PBDB Collection

Florissant (USNM Collection) (Eocene of the United States)

Where: Teller County, Colorado (38.9° N, 105.3° W: paleocoordinates 40.4° N, 97.0° W)

• coordinate based on nearby landmark

• small collection-level geographic resolution

When: Florissant Formation, Chadronian (37.2 - 33.9 Ma)

Environment/lithology: lacustrine - large; lithified, diatomaceous, tuffaceous shale

• "shales"

Size class: mesofossils

Preservation: adpression

Reposited in the USNM

Primary reference: T. D. A. Cockerell. 1913. Some fossil insects from Florissant, Colorado. Proceedings of the United States National Museum 44:341-346 [C. Labandeira/C. Labandeira/M. Clapham]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 124624: authorized by Matthew Clapham, entered by Matthew Clapham on 20.02.2012, edited by April Kinchloe, Amy Delelli and Jered Karr

Creative Commons license: CC BY (attribution)

Taxonomic list

Show authors, comments, and common names

Arachnida
Araneae - Linyphiidae	Linyphia pachygnathoides n. sp.⁵ Linyphia pachygnathoides n. sp.⁵ Petrunkevitch 1922 money spider/sheet weaver USNM 38124
Opiliones - Sclerosomatidae	"Phalangium lacoei n. sp." = Amauropilio lacoei⁵ "Phalangium lacoei n. sp." = Amauropilio lacoei⁵ Petrunkevitch 1922 harvestman USNM Lacoe 38124
Insecta
Diptera - Chironomidae	Diamesa extincta⁴ Diamesa extincta⁴ Melander 1949 non biting midge USNM 112580, 2 (Lacoe coll)
Diptera - Cylindrotomidae	"Cyttaromyia clathrata" = Cyttaromyia princetoniana⁴ "Cyttaromyia clathrata" = Cyttaromyia princetoniana⁴ Scudder 1894 long bodied crane fly Lacoe coll
Diptera - Tipulidae	Tipula limi⁴, Tipulidea distincta⁴ Tipula limi⁴ Scudder 1894 crane fly Lacoe coll Tipulidea distincta⁴ Scudder 1894 crane fly Lacoe coll
Diptera - Mycetophilidae	Boletina hypogaea n. sp.⁴ Boletina hypogaea n. sp.⁴ Melander 1949 fungus gnat USNM 112571 (Lacoe coll)
Diptera - Bibionidae	Plecia explanata n. sp.², "Hesperinus melanderi" = Plecia melanderi⁴, "Hesperinus axelianus" = Plecia axeliana⁴, Plecia orycta n. sp.⁴, Plecia tessella n. sp.⁴, Bibio jamesi⁴, Bibio capnodes n. sp.⁴, Bibio excurvatus n. sp.⁴, Bibio vulcanius n. sp.⁴, Bibio vetus⁴, Bibio cockerelli⁴, Bibio wickhami², "Hesperinus immutabilis n. sp." = Penthetria immutabilis⁴, Penthetria longifurca n. sp.⁴ Plecia explanata n. sp.² Cockerell 1917 love bug USNM 62536 "Hesperinus melanderi" = Plecia melanderi⁴ Cockerell 1911 love bug Lacoe coll (study number 246) "Hesperinus axelianus" = Plecia axeliana⁴ Cockerell 1915 love bug Lacoe coll Plecia orycta n. sp.⁴ Melander 1949 love bug USNM 112569 (Lacoe coll) Plecia tessella n. sp.⁴ Melander 1949 love bug USNM 112562 (Lacoe coll) Bibio jamesi⁴ Melander 1949 march fly USNM 112576, 9 (Lacoe coll) Bibio capnodes n. sp.⁴ Melander 1949 march fly USNM 112565 (Lacoe coll) Bibio excurvatus n. sp.⁴ Melander 1949 march fly USNM 112550 (Lacoe coll) Bibio vulcanius n. sp.⁴ Melander 1949 march fly USNM 112564, 572 (Lacoe coll) Bibio vetus⁴ James 1937 march fly Lacoe coll, study number 232 Bibio cockerelli⁴ James 1937 march fly Lacoe coll Bibio wickhami² Cockerell 1915 march fly 12 in Lacoe coll (three lots) from Melander, 1949 "Hesperinus immutabilis n. sp." = Penthetria immutabilis⁴ Melander 1949 march fly USNM 112554, 588 (Lacoe coll) Penthetria longifurca n. sp.⁴ Melander 1949 march fly USNM 112551 (Lacoe coll)
