Cassville (Permian of the United States)

Where: Monongolia County, West Virginia (39.7° N, 80.1° W: paleocoordinates 5.1° S, 16.0° W)

• coordinate based on nearby landmark

• outcrop-level geographic resolution

When: Cassville Shales Formation, Asselian (298.9 - 295.5 Ma)

• "...roof shales of the Waynesburg coal, or the very highest of the beds of the upper productive coal series, in the nomenclature of the first Pennsylvania survey, or the beds termed Permo-carboniferous by Professor Fontaine." Scudder, 1879. Asselian (or possibly earliest Sakmarian) according to Schneider et al. (2013).

Environment/lithology: terrestrial; shale

Size class: macrofossils

Preservation: mold/impression

Collected by Mr. Fontaine, Mr. White & R.D. Lacoe

Primary reference: S. H. Scudder. 1879. Palaeozoic Cockroaches: A complete revision of the species of both worlds, with an essay toward their classification. Memoirs of the Boston Society of Natural History 3:23-134 [M. Clapham/J. Karr]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 122604: authorized by Matthew Clapham, entered by Jered Karr on 03.01.2012, edited by Amy Delelli and Matthew Clapham

Creative Commons license: CC BY (attribution)

Taxonomic list

Insecta
 Dicondylia - Petrablattinidae
"Etoblattina ovata n. sp." = Petrablattina ovata4, "Gerablattina rotundata n. sp." = Penetoblatta rotundata4, "Anthracoblattina virginiensis n. sp." = Penetoblatta virginiensis4, "Gerablattina lata n. sp." = Amblyblatta lata4, "Etoblattina debilis n. sp." = Symphyoblatta debilis4, "Gerablattina permanenta n. sp." = Amoeboblatta permanenta4
"Etoblattina ovata n. sp." = Petrablattina ovata4 Scudder 1895 winged insect
Coll Lacoe 2102a (USNM 38201)
"Gerablattina rotundata n. sp." = Penetoblatta rotundata4 Scudder 1895 winged insect
Coll Lacoe 2149a
"Anthracoblattina virginiensis n. sp." = Penetoblatta virginiensis4 Scudder 1895 winged insect
Coll Lacoe 2138 ab
"Gerablattina lata n. sp." = Amblyblatta lata4 Scudder 1895 winged insect
Coll Lacoe 2148a
"Etoblattina debilis n. sp." = Symphyoblatta debilis4 Scudder 1895 winged insect
Coll Lacoe 2103 ab, cd, e
"Gerablattina permanenta n. sp." = Amoeboblatta permanenta4 Scudder 1895 winged insect
Coll Lacoe 2144 ab
 Dicondylia - Schizoblattidae
"Poroblattina fossa n. sp." = Apempherus fossus4, "Poroblattina complexinervis n. sp." = Apempherus complexinervis4
"Poroblattina fossa n. sp." = Apempherus fossus4 Scudder 1895 winged insect
Coll Lacoe 2083 ab, c
"Poroblattina complexinervis n. sp." = Apempherus complexinervis4 Scudder 1895 winged insect
Coll Lacoe 2135 ab
 Dicondylia - Bradyblattidae
"Etoblattina mucronata n. sp." = Bradyblatta mucronata4, "Etoblattina sagittaria n. sp." = Bradyblatta sagittaria4, "Etoblattina mediana n. sp." = Bradyblatta sagittaria4
"Etoblattina mucronata n. sp." = Bradyblatta mucronata4 Scudder 1895 winged insect
Coll lacoe 2105 ab (USNM 38199)
"Etoblattina sagittaria n. sp." = Bradyblatta sagittaria4 Scudder 1895 winged insect
Coll Lacoe 2100a, b, c, d
"Etoblattina mediana n. sp." = Bradyblatta sagittaria4 Scudder 1895 winged insect
Coll Lacoe 2101 ab (USNM 38198)
 Dicondylia - Spiloblattinidae
