Cassville (Permian of the United States)

Where: Monongolia County, West Virginia (39.7° N, 80.1° W: paleocoordinates 5.1° S, 16.0° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Cassville Shales Formation, Asselian (298.9 - 295.5 Ma)

• "...roof shales of the Waynesburg coal, or the very highest of the beds of the upper productive coal series, in the nomenclature of the first Pennsylvania survey, or the beds termed Permo-carboniferous by Professor Fontaine." Scudder, 1879. Asselian (or possibly earliest Sakmarian) according to Schneider et al. (2013).

Environment/lithology: terrestrial; shale

Size class: macrofossils

Preservation: mold/impression

Collected by Mr. Fontaine, Mr. White & R.D. Lacoe

Primary reference: S. H. Scudder. 1879. Palaeozoic Cockroaches: A complete revision of the species of both worlds, with an essay toward their classification. Memoirs of the Boston Society of Natural History 3:23-134 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 122604: authorized by Matthew Clapham, entered by Jered Karr on 03.01.2012, edited by Amy Delelli and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Blattodea - Archimylacridae
Etoblattina sp.4, "Etoblattina arcta n. sp." = Scudderula arcta4, "Gerablattina ovata n. sp." = Liparoblatta ovata4, "Gerablattina radiata n. sp." = Liparoblatta radiata4, Gerablattina eversa n. sp.4, "Gerablattina permacra n. sp." = Acosmoblatta permacra4, "Etoblattina eakiniana n. sp." = Acosmoblatta eakiniana4, Gerablattina inculta n. sp.4
Etoblattina sp.4 Scudder 1879 cockroach
Coll Lacoe 2084 ab; hindwing
"Etoblattina arcta n. sp." = Scudderula arcta4 Scudder 1895 cockroach
Coll Lacoe 2118a
"Gerablattina ovata n. sp." = Liparoblatta ovata4 Scudder 1895 cockroach
Coll Lacoe 2150 ab
"Gerablattina radiata n. sp." = Liparoblatta radiata4 Scudder 1895 cockroach
Coll Lacoe 2147a
Gerablattina eversa n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2145a
"Gerablattina permacra n. sp." = Acosmoblatta permacra4 Scudder 1895 cockroach
USGS 2187(2188) collected by Scudder
"Etoblattina eakiniana n. sp." = Acosmoblatta eakiniana4 Scudder 1895 cockroach
Coll Lacoe 2111 ab
Gerablattina inculta n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2139a
 Blattodea -
Gerablattina perita n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2140 ab
Etoblattina exigua n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2107 ab
 Blattodea - Spiloblattinidae
Etoblattina aperta n. sp.4, "Etoblattina detecta n. sp." = Arrhythmoblatta detecta4, Arrhythmoblatta scudderiana n. sp.2, "Gerablattina balteata n. sp." = Sysciophlebia balteata, "Sysciophlebia balteata" = Sysciophlebia balteata3, "Gerablattina cassvici n. sp." = Sysciophlebia balteata4, "Gerablattina diversinervis n. sp." = Sysciophlebia balteata4, "Etoblattina occulta n. sp." = Sysciophlebia balteata4, "Etoblattina patiens n. sp." = Sysciophlebia patiens4, "Etoblattina recidiva n. sp." = Sysciophlebia recidiva4, "Etoblattina invisa n. sp." = Sysciophlebia invisa4, "Etoblattina defossa n. sp." = Dicladoblatta defossa4
Etoblattina aperta n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2066 ab
"Etoblattina detecta n. sp." = Arrhythmoblatta detecta4 Scudder 1895 cockroach
Coll Lacoe 2106 ab, c
Arrhythmoblatta scudderiana n. sp.2 Handlirsch 1906 cockroach