Diptera - Asilidae	Asilus curculionis n. sp.³, Leptogaster prior n. sp.³ Asilus curculionis n. sp.³ Melander 1946 robber fly USNM 112558 (Lacoe coll) Leptogaster prior n. sp.³ Melander 1946 grass fly USNM 112560 (Lacoe coll)
Diptera - Bombyliidae	Apolysis magister⁴, "Protophthiria atra n. sp." = Tithonomyia atra⁴, "Acreotrichites scopulicornis n. gen. n. sp." = Desmatomyia scopulicornis², "Melanderella testea n. sp." = Dolichomyia testea⁴ Apolysis magister⁴ Melander 1946 bee fly USNM 112573 (Lacoe coll) "Protophthiria atra n. sp." = Tithonomyia atra⁴ Melander 1949 bee fly USNM 112552 (Lacoe coll) "Acreotrichites scopulicornis n. gen. n. sp." = Desmatomyia scopulicornis² Cockerell 1917 bee fly USNM 62537 "Melanderella testea n. sp." = Dolichomyia testea⁴ Melander 1949 bee fly USNM 112553 (Lacoe coll)
Diptera - Lauxaniidae	Sapromyza veterana n. sp.⁴ Sapromyza veterana n. sp.⁴ Melander 1949 fly USNM 112557 (Lacoe coll)
Diptera - Ulidiidae	Melieria atavina n. sp.² Melieria atavina n. sp.² Cockerell 1917 picture-winged fly USNM 62540
Diptera - Richardiidae	Urortalis caudatus n. sp.² Urortalis caudatus n. sp.² Cockerell 1917 fly USNM 62539
Diptera - Phoridae	"Phora cockerelli" = Spiniphora cockerelli⁴ "Phora cockerelli" = Spiniphora cockerelli⁴ Brues 1908 scuttle fly Lacoe coll, study number 154
Diptera - Hybotidae	Tachypeza primitiva n. sp.⁴ Tachypeza primitiva n. sp.⁴ Melander 1949 dance fly USNM 112566 (Lacoe coll)
Diptera - Empididae	Rhamphomyia inanimata n. sp.⁴, Rhamphomyia infernalis n. sp.⁴, Rhamphomyia fossa n. sp.⁴, Rhamphomyia craterae n. sp.⁴, Rhamphomyia hypolitha n. sp.², Empis infossa n. sp.⁴ Rhamphomyia inanimata n. sp.⁴ Melander 1949 dagger fly USNM 112570(586) (Lacoe coll) Rhamphomyia infernalis n. sp.⁴ Melander 1949 dagger fly USNM 112556 (Lacoe coll) Rhamphomyia fossa n. sp.⁴ Melander 1949 dagger fly USNM 112567, 583 (Lacoe coll) Rhamphomyia craterae n. sp.⁴ Melander 1949 dagger fly USNM 112561 (Lacoe coll) Rhamphomyia hypolitha n. sp.² Cockerell 1917 dagger fly USNM 62538 Empis infossa n. sp.⁴ Melander 1949 dagger fly USNM 112625 (Lacoe coll)
Diptera - Xylomyidae	Solva (Xylomya) inornata n. sp.⁴ Solva (Xylomya) inornata n. sp.⁴ Melander 1949 fly USNM 112568 (Lacoe coll)
Diptera - Nemestrinidae	Nemestrinidae indet.⁴ Nemestrinidae indet.⁴ Macquart 1834 tangle-veined fly Lacoe coll
Diptera - Rhagionidae	Rhagio wheeleri⁴, Rhagio fossitius n. sp.⁴ Rhagio wheeleri⁴ Melander 1949 snipe fly USNM 112574 (Lacoe coll) Rhagio fossitius n. sp.⁴ Melander 1949 snipe fly USNM 112626 (Lacoe coll)
Diptera -	Anthomyia persepulta n. sp.² Anthomyia persepulta n. sp.² Cockerell 1917 fly USNM 62541
Diptera - Scatopsidae	Reichertella fasciata n. sp.⁴ Reichertella fasciata n. sp.⁴ Melander 1949 minute black scavenger fly USNM 112563 (Lacoe coll)