"Etoblattina defossa n. sp." = Dicladoblatta defossa4, Arrhythmoblatta scudderiana n. sp.2, "Etoblattina detecta n. sp." = Arrhythmoblatta detecta4, Etoblattina aperta n. sp.4, "Etoblattina patiens n. sp." = Sysciophlebia patiens4, "Etoblattina recidiva n. sp." = Sysciophlebia recidiva4, "Etoblattina invisa n. sp." = Sysciophlebia invisa4, "Gerablattina balteata n. sp." = Sysciophlebia balteata, "Sysciophlebia balteata" = Sysciophlebia balteata3, "Gerablattina cassvici n. sp." = Sysciophlebia balteata4, "Etoblattina occulta n. sp." = Sysciophlebia balteata4, "Gerablattina diversinervis n. sp." = Sysciophlebia balteata4
"Etoblattina defossa n. sp." = Dicladoblatta defossa4 Scudder 1895 winged insect
Coll Lacoe 2128 ab
Arrhythmoblatta scudderiana n. sp.2 Handlirsch 1906 winged insect
USNM 38894 (specimen of E. detecta in Scudder, 1895; pl 4, fig 13)
"Etoblattina detecta n. sp." = Arrhythmoblatta detecta4 Scudder 1895 winged insect
Coll Lacoe 2106 ab, c
Etoblattina aperta n. sp.4 Scudder 1895 winged insect
Coll Lacoe 2066 ab
"Etoblattina patiens n. sp." = Sysciophlebia patiens4 Scudder 1895 winged insect
Coll Lacoe 2104 ab
"Etoblattina recidiva n. sp." = Sysciophlebia recidiva4 Scudder 1895 winged insect
Coll Lacoe 2129 a
"Etoblattina invisa n. sp." = Sysciophlebia invisa4 Scudder 1895 winged insect
Coll Lacoe 2126 ab
"Gerablattina balteata n. sp." = Sysciophlebia balteata Scudder 1879 winged insect
"Sysciophlebia balteata" = Sysciophlebia balteata3 Scudder 1879 winged insect
"Gerablattina cassvici n. sp." = Sysciophlebia balteata4 Scudder 1879 winged insect
Coll Lacoe 2142 ab, cd
"Etoblattina occulta n. sp." = Sysciophlebia balteata4 Scudder 1879 winged insect
Coll Lacoe 2127 ab
"Gerablattina diversinervis n. sp." = Sysciophlebia balteata4 Scudder 1879 winged insect
Coll Lacoe 2141 ab
 Blattodea -
Etoblattina exigua n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2107 ab
Gerablattina perita n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2140 ab
 Blattodea - Mesoblattinidae
"Etoblattina expuncta n. sp." = Pareinoblatta expuncta4
"Etoblattina expuncta n. sp." = Pareinoblatta expuncta4 Scudder 1895 cockroach
Coll Lacoe 2109 ab
 Blattodea - Archimylacridae
"Gerablattina radiata n. sp." = Liparoblatta radiata4, "Gerablattina ovata n. sp." = Liparoblatta ovata4, Gerablattina eversa n. sp.4, "Etoblattina arcta n. sp." = Scudderula arcta4, Gerablattina inculta n. sp.4, "Gerablattina permacra n. sp." = Acosmoblatta permacra4, "Etoblattina eakiniana n. sp." = Acosmoblatta eakiniana4, Etoblattina sp.4
"Gerablattina radiata n. sp." = Liparoblatta radiata4 Scudder 1895 cockroach
Coll Lacoe 2147a
"Gerablattina ovata n. sp." = Liparoblatta ovata4 Scudder 1895 cockroach
Coll Lacoe 2150 ab
Gerablattina eversa n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2145a
"Etoblattina arcta n. sp." = Scudderula arcta4 Scudder 1895 cockroach
Coll Lacoe 2118a
Gerablattina inculta n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2139a
"Gerablattina permacra n. sp." = Acosmoblatta permacra4 Scudder 1895 cockroach
USGS 2187(2188) collected by Scudder
"Etoblattina eakiniana n. sp." = Acosmoblatta eakiniana4 Scudder 1895 cockroach
Coll Lacoe 2111 ab
Etoblattina sp.4 Scudder 1879 cockroach
Coll Lacoe 2084 ab; hindwing
 Blattodea - Poroblattinidae
"Petrablattina hastata n. sp." = Poroblattina parvula4
"Petrablattina hastata n. sp." = Poroblattina parvula4 Goldenberg 1869 cockroach
Coll Lacoe 2133 ab
 Blattodea -
