Grube Messel Pit (Senckenberg SMF collection) (Eocene of Germany)

Where: Hessen, Germany (49.9° N, 8.8° E: paleocoordinates 46.9° N, 4.8° E)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Messel Formation, MP 11 (48.6 - 40.4 Ma)

• "lower Middle Eocene, lower-most Geiseltalian, Mammal Paleogene level 11… .575 to .77 m above local stratigraphic marker Alpha."

• bed-level stratigraphic resolution

Environment/lithology: crater lake; lithified claystone

• "A drilling project conducted in 2001 showed that the former Lake Messel was a maar lake, which is a small deep lake that had its origin in an explosive volcanic eruption. The former lake basin had an original diameter of about 1.5 km, and a depth of about 300–400 m (Harms, 2002; Felder & Harms, 2004)."
• "The fossils are embedded in darkly coloured, finely laminated claystone sediments rich in organic substances (‘oil shale’), which were deposited in a meromictic lake (e.g. Goth, 1990)."

Size classes: macrofossils, mesofossils

Preservation: adpression, soft parts

Reposited in the SMF

Collection methods: quarrying,

Primary reference: R. K. Kinzelbach and H. Pohl. 1994. The Fossil Strepsiptera (Insecta: Strepsiptera). Annals of the Entomological Society of America 87(1):59-70 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 120798: authorized by Matthew Clapham, entered by Jered Karr on 21.11.2011, edited by Matthew Clapham and Jenna Sandlin