Mantodea - Mantidae	Eobruneria tessellata n. gen. n. sp. Eobruneria tessellata n. gen. n. sp. Cockerell 1913 praying mantis USNM 58583 (Gustav Hambach coll))
Hymenoptera - Formicidae	Lasius peritulus¹, Formica robusta¹, Formica cockerelli¹, Formica grandis n. sp.¹, Elaeomyrmex coloradensis¹, Elaeomyrmex gracilis¹, Protazteca elongata¹, Protazteca capitata¹, Protazteca quadrata¹, Iridomyrmex obscurans n. sp.¹, Petraeomyrmex minimus¹, Miomyrmex impactus¹, Liometopum scudderi¹, Liometopum miocenicum¹, Aphaenogaster mayri¹, Messor sculpturatus¹ Lasius peritulus¹ Cockerell 1927 ant USNM 78802 Formica robusta¹ Carpenter 1930 ant USNM 78800 Formica cockerelli¹ Carpenter 1930 ant USNM 78806 Formica grandis n. sp.¹ Carpenter 1930 ant USNM 78801 (Lacoe collection) Elaeomyrmex coloradensis¹ Carpenter 1930 ant USNM 78798 Elaeomyrmex gracilis¹ Carpenter 1930 ant USNM 78797 Protazteca elongata¹ Carpenter 1930 ant USNM 78791 Protazteca capitata¹ Carpenter 1930 ant USNM 78793 Protazteca quadrata¹ Carpenter 1930 ant USNM 78792 Iridomyrmex obscurans n. sp.¹ Carpenter 1930 ant USNM 78799 (Lacoe collection) Petraeomyrmex minimus¹ Carpenter 1930 ant USNM 78805 Miomyrmex impactus¹ Cockerell 1927 ant USNM 78796 Liometopum scudderi¹ Carpenter 1930 ant USNM 78795 Liometopum miocenicum¹ Carpenter 1930 ant USNM 78794 Aphaenogaster mayri¹ Carpenter 1930 ant USNM 78803 Messor sculpturatus¹ Carpenter 1930 ant USNM 78804
Hymenoptera - Megachilidae	Heriades priscus n. sp.² Heriades priscus n. sp.² Cockerell 1917 mason bee USNM 62543
Hymenoptera -	Andrena lagopus¹¹ Andrena lagopus¹¹ Latreille 1809 bee
Hymenoptera - Tenthredinidae	Taeniurites fortis n. gen. n. sp.² Taeniurites fortis n. gen. n. sp.² Cockerell 1917 wood wasp USNM 62542
Hymenoptera - Xyelidae	Xyela (Xyela) florissantensis n. sp.⁶ Xyela (Xyela) florissantensis n. sp.⁶ Rasnitsyn 1995 sawfly USNM 127677
Neuroptera - Osmylidae	Osmylidia requieta Osmylidia requieta Scudder 1890 lance lacewing USNM 58681 (Gustav Hambach coll)
Coleoptera - Carabidae	Platynus insculptipennis n. sp.¹⁰, Amara veterata⁸, Amara powellii⁸, Pterostichus walcotti⁸, Harpalus ulomaeformis n. sp.¹⁰, Harpalus redivivus n. sp.¹⁰, Cratacanthus florissantensis n. sp.¹⁰ Platynus insculptipennis n. sp.¹⁰ Wickham 1917 ground beetle USNM 63441 Amara veterata⁸ Scudder 1900 ground beetle USNM 59645 Amara powellii⁸ Scudder 1900 ground beetle USNM 59646 Pterostichus walcotti⁸ Scudder 1900 ground beetle USNM 59644 Harpalus ulomaeformis n. sp.¹⁰ Wickham 1917 ground beetle USNM 63443 Harpalus redivivus n. sp.¹⁰ Wickham 1917 ground beetle USNM 63444 Cratacanthus florissantensis n. sp.¹⁰ Wickham 1917 ground beetle USNM 63442
Coleoptera - Dytiscidae	Agabus florissantensis n. sp.⁸ Agabus florissantensis n. sp.⁸ Wickham 1913 predaceous diving beetle USNM 59632
Coleoptera - Lucanidae	Lucanus fossilis n. sp.⁸ Lucanus fossilis n. sp.⁸ Wickham 1913 stag beetle USNM 59643
Coleoptera - Scarabaeidae	Diplotaxis aurora n. sp.⁸, Macrodactylus pluto⁸, Aphodius granarioides n. sp.⁸, Ataenius patescens⁸ Diplotaxis aurora n. sp.⁸ Wickham 1913 scarab beetle USNM 59657 Macrodactylus pluto⁸ Wickham 1912 scarab beetle USNM 59656 Aphodius granarioides n. sp.⁸ Wickham 1913 scarab beetle USNM 59655 Ataenius patescens⁸ Scudder 1900 scarab beetle USNM 59654