USNM 38894 (specimen of E. detecta in Scudder, 1895; pl 4, fig 13)
"Gerablattina balteata n. sp." = Sysciophlebia balteata Scudder 1879 cockroach
"Sysciophlebia balteata" = Sysciophlebia balteata3 Scudder 1879 cockroach
"Gerablattina cassvici n. sp." = Sysciophlebia balteata4 Scudder 1879 cockroach
Coll Lacoe 2142 ab, cd
"Gerablattina diversinervis n. sp." = Sysciophlebia balteata4 Scudder 1879 cockroach
Coll Lacoe 2141 ab
"Etoblattina occulta n. sp." = Sysciophlebia balteata4 Scudder 1879 cockroach
Coll Lacoe 2127 ab
"Etoblattina patiens n. sp." = Sysciophlebia patiens4 Scudder 1895 cockroach
Coll Lacoe 2104 ab
"Etoblattina recidiva n. sp." = Sysciophlebia recidiva4 Scudder 1895 cockroach
Coll Lacoe 2129 a
"Etoblattina invisa n. sp." = Sysciophlebia invisa4 Scudder 1895 cockroach
Coll Lacoe 2126 ab
"Etoblattina defossa n. sp." = Dicladoblatta defossa4 Scudder 1895 cockroach
Coll Lacoe 2128 ab
 Blattodea - Poroblattinidae
Poroblattina gratiosa n. sp.4, "Petrablattina hastata n. sp." = Poroblattina parvula4
Poroblattina gratiosa n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2134 ab (USNM 38166)
"Petrablattina hastata n. sp." = Poroblattina parvula4 Goldenberg 1869 cockroach
Coll Lacoe 2133 ab
 Dicondylia - Phyloblattidae
"Gerablattina deducta n. sp." = Phyloblatta deducta4, Phyloblatta scudderiana n. sp.1, ? Phyloblatta rebaptizata n. sp.1, "Etoblattina immolata n. sp." = Phyloblatta immolata4, "Etoblattina praedulcis n. sp." = Phyloblatta praedulcis4, Phyloblatta arcuata n. sp.1, Phyloblatta abbreviata n. sp.1, "Etoblattina expulsata n. sp." = Phyloblatta expulsata4, "Etoblattina angusta n. sp." = Phyloblatta angusta4, "Etoblattina imperfecta n. sp." = Phyloblatta imperfecta4, "Etoblattina gratiosa n. sp." = Phyloblatta gratiosa4, "Etoblattina exsecuta n. sp." = Phyloblatta exsecuta4, Phyloblatta debilis n. sp.1, "Etoblattina mactata n. sp." = Phyloblatta mactata4, "Etoblattina macilenta n. sp." = Phyloblatta macilenta4, "Etoblattina secreta n. sp." = Phyloblatta secreta4, "Etoblattina rogi n. sp." = Phyloblatta rogi4, "Etoblattina accubita n. sp." = Phyloblatta accubita4, "Etoblattina communis n. sp." = Phyloblatta communis4, "Etoblattina macerata n. sp." = Phyloblatta macerata4, Phyloblatta vulgata n. sp.1, "Gerablattina uniformis n. sp." = Phyloblatta uniformis4, "Etoblattina funeraria n. sp." = Phyloblatta funeraria4, "Gerablattina abdicata n. sp." = Phyloblatta abdicata4, "Etoblattina residua n. sp." = Phyloblatta residua4, Phyloblatta macroptera n. sp.1, ? Phyloblatta dimidiata n. sp.1, Phyloblatta elatior n. sp.1, "Gerablattina concinna n. sp." = Phyloblatta concinna4, Phyloblatta regularis n. sp.1, Phyloblatta dichotoma n. sp.1, Phyloblatta virginiana n. sp.1, "Etoblattina lata n. sp." = Phyloblatta lata4, Phyloblatta cassvilleana n. sp.1, "Etoblattina expugnata n. sp." = Phyloblatta expugnata4, "Etoblattina obatra n. sp." = Phyloblatta obatra4, Phyloblatta fracta n. sp.1, Phyloblatta mortua n. sp.1
"Gerablattina deducta n. sp." = Phyloblatta deducta4 Scudder 1895 winged insect
Coll Lacoe 2146 a (USNM 38063)
Phyloblatta scudderiana n. sp.1 Handlirsch 1906 winged insect
USNM 38904 (specimen of G. concinna in Scudder, 1895, pl 10, fig 5)
? Phyloblatta rebaptizata n. sp.1 Handlirsch 1906 winged insect