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Heterocera indet.9
SMF MeI 296
Insecta
 Hemiptera - Dictyopharidae
Wedelphus dichopteroides n. gen. n. sp.10
Wedelphus dichopteroides n. gen. n. sp.10 Szwedo and Wappler 2006 planthopper
SMF Mel 1512
 Hemiptera - Diaspididae
Aspidiotinae "indet. 1"11 Ferris 1942 armoured scale insect
FIS SM.B Me 3958
 Hemiptera -
? Auchenorrhyncha indet.9 true bug
SMF MeI 1211
Cicadomorpha indet.9 Evans 1946 true bug
SMF MeI 204, 295, 524, 566, 816, 1160, 1209, 1210
 Hemiptera - Cicadidae
Cicadidae indet.9 Latreille 1802 cicada
SMF MeI 97, 127, 739, 743, 744, 943, 944, 1046, 1213
 Hemiptera - Membracidae
? Membracidae indet.9 Rafinesque 1815 treehopper
SMF MeI 114
 Hemiptera - Pentatomidae
Pentatomidae indet.9 Leach 1815 stink bug
SMF MeI 649
 Hemiptera - Cydnidae
Cydnidae indet.9 Billberg 1820 burrowing bug
SMF MeI 79
 Hemiptera - Miridae
Miridae indet.9 Hahn 1833 plant bug
SMF MeI 132, 306, 522, 1095
 Blattodea -
Isoptera indet.9 termite
SMF MeI 952
 Blattodea - Blattoidae
Blattidae indet.9 Latreille 1810 cockroach
SMF MeI 1226
 Orthoptera -
Ensifera indet.9 Chopard 1920 grasshopper
SMF MeI 112, 188, 676, 684, 818
 Hymenoptera -
Apocrita indet.9 Gerstaecker 1867 wasp
SMF MeI 111, 203, 588, 589, 593, 595, 647, 680, 766
 Hymenoptera - Scoliidae
Scoliidae indet.9 Latreille 1802 wasp
SMF MeI 1500
 Hymenoptera -
Vespoidea indet.9 Latreille 1802 wasp
SMF MeI 427, 548, 651
 Hymenoptera - Tiphiidae
Tiphiidae indet.9 Leach 1815 wasp
SMF MeI 397, 513
 Hymenoptera - Formicidae
Formicidae indet.9 Latreille 1802 ant
SMF MeI 192, 1165, 57, 196, 198, 202, 549, 953, 1117, 113, 161, 428, 514, 515, 547, 599, 768, 1039, 1094, 1208
Formicinae indet.9 Latreille 1802 ant
SMF MeI 56, 122a-c, 468, 890, 923, 1204, 17, 314, 317, 320, 512, 561, 603, 654, 803, 812, 814, 909, 962, 963, 1108
Gesomyrmex breviceps n. sp.4 Dlussky et al. 2009 ant
SMF MeI 2305, 11054, 13151
Gesomyrmex curiosus n. sp.4 Dlussky et al. 2009 ant
SMF MeI 11953
Gesomyrmex pulcher n. sp.4 Dlussky et al. 2009 ant
SMF MeI 10999
Camponotus sp. Mayr 1861 ant
SMF-MeI 196 host of Stichotrema
Oecophylla longiceps n. sp.3 Dlussky et al. 2008 weaver ant
Casaleia eocenica n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 5565
Dolichoderinae indet.9 Forel 1878 ant
SMF MeI 321, 429, 1098, 1206
Myrmicinae indet.9 Lepeletier 1835 ant
SMF MeI 456, 517, 815, 929
Pseudectatomma eocenica n. gen. n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 400
Pseudectatomma striatula n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 5762, 8153, 10798
Ponerinae indet.5 Lepeletier de Saint-Fargeau 1835 ant
FIS MeI 202, 662, 7055, 7222, 7369, 7751, 7758, 8298, 8551, 10805, 11007, 12307, 12377, 12628, 12641, 13220, 13339, 13585, 13802
Cyrtopone striata n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 1392
Cyrtopone curiosa n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 6075
Cyrtopone microcephala n. gen. n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 2407
Cyrtopone elongata n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 10167
Protopone oculata n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 2334
? Protopone dubia n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 10142
Protopone sepulta n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 5329
Protopone germanica n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 10841, 12360
Protopone vetula n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 7679
Protopone magna n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 8790, 8789
Messelepone leptogenoides n. gen. n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 4808, 7198
Pachycondyla petrosa n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 12273
Pachycondyla petiolosa n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 1893, 1427
"Pachycondyla minuta n. sp." = Pachycondyla parvula5 Dlussky et al. 2015 ant
FIS MeI 10638, 10167, 12250, 10093, 13789, 14219
Pachycondyla lutzi n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 11958, 11481, 1361, 3553, 2078, 10526, 11679, 10564, 12975, 13507
Pachycondyla eocenica n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 10889, 8710
? Pachycondyla messeliana n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 4744
Cephalopone grandis n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 5361
Cephalopone potens n. gen. n. sp.5 Dlussky and Wedmann 2012 ant