Coleoptera - Leiodidae	Anisotoma sibylla n. sp.⁸ Anisotoma sibylla n. sp.⁸ Wickham 1913 round fungus beetle USNM 59633
Coleoptera - Staphylinidae	Miolithocharis lithographica n. gen. n. sp.⁸, Quedius chamberlini⁸, Philonthus marcidulus⁸, Staphylinus vulcan n. sp.⁸, Aleocharopsis caseyi n. sp.⁸, Aleocharopsis secunda n. sp.⁸, Stenus morsei⁸ Miolithocharis lithographica n. gen. n. sp.⁸ Wickham 1913 rove beetle USNM 59638 Quedius chamberlini⁸ Scudder 1890 rove beetle USNM 59647 Philonthus marcidulus⁸ Scudder 1900 rove beetle USNM 59648 Staphylinus vulcan n. sp.⁸ Wickham 1913 rove beetle USNM 59636 Aleocharopsis caseyi n. sp.⁸ Wickham 1913 rove beetle USNM 59634 Aleocharopsis secunda n. sp.⁸ Wickham 1913 rove beetle USNM 59635 Stenus morsei⁸ Scudder 1900 rove beetle USNM 59637
Coleoptera - Ptinidae	Vrilletta monstrosa n. sp.¹⁰ Vrilletta monstrosa n. sp.¹⁰ Wickham 1917 spider beetle USNM 63447
Coleoptera - Trogossitidae	Tenebroides corrugata n. sp.⁸ Tenebroides corrugata n. sp.⁸ Wickham 1913 bark gnawing beetle USNM 59641
Coleoptera - Tenebrionidae	Blapstinus linellii n. sp.⁸, Miostenosis lacordairei n. gen. n. sp.⁸, Platydema bethunei n. sp.⁸ Blapstinus linellii n. sp.⁸ Wickham 1913 darkling beetle USNM 59664 Miostenosis lacordairei n. gen. n. sp.⁸ Wickham 1913 darkling beetle USNM 59663 Platydema bethunei n. sp.⁸ Wickham 1913 darkling beetle USNM 59665
Coleoptera - Coccinellidae	Anatis resurgens n. sp.¹⁰ Anatis resurgens n. sp.¹⁰ Wickham 1917 ladybird beetle USNM 63445
Coleoptera - Cryptophagidae	Cryptophagus bassleri n. sp.⁸ Cryptophagus bassleri n. sp.⁸ Wickham 1913 silken fungus beetle USNM 59640
Coleoptera - Laemophloeidae	Lithocoryne arcuata n. sp.⁸ Lithocoryne arcuata n. sp.⁸ Wickham 1913 beetle USNM 59639
Coleoptera - Attelabidae	"Rhynchites subterraneus" = Prodeporaides subterraneus⁸ "Rhynchites subterraneus" = Prodeporaides subterraneus⁸ Scudder 1893 tooth-nosed snout weevil USNM 59666
Coleoptera - Curculionidae	Rhysosternum longirostre⁸, "Baris primalis n. sp." = Lithogeraeus primalis¹⁰, Geralophus repositus⁸, Geralophus antiquarius⁸, Geralophus fossicius⁸, Geralophus lassatus⁸, Geralophus saxuosus⁸, Geralophus retritus⁸, Pandeleteinus nudus n. sp.¹⁰, Evopes veneratus⁸, "Coniatus evisceratus" = Listroderes evisceratus⁸, Dorytomus williamsi⁸, "Balaninus minusculus" = Curculio minusculus⁸, Tychius ferox n. sp.¹⁰, Cryptorhynchus profusus⁸, Cleonus foersteri⁸, Cleonus exterraneus⁸ Rhysosternum longirostre⁸ Scudder 1893 snout beetle USNM 59678 "Baris primalis n. sp." = Lithogeraeus primalis¹⁰ Wickham 1917 snout beetle USNM 63453 Geralophus repositus⁸ Scudder 1893 snout beetle USNM 59671 Geralophus antiquarius⁸ Scudder 1893 snout beetle USNM 59668 Geralophus fossicius⁸ Scudder 1893 snout beetle USNM 59670 Geralophus lassatus⁸ Scudder 1893 snout beetle USNM 59672 Geralophus saxuosus⁸ Scudder 1893 snout beetle USNM 59669 Geralophus retritus⁸ Scudder 1893 snout beetle USNM 59673 Pandeleteinus nudus n. sp.