Poroblattina gratiosa n. sp.4 Scudder 1895 cockroach
Coll Lacoe 2134 ab (USNM 38166)
 Dicondylia - Phyloblattidae
"Etoblattina secreta n. sp." = Phyloblatta secreta4, "Etoblattina expulsata n. sp." = Phyloblatta expulsata4, "Gerablattina concinna n. sp." = Phyloblatta concinna4, Phyloblatta cassvilleana n. sp.1, Phyloblatta vulgata n. sp.1, "Etoblattina residua n. sp." = Phyloblatta residua4, "Etoblattina rogi n. sp." = Phyloblatta rogi4, "Etoblattina mactata n. sp." = Phyloblatta mactata4, Phyloblatta fracta n. sp.1, Phyloblatta debilis n. sp.1, "Etoblattina praedulcis n. sp." = Phyloblatta praedulcis4, Phyloblatta macroptera n. sp.1, ? Phyloblatta dimidiata n. sp.1, Phyloblatta mortua n. sp.1, "Etoblattina exsecuta n. sp." = Phyloblatta exsecuta4, ? Phyloblatta rebaptizata n. sp.1, Phyloblatta scudderiana n. sp.1, "Etoblattina lata n. sp." = Phyloblatta lata4, "Etoblattina angusta n. sp." = Phyloblatta angusta4, "Etoblattina expugnata n. sp." = Phyloblatta expugnata4, "Etoblattina gratiosa n. sp." = Phyloblatta gratiosa4, Phyloblatta regularis n. sp.1, "Gerablattina abdicata n. sp." = Phyloblatta abdicata4, Phyloblatta abbreviata n. sp.1, "Etoblattina imperfecta n. sp." = Phyloblatta imperfecta4, Phyloblatta elatior n. sp.1, "Etoblattina macerata n. sp." = Phyloblatta macerata4, "Etoblattina macilenta n. sp." = Phyloblatta macilenta4, "Gerablattina uniformis n. sp." = Phyloblatta uniformis4, "Etoblattina funeraria n. sp." = Phyloblatta funeraria4, "Etoblattina communis n. sp." = Phyloblatta communis4, Phyloblatta arcuata n. sp.1, "Gerablattina deducta n. sp." = Phyloblatta deducta4, "Etoblattina accubita n. sp." = Phyloblatta accubita4, Phyloblatta virginiana n. sp.1, "Etoblattina obatra n. sp." = Phyloblatta obatra4, "Etoblattina immolata n. sp." = Phyloblatta immolata4, Phyloblatta dichotoma n. sp.1
"Etoblattina secreta n. sp." = Phyloblatta secreta4 Scudder 1895 winged insect
Coll Lacoe 2125 ab, cd, ef, g (USNM 38167; Handlirsch assigns 1 to P. virginiana)
"Etoblattina expulsata n. sp." = Phyloblatta expulsata4 Scudder 1895 winged insect
Coll Lacoe 2113 ab, cd, ef (USNM 38178; Handlirsch assigns 1 to P. vulgata)
"Gerablattina concinna n. sp." = Phyloblatta concinna4 Scudder 1895 winged insect
Coll Lacoe 2077 ab, cd (USNM 38172; Handlirsch assigns 1 to P. scudderiana)
Phyloblatta cassvilleana n. sp.1 Handlirsch 1906 winged insect
USNM 38892 (specimen of G. uniformis in Scudder, 1895, pl 10, fig 10)
Phyloblatta vulgata n. sp.1 Handlirsch 1906 winged insect
USNM 38901 (specimen of E. expulsata in Scudder, 1895, pl. 9, fig 4)
"Etoblattina residua n. sp." = Phyloblatta residua4 Scudder 1895 winged insect
Coll Lacoe 2131a (USNM 38179)
"Etoblattina rogi n. sp." = Phyloblatta rogi4 Scudder 1895 winged insect
Coll Lacoe 2122 a, bc (USNM 38088)
"Etoblattina mactata n. sp." = Phyloblatta mactata4 Scudder 1895 winged insect
Coll Lacoe 2116a (USNM 38081)
Phyloblatta fracta n. sp.1 Handlirsch 1906 winged insect
USNM 38897 (specimen of E. communis in Scudder, 1895, pl 7, fig 12)
Phyloblatta debilis n. sp.1 Handlirsch 1906 winged insect
USNM 38094 (specimen of E. immolata in Scudder, 1895, pl. 7, fig 8)
"Etoblattina praedulcis n. sp." = Phyloblatta praedulcis4 Scudder 1895 winged insect
Coll Lacoe 2119 ab (USNM 38165)
Phyloblatta macroptera n. sp.1 Handlirsch 1906 winged insect