FIS MeI 10860, 11830
Archimyrmex wedmannae n. sp.2 Dlussky 2012 bulldog ant
SMF Mel 2016
"Formicium giganteum n. sp." = Titanomyrma gigantea8 Lutz 1986 ant
"Formicium simillimum n. sp." = Titanomyrma simillima8 Lutz 1986 ant
26 in Lutz 1986; SMF MeI 1548a-c in Lutz 1990
 Hymenoptera - Apidae
"Anthophoridae indet." = Anthophorini9, Electrapis sp.13, Protobombus messelensis n. sp.12
"Anthophoridae indet." = Anthophorini9 Kirby 1837 bee
SMF MeI 222
Electrapis sp.13 Cockerell 1908 bee
FIS MeI 3300, 10890, 12151
Protobombus messelensis n. sp.12 Wappler and Engel 2003 bee
SMF MeI 6388
 Hymenoptera - Sphecidae
Sphecidae indet.9 Latreille 1802 digger wasp
SMF MeI 319, 1107
 Hymenoptera - Braconidae
Braconidae indet.9 Nees 1811 wasp
SMF MeI 1021, 1205
 Hymenoptera - Ichneumonidae
Ichneumonidae indet.9 Latreille 1802 ichneumon wasp
SMF MeI 195, 594
 Hymenoptera - Chalcididae
Chalcididae indet.9 Leach 1815 chalcid wasp
SMF MeI 1207
 Hymenoptera -
Proctotrupoidea indet.9 Latreille 1802 wasp
SMF MeI 596
 Dicondylia -
Coleoptera indet.9 Linnaeus 1758 beetle
SMF MeI 360, 435
 Coleoptera - Psephenidae
Eubrianacinae "indet. 1"14 Jacobson 1913 water penny beetle
"range from 5 m below to 5 m above marker alpha"
 Coleoptera - Hydrophilidae
Hydrophilina indet.6 Latreille 1802 water scavenger beetle
SMF MeI 3903, 8114, 11236
Hydrobiomorpha "sp. 1"6 Blackburn 1888 water scavenger beetle
SMF MeI 5085, 10530
Hydrobiomorpha eopalpalis6 Fikacek et al. 2010 water scavenger beetle
SMF MeI 459, 3893, 3923, 4204, 4272, 4445, 5040, 5577, 7331, 8590, 8801, 8944, 10662, 12304, 14099, 14101, 14614
Hydrophilus sp.6 Geoffroy 1762 water scavenger beetle
SMF MeI 806, 1524, 4648, 8936, 11127, 11804, 14052
Hydrochara sp.6 Berthold 1827 water scavenger beetle
SMF MeI 249, 1766, 5069, 6591, 7837
 Coleoptera - Staphylinidae
Staphylinidae indet.9 Latreille 1802 rove beetle
SMF MeI 744, 919
 Coleoptera - Cerambycidae
Lamiinae indet.9 Latreille 1825 longhorned beetle
SMF MeI 685, 969, 1015, 1016, 1027
 Coleoptera - Chrysomelidae
Donaciinae indet.9 Kirby 1837 leaf beetle
SMF MeI 181, 327
Sagrinae indet.9 Leach 1815 frog-legged beetle
SMF MeI 31, 153, 218, 259, 302, 388, 519, 551, 715, 731, 823, 827, 828, 843, 851, 884, 955, 959, 966, 1018, 1080, 1085, 1113, 1162, 1194
 Coleoptera - Brentidae
Brentidae indet.9 Billberg 1820 weevil
SMF MeI 248
 Coleoptera - Apionidae
Apionidae indet.9 Schoenherr 1823 weevil
SMF MeI 604
 Coleoptera - Lucanidae
Protognathinus spielbergi n. gen. n. sp.1 Chalumeau and Brochier 2001 stag beetle
Specimen illustrated in Feb. 2000 National Geographic magazine
aff. Dorcus sp.9 MacLeay 1819 stag beetle
SMF MeI 711
Lampriminae indet.9 MacLeay 1819 stag beetle
SMF MeI 3, 75, 177, 199, 313, 458, 520, 745, 811, 1072, 1131, 1174, 1549
 Coleoptera - Scarabaeidae
Onitini indet.9 Laporte 1840 scarab beetle
SMF MeI 172 (similar to Onitis)
Onthophagini indet.9 Burmeister 1846 scarab beetle
SMF MeI 646 (similar to Onthophagus)
 Coleoptera - Geotrupidae
Geotrupinae indet.9 Latreille 1802 earth-boring dung beetle
SMF MeI 660
 Coleoptera - Carabidae
Carabidae indet.9 Latreille 1802 ground beetle
SMF MeI 128, 307, 359, 372, 437, 477, 737, 858, 862
Scaritinae indet.9 Bonelli 1810 ground beetle
SMF MeI 126, 707, 1170
 Strepsiptera - Myrmecolacidae
Stichotrema sp. Hofeneder 1910 twisted-winged parasite
SMF-MeI 196b, c parasite of Camponotus
 Diptera -
Brachycera indet.9 Zetterstedt 1842 fly
SMF MeI 652
 Diptera - Syrphidae
Syrphidae indet.9 Latreille 1802 hover fly
SMF MeI 1338
 Diptera -
Tabanomorpha indet.9 Hennig 1948 fly
SMF MeI 1218 (Rhagionidae or Athericidae)
 Diptera - Tabanidae
Tabanidae indet.9 Latreille 1802 horse fly
SMF MeI 653
 Diptera -
Nematocera indet.9 Latreille 1825 fly
SMF MeI 677
Tipuloidea indet.9 Latreille 1802 crane fly
SMF MeI 1217 (Tipulidae or Limoniidae)
 Odonata - Dysagrionidae
Petrolestes messelensis n. sp.7 Garrouste and Nel 2015 damselfly
SMF MeI 11777
 Odonata -
Anisoptera indet.9 true dragonfly
SMF MeI 523
 Odonata - Aeshnidae
"Aeschnidae indet." = Aeshnidae9
"Aeschnidae indet." = Aeshnidae9 Leach 1815 darner dragonfly
SMF MeI 817
 Odonata - Corduliidae
cf. Corduliidae indet.9 Selys-Longchamps 1850 green-eyed skimmer
SMF MeI 1225
 Odonata - Pseudostenolestidae
Pseudostenolestes bechlyi n. gen. n. sp.7
Pseudostenolestes bechlyi n. gen. n. sp.7 Garrouste and Nel 2015 dragonfly
SMF MeI 14781, 13209, 14712, 14794