¹⁰ Wickham 1917 snout beetle USNM 63451 Evopes veneratus⁸ Scudder 1893 snout beetle USNM 59667 "Coniatus evisceratus" = Listroderes evisceratus⁸ Scudder 1893 snout beetle USNM 59674 Dorytomus williamsi⁸ Scudder 1893 snout beetle USNM 59677 "Balaninus minusculus" = Curculio minusculus⁸ Scudder 1893 snout beetle USNM 59680 Tychius ferox n. sp.¹⁰ Wickham 1917 snout beetle USNM 63452 Cryptorhynchus profusus⁸ Scudder 1893 snout beetle USNM 59679 Cleonus foersteri⁸ Scudder 1893 snout beetle USNM 59676 Cleonus exterraneus⁸ Scudder 1893 snout beetle USNM 59675
Coleoptera - Chrysomelidae	Systena florissantensis n. sp.⁸, Bruchus dormescens n. sp.⁸, Bruchus antaeus n. sp.¹⁰, Bruchus exhumatus⁸ Systena florissantensis n. sp.⁸ Wickham 1913 leaf beetle USNM 59660 Bruchus dormescens n. sp.⁸ Wickham 1913 leaf beetle USNM 59661 Bruchus antaeus n. sp.¹⁰ Wickham 1917 leaf beetle USNM 63450 Bruchus exhumatus⁸ Wickham 1912 leaf beetle USNM 59661
Coleoptera - Cerambycidae	"Callidium grandaevum n. sp." = Phymatodes grandaevus¹⁰, "Leptura antecurrens n. sp." = Judolia antecurrens⁸, Leptura ponderosissima n. sp.⁸, "Spondylis tertiarius n. sp." = Neandra florissantensis¹⁰ "Callidium grandaevum n. sp." = Phymatodes grandaevus¹⁰ Wickham 1917 longhorned beetle USNM 63449 "Leptura antecurrens n. sp." = Judolia antecurrens⁸ Wickham 1913 longhorned beetle USNM 59658 Leptura ponderosissima n. sp.⁸ Wickham 1913 longhorned beetle USNM 59659 "Spondylis tertiarius n. sp." = Neandra florissantensis¹⁰ Wickham 1920 longhorned beetle USNM 63448
Coleoptera - Buprestidae	Anthaxia exhumata n. sp.⁸ Anthaxia exhumata n. sp.⁸ Wickham 1913 jewel beetle USNM 59642
Coleoptera - Elateridae	Cardiophorus lithographicus⁹, Cardiophorus florissantensis⁹ Cardiophorus lithographicus⁹ Wickham 1916 click beetle Cardiophorus florissantensis⁹ Wickham 1916 click beetle
Coleoptera - Cantharidae	Chauliognathus pristinus⁸, Podabrus wheeleri⁸, Podabrus cupesoides n. sp.¹⁰ Chauliognathus pristinus⁸ Scudder 1876 soldier beetle USNM 59652 Podabrus wheeleri⁸ Wickham 1909 soldier beetle USNM 59653 Podabrus cupesoides n. sp.¹⁰ Wickham 1917 soldier beetle USNM 63446
Coleoptera - Byrrhidae	Byrrhus romingeri⁸, Amphicyrta inhaesa⁸, Nosotetocus debilis⁸ Byrrhus romingeri⁸ Scudder 1900 pill beetle USNM 59651 Amphicyrta inhaesa⁸ Scudder 1900 pill beetle USNM 59650 Nosotetocus debilis⁸ Scudder 1900 pill beetle USNM 59649
Trichoptera -	Nebritus willistoni n. sp.⁴ Nebritus willistoni n. sp.⁴ Melander 1949 caddisfly USNM 112553 (Lacoe coll)
Lepidoptera - Nymphalidae	"Jupiteria charon n. gen. n. sp." = Jupitellia charon⁷, "Chlorippe wilmattae" = Doxocopa wilmattae "Jupiteria charon n. gen. n. sp." = Jupitellia charon⁷ Scudder 1889 brush-footed butterfly Lacoe 2100 "Chlorippe wilmattae" = Doxocopa wilmattae Cockerell 1907 brush-footed butterfly USNM 58682 (Gustav Hambach coll)

¹Carpenter 1930; ²Cockerell 1917; ³Melander 1946; ⁴Melander 1949; ⁵Petrunkevitch 1922; ⁶Rasnitsyn 1995; ⁷Scudder 1889; ⁸Wickham 1913; ⁹Wickham 1916; ¹⁰Wickham 1917; ¹¹Zeuner and Manning 1976