USNM 38106 (specimen of P. gratiosa in Scudder, 1895, pl 11, fig 13)
"Etoblattina immolata n. sp." = Phyloblatta immolata4 Scudder 1895 winged insect
Coll Lacoe 2115 ab, cd (USNM 38079; other specimen assigned to P. debilis Handlirsch)
"Etoblattina praedulcis n. sp." = Phyloblatta praedulcis4 Scudder 1895 winged insect
Coll Lacoe 2119 ab (USNM 38165)
Phyloblatta arcuata n. sp.1 Handlirsch 1906 winged insect
USNM 38898 (specimen of E. communis in Scudder, 1895, pl 7, fig 13)
Phyloblatta abbreviata n. sp.1 Handlirsch 1906 winged insect
USNM 38588
"Etoblattina expulsata n. sp." = Phyloblatta expulsata4 Scudder 1895 winged insect
Coll Lacoe 2113 ab, cd, ef (USNM 38178; Handlirsch assigns 1 to P. vulgata)
"Etoblattina angusta n. sp." = Phyloblatta angusta4 Scudder 1895 winged insect
Coll Lacoe 2080 ab (USNM 38185)
"Etoblattina imperfecta n. sp." = Phyloblatta imperfecta4 Scudder 1895 winged insect
Coll Lacoe 2124 ab (USNM 38083)
"Etoblattina gratiosa n. sp." = Phyloblatta gratiosa4 Scudder 1895 winged insect
Coll Lacoe 2114 ab
"Etoblattina exsecuta n. sp." = Phyloblatta exsecuta4 Scudder 1895 winged insect
Coll Lacoe 2120 ab, cd, e (USNM 38180; others assigned to new species by Handlirsch)
Phyloblatta debilis n. sp.1 Handlirsch 1906 winged insect
USNM 38094 (specimen of E. immolata in Scudder, 1895, pl. 7, fig 8)
"Etoblattina mactata n. sp." = Phyloblatta mactata4 Scudder 1895 winged insect
Coll Lacoe 2116a (USNM 38081)
"Etoblattina macilenta n. sp." = Phyloblatta macilenta4 Scudder 1895 winged insect
Coll Lacoe 2121 ab (USNM 38163)
"Etoblattina secreta n. sp." = Phyloblatta secreta4 Scudder 1895 winged insect
Coll Lacoe 2125 ab, cd, ef, g (USNM 38167; Handlirsch assigns 1 to P. virginiana)
"Etoblattina rogi n. sp." = Phyloblatta rogi4 Scudder 1895 winged insect
Coll Lacoe 2122 a, bc (USNM 38088)
"Etoblattina accubita n. sp." = Phyloblatta accubita4 Scudder 1895 winged insect
Coll Lacoe 2112a (USNM 38169)
"Etoblattina communis n. sp." = Phyloblatta communis4 Scudder 1895 winged insect
Coll Lacoe 2187 ab, cd, e, f, g, h, i, j, kl, mn (USNM 38188; Handlirsch assigns 6 specimens to new species)
"Etoblattina macerata n. sp." = Phyloblatta macerata4 Scudder 1895 winged insect
Coll Lacoe 2081 ab, c (USNM 38183)
Phyloblatta vulgata n. sp.1 Handlirsch 1906 winged insect
USNM 38901 (specimen of E. expulsata in Scudder, 1895, pl. 9, fig 4)
"Gerablattina uniformis n. sp." = Phyloblatta uniformis4 Scudder 1895 winged insect
Coll Lacoe 2082 ab, cd, e, fg (USNM 38177; Handlirsch assigns 3 to other species)
"Etoblattina funeraria n. sp." = Phyloblatta funeraria4 Scudder 1895 winged insect
Coll Lacoe 2108 ab, c (USNM 38078)
"Gerablattina abdicata n. sp." = Phyloblatta abdicata4 Scudder 1895 winged insect
Coll Lacoe 2143a (USNM 38065)
"Etoblattina residua n. sp." = Phyloblatta residua4 Scudder 1895 winged insect
Coll Lacoe 2131a (USNM 38179)
Phyloblatta macroptera n. sp.1 Handlirsch 1906 winged insect
USNM 38891 (specimen of E. communis in Scudder, 1895, pl 7, fig 17)
? Phyloblatta dimidiata n. sp.1 Handlirsch 1906 winged insect
USNM 38905 (specimen of G. uniformis in Scudder, 1895, pl 10, fig 11))
Phyloblatta elatior n. sp.1 Handlirsch 1906 winged insect
USNM 38087 (specimen of E. communis in Scudder, 1895, pl 7, fig 14)
"Gerablattina concinna n. sp." = Phyloblatta concinna4 Scudder 1895 winged insect
