"Ischyrodidelphis castelllanosi n. gen. n. sp." = Patene simpsoni19 Paula Couto 1952 metatherian
MNRJ 1351-V - holotype (median fragment of right mandibular ramus); paratypes: MNRJ 1352 and 1353 (right m2s), 1354 (left m2) and 1425 (right p3)
 Theriamorpha -
Austropediomys marshalli n. gen. n. sp.6
Austropediomys marshalli n. gen. n. sp.6 Carneiro et al. 2018 metatherian
Holotype DGM 808-M, incomplete left maxilla with M2–4; MCT 2798-M, incomplete left M1; MCT 2799-M, complete left M3; MCT 2800- M, incomplete right M4; MCT 4386-M, left DP3
 Placentalia -
Notoungulata indet.4 Roth 1903 notoungulate
MNHN-F-BRD 23 (left isolated petrosal)
 Notoungulata - Oldfieldthomasiidae
"Homalostylops atavus n. sp." = Itaboraitherium atavum23, "Henricosbornia magellanica n. sp." = Colbertia magellanica21, Camargomendesia pristina n. gen. n. sp.3
"Homalostylops atavus n. sp." = Itaboraitherium atavum23 Couto 1954 notoungulate
MNRJ 1992-V - type (posterior fragment of a right mandibular ramus with m1-3); paratypes: MNRJ 1990-V (2 right upper molars) and MNRJ 2245-V (one right and 3 left upper molars
"Henricosbornia magellanica n. sp." = Colbertia magellanica21 Price and Paula Couto 1950 notoungulate
DGM 152-M - type (left m3); paratype: DGM 153-M (fragment of left lower jaw); referred material: MNRJ 1865-V to 1909-V and 1944-V to 1977-V (partial right and left lower jaws with cheek teeth), 1910-V to 1925-V (fragmentary left and right maxillae with cheek teeth), 1926-V and 1927-V (series of associated left and right P2-M3 and P2-M2, respectively), 1928-V to 1943-V (isolated right and left upper cheek teeth), 1978-V to 1987-V (isolated right and left lower cheek teeth), DGM 236-M, 242-M, 252-M, 254-M, 256-M to 259-M, 264-M to 266-M, 269-M, 272-M, 274-M to 276-M, 279-M to 283-M, 285-M and 286-M, 289-M, 291-M, 293-M, 295-M, 298-M and 299-M, 316-M, 322-M, 377-M to 382-M (incomplete right and left lower jaws with cheek teeth), 310-M (almost complete mandible with right P2, P4-M3, and left P2-M3 [cast AMNH 49852]), 284-M, 288-M, 302-M, 306-M, 357-M, 372-M to 376-M (partial right and left maxillae with cheek teeth), 370-M and 371-M (fragmentary skulls with right P2-M3 and left P2-4, and right P1-Mi and left P4-M3, respectively), AMNH 49840, 49867 to 49877 (incomplete right and left maxillae with cheek teeth), 49841 to 49846, 49864 to 49866, 49878 to 49890 (partial right and left lower jaws with cheek teeth)
Camargomendesia pristina n. gen. n. sp.3 notoungulate
 Notoungulata - Henricosborniidae
Nanolophodon tutuca n. gen. n. sp.7
Nanolophodon tutuca n. gen. n. sp.7 Castro et al. 2021 notoungulate
Holotype: MCT 4419M, a fragment of left maxilla with P3-M2 (moderate to advanced wear); MCT 4400M, isolated right P2; MCT 4397M, isolated left P3; MCT 4396M, isolated left P4; MCT 4390M, isolated right M1; MCT 4388M, isolated right M2; MCT 4389M, isolated right M2; MCT 4420M, isolated right M2; MCT 4421M, isolated left M3; MCT 4403M, isolated left lower molar; MCT 4422M, fragment of left jaw with dp2, dp3, dp4 and m1 (in alveolus).