USNM 38891 (specimen of E. communis in Scudder, 1895, pl 7, fig 17)
? Phyloblatta dimidiata n. sp.1 Handlirsch 1906 winged insect
USNM 38905 (specimen of G. uniformis in Scudder, 1895, pl 10, fig 11))
Phyloblatta mortua n. sp.1 Handlirsch 1906 winged insect
USNM 38899 (specimen of E. communis in Scudder, 1895, pl 7, fig 15)
"Etoblattina exsecuta n. sp." = Phyloblatta exsecuta4 Scudder 1895 winged insect
Coll Lacoe 2120 ab, cd, e (USNM 38180; others assigned to new species by Handlirsch)
? Phyloblatta rebaptizata n. sp.1 Handlirsch 1906 winged insect
USNM 38106 (specimen of P. gratiosa in Scudder, 1895, pl 11, fig 13)
Phyloblatta scudderiana n. sp.1 Handlirsch 1906 winged insect
USNM 38904 (specimen of G. concinna in Scudder, 1895, pl 10, fig 5)
"Etoblattina lata n. sp." = Phyloblatta lata4 Scudder 1895 winged insect
Coll Lacoe 2099a (USNM 38200)
"Etoblattina angusta n. sp." = Phyloblatta angusta4 Scudder 1895 winged insect
Coll Lacoe 2080 ab (USNM 38185)
"Etoblattina expugnata n. sp." = Phyloblatta expugnata4 Scudder 1895 winged insect
Coll Lacoe 2085 ab (USNM 38193)
"Etoblattina gratiosa n. sp." = Phyloblatta gratiosa4 Scudder 1895 winged insect
Coll Lacoe 2114 ab
Phyloblatta regularis n. sp.1 Handlirsch 1906 winged insect
USNM 38893 (specimen of G. uniformis in Scudder, 1895, pl. 10, fig 9)
"Gerablattina abdicata n. sp." = Phyloblatta abdicata4 Scudder 1895 winged insect
Coll Lacoe 2143a (USNM 38065)
Phyloblatta abbreviata n. sp.1 Handlirsch 1906 winged insect
USNM 38588
"Etoblattina imperfecta n. sp." = Phyloblatta imperfecta4 Scudder 1895 winged insect
Coll Lacoe 2124 ab (USNM 38083)
Phyloblatta elatior n. sp.1 Handlirsch 1906 winged insect
USNM 38087 (specimen of E. communis in Scudder, 1895, pl 7, fig 14)
"Etoblattina macerata n. sp." = Phyloblatta macerata4 Scudder 1895 winged insect
Coll Lacoe 2081 ab, c (USNM 38183)
"Etoblattina macilenta n. sp." = Phyloblatta macilenta4 Scudder 1895 winged insect
Coll Lacoe 2121 ab (USNM 38163)
"Gerablattina uniformis n. sp." = Phyloblatta uniformis4 Scudder 1895 winged insect
Coll Lacoe 2082 ab, cd, e, fg (USNM 38177; Handlirsch assigns 3 to other species)
"Etoblattina funeraria n. sp." = Phyloblatta funeraria4 Scudder 1895 winged insect
Coll Lacoe 2108 ab, c (USNM 38078)
"Etoblattina communis n. sp." = Phyloblatta communis4 Scudder 1895 winged insect
Coll Lacoe 2187 ab, cd, e, f, g, h, i, j, kl, mn (USNM 38188; Handlirsch assigns 6 specimens to new species)
Phyloblatta arcuata n. sp.1 Handlirsch 1906 winged insect
USNM 38898 (specimen of E. communis in Scudder, 1895, pl 7, fig 13)
"Gerablattina deducta n. sp." = Phyloblatta deducta4 Scudder 1895 winged insect
Coll Lacoe 2146 a (USNM 38063)
"Etoblattina accubita n. sp." = Phyloblatta accubita4 Scudder 1895 winged insect
Coll Lacoe 2112a (USNM 38169)
Phyloblatta virginiana n. sp.1 Handlirsch 1906 winged insect
USNM 38902 (specimen of E. secreta in Scudder, 1895, pl. 9, fig 7)
"Etoblattina obatra n. sp." = Phyloblatta obatra4 Scudder 1895 winged insect
Coll Lacoe 2123 ab (USNM 38087)
"Etoblattina immolata n. sp." = Phyloblatta immolata4 Scudder 1895 winged insect
Coll Lacoe 2115 ab, cd (USNM 38079; other specimen assigned to P. debilis Handlirsch)
Phyloblatta dichotoma n. sp.1 Handlirsch 1906 winged insect
USNM 38896 (specimen of E. communis in Scudder, 1895, pl 7, fig 11)