Coll Lacoe 2077 ab, cd (USNM 38172; Handlirsch assigns 1 to P. scudderiana)
Phyloblatta regularis n. sp.1 Handlirsch 1906 winged insect
USNM 38893 (specimen of G. uniformis in Scudder, 1895, pl. 10, fig 9)
Phyloblatta dichotoma n. sp.1 Handlirsch 1906 winged insect
USNM 38896 (specimen of E. communis in Scudder, 1895, pl 7, fig 11)
Phyloblatta virginiana n. sp.1 Handlirsch 1906 winged insect
USNM 38902 (specimen of E. secreta in Scudder, 1895, pl. 9, fig 7)
"Etoblattina lata n. sp." = Phyloblatta lata4 Scudder 1895 winged insect
Coll Lacoe 2099a (USNM 38200)
Phyloblatta cassvilleana n. sp.1 Handlirsch 1906 winged insect
USNM 38892 (specimen of G. uniformis in Scudder, 1895, pl 10, fig 10)
"Etoblattina expugnata n. sp." = Phyloblatta expugnata4 Scudder 1895 winged insect
Coll Lacoe 2085 ab (USNM 38193)
"Etoblattina obatra n. sp." = Phyloblatta obatra4 Scudder 1895 winged insect
Coll Lacoe 2123 ab (USNM 38087)
Phyloblatta fracta n. sp.1 Handlirsch 1906 winged insect
USNM 38897 (specimen of E. communis in Scudder, 1895, pl 7, fig 12)
Phyloblatta mortua n. sp.1 Handlirsch 1906 winged insect
USNM 38899 (specimen of E. communis in Scudder, 1895, pl 7, fig 15)
 Dicondylia - Schizoblattidae
"Poroblattina fossa n. sp." = Apempherus fossus4, "Poroblattina complexinervis n. sp." = Apempherus complexinervis4
"Poroblattina fossa n. sp." = Apempherus fossus4 Scudder 1895 winged insect
Coll Lacoe 2083 ab, c
"Poroblattina complexinervis n. sp." = Apempherus complexinervis4 Scudder 1895 winged insect
Coll Lacoe 2135 ab
 Dicondylia - Bradyblattidae
"Etoblattina mucronata n. sp." = Bradyblatta mucronata4, "Etoblattina sagittaria n. sp." = Bradyblatta sagittaria4, "Etoblattina mediana n. sp." = Bradyblatta sagittaria4
"Etoblattina mucronata n. sp." = Bradyblatta mucronata4 Scudder 1895 winged insect
Coll lacoe 2105 ab (USNM 38199)
"Etoblattina sagittaria n. sp." = Bradyblatta sagittaria4 Scudder 1895 winged insect
Coll Lacoe 2100a, b, c, d
"Etoblattina mediana n. sp." = Bradyblatta sagittaria4 Scudder 1895 winged insect
Coll Lacoe 2101 ab (USNM 38198)
 Dicondylia - Triassoblattidae
"Etoblattina expuncta n. sp." = Pareinoblatta expuncta4
"Etoblattina expuncta n. sp." = Pareinoblatta expuncta4 Scudder 1895 winged insect
Coll Lacoe 2109 ab
 Dicondylia - Petrablattinidae
"Anthracoblattina virginiensis n. sp." = Penetoblatta virginiensis4, "Gerablattina rotundata n. sp." = Penetoblatta rotundata4, "Etoblattina debilis n. sp." = Symphyoblatta debilis4, "Etoblattina ovata n. sp." = Petrablattina ovata4, "Gerablattina permanenta n. sp." = Amoeboblatta permanenta4, "Gerablattina lata n. sp." = Amblyblatta lata4
"Anthracoblattina virginiensis n. sp." = Penetoblatta virginiensis4 Scudder 1895 winged insect
Coll Lacoe 2138 ab
"Gerablattina rotundata n. sp." = Penetoblatta rotundata4 Scudder 1895 winged insect
Coll Lacoe 2149a
"Etoblattina debilis n. sp." = Symphyoblatta debilis4 Scudder 1895 winged insect
Coll Lacoe 2103 ab, cd, e
"Etoblattina ovata n. sp." = Petrablattina ovata4 Scudder 1895 winged insect
Coll Lacoe 2102a (USNM 38201)
"Gerablattina permanenta n. sp." = Amoeboblatta permanenta4 Scudder 1895 winged insect
Coll Lacoe 2144 ab
"Gerablattina lata n. sp." = Amblyblatta lata4 Scudder 1895 winged insect
Coll Lacoe 2148a