 Xenungulata - Carodniidae
Carodnia vieirai n. sp.22 Paula Couto 1952 placental
DGM 333-M - type (fragments of right and left maxillae; fragment of the occiput with the condyles; nearly complete mandible; all the cervical and the first two dorsal vertebrae; fragmentary first left rib; proximal fragment of left scapula; part of the bones of the anterior and posterior extremities, and other fragmentary skeletal remains); referred material: DGM 334-M (mandible of an immature individual), 335-M (fragment of right maxilla and anterior part of the skull of another individual), , 336-M- and 337-M (various skeletal remains), AMNH 49828 (left m3), MNRH 1843-V-1857-V (teeth)
 Astrapotheria - Trigonostylopidae
Tetragonostylops apthomasi25 Price and Paula Couto 1950 placental
DGM 154-M - type (right m1?); paratypes: DGM 155-M and 156-M (molars); numerous referred material
Tetragonostylops apthomasi25 Price and Paula Couto 1950 placental
MNRJ 1823-V-1839-V (small teeth)
 Panameriungulata - Didolodontidae
Didolodontidae indet.22, "Ernestokokenia protocenica n. sp." = Ricardocifellia protocenica22, Lamegoia conodonta n. gen. n. sp.22
Didolodontidae indet.22 Simpson 1934 placental
MNRJ 1841-V and 1842-V (left and right p4)
"Ernestokokenia protocenica n. sp." = Ricardocifellia protocenica22 Paula Couto 1952 placental
MNRJ 1430-V - type (fragment of left maxilla); paratype: MNRJ 1431-V (part of left lower jaw); referred material: MNRJ 1432-V (small part of a right lower jaw with M3), 1433-V, 1434-V (respectively left and right M3), 1435-V, 1436-V, 1438-V (all right M2), 1439-V (left M2), 1440-V-1442-V (all right M1), 1443-V, 1444-V (left Ml), 1445-V (right P?4), 1446-V-1448-V (left P?4), 1449-V, 1450-V (left P?s), 1451-V, 1452-V (left M3), 1453-V-1455-V (right M2), 1456-V (left M2), 1457-V (right Ml), 1458-V, 1459-V (left Ml), 1460-V, 1461-V (left P3), 1462-V (right M2-3), AMNH 49819 (partial right lower jaw with M2-3), 49820 (left P3), 49821 (left P?s), 49822 (left P?4), 49823 (four M1, unassociated), 49824 (two left M3 and one right M3), 49825 (two right M2 and one left M2) 49827 (one left M2 and one left M3), 49829 (three left Ml)
Lamegoia conodonta n. gen. n. sp.22 Paula Couto 1952 placental
MNRJ 1463-V - holotype (left m2?); paratypes: MNRJ 1464-V and 1465-V (left m1 and right m2 respectively)
 Panameriungulata - Proterotheriidae
"Anisolambda prodromus n. sp." = Paranisolambda prodromus22
"Anisolambda prodromus n. sp." = Paranisolambda prodromus22 Paula Couto 1952 placental
DGM 262-M - type (right lower jaw of a very young individual); paratypes: DGM 304-M (part of a right upper jaw), 239-M (partial left lower jaw) and 273-M (fragment of right and left upper jaw); referred material: MNRJ 1479-V (left M?2), 1493-V, 1494-V (right M?2), 1495-V, 1496-V (left M?2), 1858-V (fragment of left lower jaw with DM3-4 and Ml present), 1859-V (left M3), DGM 238-M (fragment of right lower jaw with M1-2 present), 307-M (partial left upper jaw with P2, DM3-4, M1 present), 310-M (fragment of right upper jaw with M2-3, and incomplete posterior part of the skull)
 Panameriungulata - Protolipternidae
Protolipterna ellipsodontoides n. gen. n. sp.8, Asmithwoodwardia scotti n. sp.22, "Ernestokokenia parayirunhor n. sp." = Miguelsoria parayirunhor22
Protolipterna ellipsodontoides n. gen. n. sp.8 Cifelli 1983 placental
DNPM LE444A (cast, AMNH 109542), mandible with right C, alveoli of right and left lower P1-2, and with right and left P3-M3; associated right maxillary fragment with M1-3; hundreds of other specimens
Asmithwoodwardia scotti n. sp.22 Paula Couto 1952 placental
DGM 358-M - type (badly crushed but nearly complete skull, with P3, M1-3 and left P3-4, M1-3, accompanied by its nearly complete mandible with all the teeth present, the tip of an upper C, and the greater part of the axis); paratype: AMNH 49817 (fragment of right lower jaw)
"Ernestokokenia parayirunhor n. sp." = Miguelsoria parayirunhor22 Paula Couto 1952 placental
MNRJ 1468-V - type (almost complete right lower jaw); paratypes: DGM 305-M (part of left upper jaw) and 249-M (posterior part of left lower jaw); referred material: DGM 330-M (fragment of lower jaw; with the posterior part of the symphysis, and right C, P2-4, M1l2, and left C, P2 present), 246-M (part of lower jaw, with the posterior part of the symphysis, and the right P4, M1-3 present), 255-M (part of right lower jaw, with P4, M13 present), 290-M (part of left lower jaw, with M2-3 present), 296-M (part of left lower jaw, with M2-3 present), MNRJ 1469-V (anterior part of left lower jaw, with Pc-M1 present), 1470-V (part of left lower jaw, with Ml_2 present), 1466-V (part of left lower jaw, with M1-2 present), 1467-V (part of left lower jaw, with M2-3 present), 1861-V (part of left lower jaw, with M2 present), 1862-V (left M1,very worn) and AMNH 49815 (portion of left lower jaw, with M1.2 present)
 Panameriungulata - Macraucheniidae
Victorlemoineia prototypica n. sp.22 Paula Couto 1952 placental
MNRJ 1470-V - type (right m2 or 3); paratype: MNRJ 1471-V (left m3); referred material: MNRJ 1472-V-1474-V (three left upper molars), 1475-V-1478-V (four right upper molars), 1480-V (fragment of left lower jaw with P4 present), 1481-V (right P?3), 1482-V-1486-V (right lower molars), 1487-V, 1488-V (left P3?s), 1489-V, 1490-V, 1492-V (left lower molars) and DGM 268-M (fragment of left lower jaw)
 Theriamorpha -
Palaeocladosictis mosesi n. gen. n. sp.12
Palaeocladosictis mosesi n. gen. n. sp.12 Paula Couto 1961 placental
MNRJ 2671-V, DP4 or P4
 Cingulata - Dasypodidae
Riostegotherium yanei n. sp.15 Oliveira and Bergqvist 1998 armadillo
MCN-PV 1774 (holotype) and 1775, pelvic osteoderms; MCN-PV 1776, 1778, 1779, MCT 2081 M, MLP 75-XII-26-1, MLP 75-XII-26-2, isolated moveable osteoderms; MCN-PV 1777, caudal osteoderm